Herbarium LE - Specimens

FLORA OF LAOS Fam. Begoniaceae Begonia viscosa Aver., sp.nov. Type Vientiane Prov., Vang Vieng Distr., vicinity of Vang Vieng town, around point 18°54’58”N 102°24’52”E. Secondary broadleaved evergreen dry forest on very steep rocky slopes and on the top of remnant mountain composed with highly eroded crystalline marble-like limestone at elevation 400-450 m a.s.l. Lithophytic undershrub with erect shoots to 2 m tall in crevices of open vertical cliffs. Flowers white, sometimes with light punk tint, anthers and stigmas yellow. Leaves above light green to dull gray or gray-brownish with palmate whitish marks, uniform dull purple below, petioles green to purple. Locally common. CPC 2438 24 April 2011 Photos. Coll.: L.Averyanov, P.V.The. d-EXSICCATES OF VIETNAMESE FLORA 0187/CPC 2438 (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VIETNAM Begoniaceae Begonia sonlaensis Aver. Epitype. Son La Prov., Yen Chau Distr., Muong Lum Municipality, Na Hat village around point 21°00’35”N 104°29’06”E. Primary coniferous forest with Pinus kwangtungensis and Calocedrus rupestris on very steep rocky slopes of remnant mountain composed with highly eroded solid limestone at elevation 1000-1150 m a.s.l. Lithophytic herb on vertical shady cliff. Occasional. CPC 1876a 1 April 2011 Coll.: L.Averyanov, N.T.Hiep, T.B.Ngan. Flowered under cultivation in April-May 2012. Flowers light pink. Almost odorless. Coll.: L.Averyanov, 23 April 2012. CPC 1876aa. Photos. d-EXSICCATES OF VIETNAMESE FLORA 0192/CPC 1876aa (LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF LAOS Hin Nam No Nature Protected Area Begoniaceae Begonia tatianae Aver. Laos, Khammoune prov., Boualapha distr., Vang Maner village, around point 17°29'39.3"N, 105°50'24.6"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 350-430 m a.s.l. Lithophytic herb on shady rocks near hill top. Leaves above pale green with white palmately oriented white spots, below light green to pale pinkish, petioles dull pinkish. Not common. 9 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 820b. Herbarium specimen prepared in 19.04.2019, L. Averyanov, AL 820b.1. Flowers white with light pink tint, anther and stigma bright yellow.

FLORA OF VIETNAM

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM

All photos © L. Averyanov & K.S. Nguyen

FLORA OF LAOS Hin Nam No Nature Protected Area Begoniaceae Begonia handelii Irmsch. Laos, Khammouane prov., Boualapha distr., Nong Seng village, southern sandstone slopes at summit of Phou Chuang (Poujeuang) Mt. around point 17°35'18.1"N, 105°47'47.5"E. Primary evergreen mixed and coniferous dense wind-formed forest at elev. 1000-1400 m a.s.l. Lithophytic herb on mossy rocks. Leaves and petioles uniform green. Occasional. 5 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 657b (photo only).

FLORA OF VIETNAM

All photos © L. Averyanov & K.S. Nguyen

FLORA OF LAOS Hin Nam No Nature Protected Area Begoniaceae Begonia sp. Laos, Khammoune prov., Boualapha distr., Nong Seng village, Chanh Daeng mountain, around point 17°31'11.1"N, 105°46'41.4"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 400-600 m a.s.l. Lithophytic herb on shady cliff near mountain top. Leaves uniform pale green above, almost white with brownish-pink nerves below. Locally common. 11 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 971.

FLORA OF LAOS Hin Nam No Nature Protected Area Begoniaceae Begonia sp. Laos, Khammoune prov., Boualapha distr., Vang Maner village, around point 17°29'39.3"N, 105°50'24.6"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 350-430 m a.s.l. Lithophytic herb on shady rocks near hill top. Leaves above pale green with white palmately oriented white spots, below light green to pale pinkish, petioles dull pinkish. Not common. 9 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 820a.

