Herbarium LE - Specimens

Gesneriaceae

Quang Nam Province, Nong Sơn District, Elephant Species and Habitat Conservation Area. Secondary dry evergreen broad-leaved forest on step mountain slopes composed by sandstone at elevation of about 200-250 m a.s.l. Terrestrial herb to 1 m tall on shady rocky steep slope along small rocky river. Occasional.

15 February 2023, Nguyen Van Canh, AL4441 (photo only).

Convallariaceae

Aspidistra multiflora Aver. & Tillich

Quang Nam Province, Nong Sơn District, Elephant Species and Habitat Conservation Area. Secondary dry evergreen broad-leaved forest on step mountain slopes composed by sandstone at elevation of about 200-250 m a.s.l. Terrestrial and lithophytic herb with short branching rhizome on shady rocky steep slope along small rocky river. Locally common.

15 February 2023, Nguyen Van Canh, AL4440 (photo only).

Orchidaceae

Dendrobium namdongense Aver. & C.H.Nguyen

Ha Giang Province, Quan Ba District, forest on the territory of Can Ty Commune.

17 May 2025, Truong Ba Vuong, BV 1934

The plant is slightly differs from the type of D. namdongense by longer mid lobes and side lobes erect, rectangular-square, apex truncate.

Orchidaceae

Schoenorchis spatulata (Rolfe ex Downie) Seidenf. & Smitinand

Lai Chau province, Tam Duong District, forest in Binh Lu Commune, inflorescence pendulous (?), peduncle with bilateral-wings, floral bract broadly ovate, acuminate. This species is similar to Schoenorchis gemmata but truly differs by the lip margin undulate incurved (vs spreading margin).

15 June 2024, Truong Ba Vuong, BV 1935

Orchidaceae

Vanilla sp. nov.? (V. siamensis aff.)

Di Linh District (or Duc Trong District).

17 June 2025, Nguyen Van Canh, AL4437.

Selaginellaceae

Selaginella sp.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Secondary broad-leaved evergreen forest with bamboo on rocky sandstone slopes at elevation 10–50 m a.s.l. Terrestrial herb in open rocky place along forest trail. Not common.

22 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4429 (NHC222).

Arecaceae

Calamus sp.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Secondary broad-leaved evergreen forest with bamboo on rocky sandstone slopes at elevation 10–50 m a.s.l. Climbing and ascending woody vine to 5-6 m long on very steep slope of small rocky stream. Fruits greenish. Common.

21 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4428 (NHC221).

Arecaceae

Calamus sp.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Secondary broad-leaved evergreen forest with bamboo on rocky sandstone slopes at elevation 10–50 m a.s.l. Ascending suberect woody vine to 6 m