Upper specimen on the image

FLORA OF LAOS Hin Nam No Nature Protected Area Begoniaceae Begonia khammouanensis Souvann. & Lamxay ? Laos, Khammoune prov., Boualapha distr., Ban Dou village, around Khuay cave, at 17°30'35.1"N, 105°51'28.7"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 250-390 m a.s.l. Lithophytic herb on shady cliff near mountain top. Leaves uniform green above, light pink below. Not rare. 10 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 897.

FLORA OF VIETNAM Plants of Phong Nha – Ke Bang National Park Fam. Begoniaceae Begonia gesneriifolia Aver. Epitype Vietnam, Quang Binh Prov., Bo Trach Distr.,Son Trach Municipality, around point 17º30’32’’N, 106º17’45’’E at elev. 300-400 m a.s.l. Secondary closed broadleaved evergreen forest and scrub on rocky steep slopes of remnant mountains composed with solid highly eroded crystalline deep gray limestone. Lithophytic herb on vertical shady cliff. Flowers white, anthers yellow. Locally common. No HAL 6354 2 February 2005. Collectors: L.Averyanov, P.K.Loc, P.V.The, A.Averyanova, N.T.Vinh, N.Q.Vinh, N.T.Binh. d-EXSICCATES OF VIETNAMESE FLORA 0011/HAL6354 (HN) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VIETNAM Begoniaceae Begonia rugosula Aver., sp.nov. Type Bac Kan Prov., Cho Don Distr., Xuan Lac municipality, Lung Ly locality, around point 22°17'24"N, 105°30'10"E, at elev. about 950 m. In partially destroyed primary closed evergreen broad-leaved forests near top of limestone mountain. Lithophyte. CPC 1260a 6 Marchch 2011 Coll.: N.Q.Hieu, N.T.Hiep, P.K.Loc, P.V.The, N.T.Vinh Flowered under cultivation at May 2011. Male flowers white, tepals light olive-green at base, filaments and anthers yellow; female flowers white, sometimes with light pink tint, tepals light olive-green at base, stiles and stigmas yellow. Photos. d-EXSICCATES OF VIETNAMESE FLORA 0185/CPC 1260a (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF LAOS Fam. Begoniaceae Begonia viscosa Aver., sp.nov. Type Vientiane Prov., Vang Vieng Distr., vicinity of Vang Vieng town, around point 18°54’58”N 102°24’52”E. Secondary broadleaved evergreen dry forest on very steep rocky slopes and on the top of remnant mountain composed with highly eroded crystalline marble-like limestone at elevation 400-450 m a.s.l. Lithophytic undershrub with erect shoots to 2 m tall in crevices of open vertical cliffs. Flowers white, sometimes with light punk tint, anthers and stigmas yellow. Leaves above light green to dull gray or gray-brownish with palmate whitish marks, uniform dull purple below, petioles green to purple. Locally common. CPC 2438 24 April 2011 Photos. Coll.: L.Averyanov, P.V.The. d-EXSICCATES OF VIETNAMESE FLORA 0187/CPC 2438 (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VIETNAM Begoniaceae Begonia crassula Aver. sp.nov. Paratype Quang Binh Prov., Minh Hoa Distr., Thuong Hoa municipality, environs of Mo O O O village, around point 17°39’36.7”N 105°54’55.7”E. Remnants of short tall broad-leaved primary forest on rocky top of remnant mountain composed with highly eroded solid crystalline limestone at elevation about 612 m a.s.l. Lithophytic herb or undershrub to 1 m tall on exposed open rocks on rocky mountain top. Flowers light pink, anther and stigma yellow. Locally common. CPC 3775 26 July 2011 Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Vinh. d-EXSICCATES OF VIETNAMESE FLORA 0189/CPC 3775 (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VIETNAM Fam. Begoniaceae Begonia minuscula Aver., sp.nov. Type Quang Tri Prov., Huong Hoa Distr., Huong Viet municipality around point 16°51’06”N 106°34’38”E. Partially destroyed primary broadleaved evergreen forest on very steep rocky slopes of remnant hills composed with solid highly eroded crystalline limestone at elev. about 600-700 m a.s.l. Lithophytic herb on vertical tall cliff. Leaves uniform green, pinkish or brown, sometimes with whitish spots. Not common. CPC 2838 7 May 2011 Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, P.V.The, N.T.Vinh. d-EXSICCATES OF VIETNAMESE FLORA 0184/HAL 2838 (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VIETNAM Fam. Begoniaceae Begonia rubrosetosa Aver. Epitype Thua Thien – Hue Prov., Nam Dong Distr., Thuong Lo Municipality, along Cha Mang stream, around point 16°07’35’’N 107°44’53’’E. Remnants of primary evergreen broad-leaved lowland forest on very steep hill slopes composed with stratified shale and sandstone at elev. about 150 m a.s.l. Terrestrial and lithophytic herb on very steep rocky slope. Leaves uniform green with reddish nerves. Flowers red, anther and stigmas yellow. Locally very common. HAL 10883 4 Apr. 2007. Photo. Coll.: L.Averyanov, Phan Ke Loc, A.Averyanova, N.T.Vinh, N.D.Phuong, L.V.Hung. Det. N.Q.Hieu d-EXSICCATES OF VIETNAMESE FLORA 0104/HAL 10883 (HN, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF LAOS Begoniaceae Begonia quadripetiolata Aver., sp. nov. Vientiane prov., Vang Vieng distr., Oa Hom village, around point 19°06'52.4''N 102°22'35.8''E. Secondary broad-leaved evergreen and semideciduous dry forest with on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. 550-650 m a.s.l. Lithophytic herb with stem 10-20 cm tall on shady cliffs. Leaves and petioles uniform green. Locally very common. LA-VN 1510 22 March 2017 Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.Maisak, Khamla Inkhavilay, Hongthong Southyvohanh, Pinmany Sisavath, Chanthala Cheuaysamphanh. Fl. in cult. 20 March 2019. Flowers odorless, white, anther yellow, only male flowers. L. Averyanov, T. Maisak, LA-VN 1510/1. Photos.

FLORA OF VIETNAM Fam. Begoniaceae Begonia minuscula Aver., sp.nov. Type Quang Tri Prov., Huong Hoa Distr., Huong Viet municipality around point 16°51’06”N 106°34’38”E. Partially destroyed primary broadleaved evergreen forest on very steep rocky slopes of remnant hills composed with solid highly eroded crystalline limestone at elev. about 600-700 m a.s.l. Lithophytic herb on vertical tall cliff. Leaves uniform green, pinkish or brown, sometimes with whitish spots. Not common. CPC 2838 7 May 2011 Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, P.V.The, N.T.Vinh. d-EXSICCATES OF VIETNAMESE FLORA 0184/HAL 2838 (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VIETNAM Fam. Begoniaceae Begonia sp. N. Vietnam, Bac Can prov., Na Ri distr., Kim Hy municipality, vicinities of Lung Hin Con village (N 22º17' E 106º03'), 27 km to 55º of Bac Can city. Primary evergreen mixed mossy forest with Pseudotsuga brevifolia and bamboo along tops of karst remnant limestone ridge at elev. 700 m a.s.l. Adpressed lithophytic forb in crevices of shady verical cliffs. Leaves below violet, above deep green with irregular palmate white spot. Very common. Usual codominanthe of lithophytic forb communities. No NTH 3725 20 Oct. 1999 Collectors: N.T. Hiep, P.K. Loc, L. Averyanov.

FLORA OF VIETNAM Begoniaceae Begonia crassula Aver. sp.nov. Paratype Quang Binh Prov., Minh Hoa Distr., Thuong Hoa municipality, environs of Mo O O O village, around point 17°39’36.7”N 105°54’55.7”E. Remnants of short tall broad-leaved primary forest on rocky top of remnant mountain composed with highly eroded solid crystalline limestone at elevation about 612 m a.s.l. Lithophytic herb or undershrub to 1 m tall on exposed open rocks on rocky mountain top. Flowers light pink, anther and stigma yellow. Locally common. CPC 3775 26 July 2011 Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Vinh. d-EXSICCATES OF VIETNAMESE FLORA 0189/CPC 3775 (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VIETNAM Begoniaceae Begonia minuscula Aver. Paratype Vietnam, Quang Tri Prov., Huong Hoa Distr., Huong Viet Municipality, to N of Sa Mui pass, around point 16º51’14’’N 106º34’13’’E at elevation 550-650 m a.s.l. Remnants of primary broad-leaved evergreen lowland forest on rocky remnant highly eroded solid marble-like limestone hills and low mountains with very steep slopes and vertical cliffs. Lithophytic creeping herb on vertical shady cliffs. Leaves pure green to pink-brown. Very common. HLF 5910 31 March 2006. Coll. N.T.Hiep, L.Averyanov, P.K.Loc, P.V.The, N.S.Khang, P.H.Duc, L.T.Son, S.Bounphanmy, T.K.Ratana, O.Sonliya, T.Q.Dang, N.V.Hien, H.T.N.Trang, L.T.Chau. Grant from Henry Luce Foundation.

FLORA OF VIETNAM Fam. Begoniaceae Begonia minuscula Aver., sp.nov. Type Quang Tri Prov., Huong Hoa Distr., Huong Viet municipality around point 16°51’06”N 106°34’38”E. Partially destroyed primary broadleaved evergreen forest on very steep rocky slopes of remnant hills composed with solid highly eroded crystalline limestone at elev. about 600-700 m a.s.l. Lithophytic herb on vertical tall cliff. Leaves uniform green, pinkish or brown, sometimes with whitish spots. Not common. CPC 2838 7 May 2011 Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, P.V.The, N.T.Vinh. d-EXSICCATES OF VIETNAMESE FLORA 0184/HAL 2838 (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VIETNAM Fam. Begoniaceae Begonia rubrosetosa Aver. Epitype Thua Thien – Hue Prov., Nam Dong Distr., Thuong Lo Municipality, along Cha Mang stream, around point 16°07’35’’N 107°44’53’’E. Remnants of primary evergreen broad-leaved lowland forest on very steep hill slopes composed with stratified shale and sandstone at elev. about 150 m a.s.l. Terrestrial and lithophytic herb on very steep rocky slope. Leaves uniform green with reddish nerves. Flowers red, anther and stigmas yellow. Locally very common. HAL 10883 4 Apr. 2007. Photo. Coll.: L.Averyanov, Phan Ke Loc, A.Averyanova, N.T.Vinh, N.D.Phuong, L.V.Hung. Det. N.Q.Hieu d-EXSICCATES OF VIETNAMESE FLORA 0104/HAL 10883 (HN, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VIETNAM Begoniaceae Begonia crassula Aver. sp.nov. Paratype Quang Binh Prov., Minh Hoa Distr., Thuong Hoa municipality, environs of Mo O O O village, around point 17°39’36.7”N 105°54’55.7”E. Remnants of short tall broad-leaved primary forest on rocky top of remnant mountain composed with highly eroded solid crystalline limestone at elevation about 612 m a.s.l. Lithophytic herb or undershrub to 1 m tall on exposed open rocks on rocky mountain top. Flowers light pink, anther and stigma yellow. Locally common. CPC 3775 26 July 2011 Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Vinh. d-EXSICCATES OF VIETNAMESE FLORA 0189/CPC 3775 (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VIETNAM Begoniaceae Begonia rigidifolia Aver., sp.nov. Type Quang Binh Prov., Tuyen Hoa Distr., Lam Hoa municipality, Chuoi village, around point 17°56’51”N 105°49’18”E. Primary broadleaved evergreen dry forest on very steep slopes and on tops of rocky ridge composed with highly eroded crystalline limestone at elev. 200-250 m a.s.l. Lithophytic rosulate herb on shady vertical cliffs. Flowers light pinkish, anther and stigmas yellow, leaves dark to light green with irregular dirty brownish or dark green irregular palmate markings above, light green, nearly white below, with unclear dirty purple marks. Petioles dull brownish-purple. Not rare. CPC 2733 3 May 2011 Photos. Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, P.V.The, N.T.Vinh. d-EXSICCATES OF VIETNAMESE FLORA 0186/CPC 2733 (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF LAOS Fam. Begoniaceae Begonia viscosa Aver., sp.nov. Type Vientiane Prov., Vang Vieng Distr., vicinity of Vang Vieng town, around point 18°54’58”N 102°24’52”E. Secondary broadleaved evergreen dry forest on very steep rocky slopes and on the top of remnant mountain composed with highly eroded crystalline marble-like limestone at elevation 400-450 m a.s.l. Lithophytic undershrub with erect shoots to 2 m tall in crevices of open vertical cliffs. Flowers white, sometimes with light punk tint, anthers and stigmas yellow. Leaves above light green to dull gray or gray-brownish with palmate whitish marks, uniform dull purple below, petioles green to purple. Locally common. CPC 2438 24 April 2011 Photos. Coll.: L.Averyanov, P.V.The. d-EXSICCATES OF VIETNAMESE FLORA 0187/CPC 2438 (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF LAOS Begoniaceae Begonia quadripetiolata Aver., sp. nov. Vientiane prov., Vang Vieng distr., Oa Hom village, around point 19°06'52.4''N 102°22'35.8''E. Secondary broad-leaved evergreen and semideciduous dry forest with on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. 550-650 m a.s.l. Lithophytic herb with stem 10-20 cm tall on shady cliffs. Leaves and petioles uniform green. Locally very common. LA-VN 1510 22 March 2017 Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.Maisak, Khamla Inkhavilay, Hongthong Southyvohanh, Pinmany Sisavath, Chanthala Cheuaysamphanh. Fl. in cult. 20 March 2019. Flowers odorless, white, anther yellow, only male flowers. L. Averyanov, T. Maisak, LA-VN 1510/1. Photos.

FLORA OF VIETNAM Begoniaceae Begonia acetosella Craib Ha Giang prov., Quan Ba distr., Tung Vai commune, Thang village, around point 23º03ʹ13.4ʺN 104º51ʹ48.8ʺE, at elevation 1050-1150 m a.s.l., steep slopes of stream valley composed with eroded stratified highly eroded limestone. Primary evergreen broad-leaved very humid forest. Terrestrial and lithophytic herb to 1 m tall. Leaves and petioles uniform green. Flowers light pink to almost white. Very common. 21 April 2018, L. Averyanov, Nguyen Sinh Khang, Nguyen Tien Hiep, Nguyen Quang Hieu, Chuong Quang Ngan, T. Maisak, VR 527.

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM Begoniaceae Begonia cavaleriei H. Lév. Ha Giang prov., Quan Ba distr., Tung Vai commune, Thang village, around point 23º03ʹ41.5ʺN 104º50ʹ41.6ʺE, steep rocky slopes near mountain top composed with eroded stratified highly eroded limestone at elevation 1200-1400 m a.s.l. Primary evergreen broad-leaved very humid forest. Lithophytic herb on open mossy rocks. Flowers pink, leaves and petioles uniformly green. Very common. 22 April 2018, L. Averyanov, Nguyen Sinh Khang, Nguyen Tien Hiep, Nguyen Quang Hieu, Chuong Quang Ngan, T. Maisak, VR 644.

All photos © L. Averyanov & K.S. Nguyen

FLORA OF LAOS Hin Nam No Nature Protected Area Begoniaceae Begonia tatianae Aver. Laos, Khammoune prov., Boualapha distr., Vang Maner village, around point 17°29'39.3"N, 105°50'24.6"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 350-430 m a.s.l. Lithophytic herb on shady rocks near hill top. Leaves above pale green with white palmately oriented white spots, below light green to pale pinkish, petioles dull pinkish. Not common. 9 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 820. Herbarium specimen prepared in 19.04.2019, L. Averyanov, AL 820.1. Flowers white with light pink tint, anther and stigma bright yellow.

FLORA OF LAOS

FLORA OF LAOS Begoniaceae Begonia quadripetiolata Aver., sp. nov. Vientiane prov., Vang Vieng distr., Oa Hom village, around point 19°06'52.4''N 102°22'35.8''E. Secondary broad-leaved evergreen and semideciduous dry forest with on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. 550-650 m a.s.l. Lithophytic herb with stem 10-20 cm tall on shady cliffs. Leaves and petioles uniform green. Locally very common. LA-VN 1510 22 March 2017 Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.Maisak, Khamla Inkhavilay, Hongthong Southyvohanh, Pinmany Sisavath, Chanthala Cheuaysamphanh.

FLORA OF VIETNAM Begoniaceae Begonia sp. Nghe An Prov., Ky Son Distr., Na Ngoi Municipality, Phu Kha village, eastern slopes of Phu Xai Lai Leng mountain system. Primary broad-leaved humid forest on steep granite slopes at elevation about 2350 m a.s.l. around point 19°12’21.2’’N 104°11’28.9’’E. Tuberiferous herb 5-10 cm tall. Rare. 23 October 2013 CPC 6072 Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, L.M.Tuan, N.A.Trang, L.H.Dang

FLORA OF VIETNAM Begoniaceae Begonia sp. Cao Bang Prov., Nguyen Binh Distr., Yen Lac Municipality, Chi Doi village, around point 22°45'34"N, 105°51'48.7"E. Primary fractionaly logged coniferous forest with Pseudotsuga sinensis along highly eroded rocky limestone ridge at elevation about 1400 m a.s.l. Lithophytic herb on shady cliffs on miffle part of mountain slope. Leaves dark green above, light green with purple veins below. Flowers pink, anthers yellow. Very common. 5 October 2013 CPC 5421 L.Averyanov, N.T.Hiep, L.M.Tuan, N.S.Khang, T.Maisak, L.Osinovets Photos

FLORA OF VIETNAM Begoniaceae Begonia sp. Cao Bang Prov., Thong Nong Distr., Yen Son Municipality, Nhieu Lung village, around point 22°46'07.9"N, 105°53'59.4"E. Primary fractionaly logged coniferous forest with Pseudotsuga sinensis along highly eroded rocky limestone ridge at elevation 1150-1250 m a.s.l. Lithophytic rosulate herb on vertical cliffs in shady humid place in rocky valley. Leaves dark green with light green palmate markings above, light green with purple nerves below. Flowers pink, anther yellow. Locally very common. 8 October 2013 CPC 5472 L.Averyanov, N.T.Hiep, L.M.Tuan, N.S.Khang, T.Maisak, L.Osinovets

FLORA OF VIETNAM Begoniaceae Begonia “arachna” sp.nov.? Ha Giang Prov., Bac Me Distr., Phieng Luong municipality, around Phieng Luong village. Primary broad-leaved humid evergreen forest on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation 1100-1150 m a.s.l. around point 22°38’07.1’’N 105°19’22.1’’E. Lithophytic rosulate herb on shady vertical cliffs. Flowers light pink to almost white, ant hers and stigma yellow, Fruits purple. Leaves green, dark red-purple below, petioles dull purple. Very common. 13 November 2014 CPC 7391 Photos. Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz

FLORA OF VIETNAM Begoniaceae Begonia cavaleriei H. Lév. Ha Giang prov., Quan Ba distr., Tung Vai commune, Thang village, around point 23º03ʹ41.5ʺN 104º50ʹ41.6ʺE, steep rocky slopes near mountain top composed with eroded stratified highly eroded limestone at elevation 1200-1400 m a.s.l. Primary evergreen broad-leaved very humid forest. Lithophytic herb on open mossy rocks. Flowers pink, leaves and petioles uniformly green. Very common. 22 April 2018, L. Averyanov, Nguyen Sinh Khang, Nguyen Tien Hiep, Nguyen Quang Hieu, Chuong Quang Ngan, T. Maisak, VR 644.

All photos © L. Averyanov & K.S. Nguyen

FLORA OF LAOS Begoniaceae Begonia sp. Vientiane Prov., Vang Vieng Distr., Na Po village, about 12 km to the W of Vang Vieng town, Pa Nang Oua Mt., around point 18°55’44.9”N 102°20’20.1”E. Dry broad-leaved primary and secondary evergreen and semideciduous forest on very steep rocky slopes and on vertical cliffs of remnant highly eroded rocky limestone mountains composed by solid crystalline limestone at elevation 700-1000 m. Creeping lithophyte on shady cliff near mountain top. Leaves dark green to olive-brown with white spots above, dull violet below with unclear white spots, petiole purple. Locally common. LA-VN 521 17 March 2013 Coll.: N.T.Hiep, L.Averyanov, K.Chantthavongsa, N.S.Khang, P.V.The, S.Lorphengsy

FLORA OF LAOS Hin Nam No Nature Protected Area Begoniaceae Begonia sp. Laos, Khammoune prov., Boualapha distr., Vang Maner village, Palai mountain area, around point 17°30'13.6"N, 105°49'23.9"E, 520 m a.s.l. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 300-500 m a.s.l. Lithophytic clustering herb on small shady shelves of vertical cliff near mountain top. Leaves uniform green. Occasional. 8 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 795.

FLORA OF VIETNAM Begoniaceae Begonia khangii sp.nov.? Nghe An Prov., Ky Son Distr., Na Ngoi Municipality, eastern slopes of Phu Xai Lai Leng Mountain. Primary very humid broad-leaved forest on steep mountain slopes composed of sandstone and gray shale at elevation 2100-2200 m a.s.l. around point 19°11’59.3’’N 104°11’23.9’’E. Lithophytic herb on mossy rocks. Flowers pink, anthers and stigmas yellow. Leaves below with dull dirty purple spots. Very common. 25 October 2013 CPC 6281 Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, L.M.Tuan, N.A.Trang, L.H.Dan

FLORA OF VIETNAM Begoniaceae Begonia sp. Tuyen Quang Prov., Na Hang Distr., Sinh Long Municipality, Khuoi Phin village, around point 22°38'21.9"N, 105°20'43.5"E. Primary evergreen mixed humid forest on rocky limestone on highly eroded karstic hills composed with gray solid marble-like limestone at elevation 1100-1200 m a.s.l. Lithophytic herb on vertical cliffs. Leaves above uniform green, below brightly red. Flowers pink. Abundant. 29 September 2013 CPC 5317 L.Averyanov, N.T.Hiep, L.M.Tuan, N.S.Khang, D.V.Cuong, N.V.Truong.T.Maisak, L.Osinovets

FLORA OF VIETNAM Begoniaceae Begonia sp. Cao Bang Prov., Thong Nong Distr., Yen Son Municipality, Nhieu Lung village, around point 22°46'07.9"N, 105°53'59.4"E. Primary fractionaly logged coniferous forest with Pseudotsuga sinensis along highly eroded rocky limestone ridge at elevation 1150-1250 m a.s.l. Lithophytic rosulate herb on vertical cliffs in shady humid place in rocky valley. Leaves dark green with light green palmate markings above, light green with purple nerves below. Flowers pink, anther yellow. Locally very common. 8 October 2013 CPC 5472 L.Averyanov, N.T.Hiep, L.M.Tuan, N.S.Khang, T.Maisak, L.Osinovets

FLORA OF VIETNAM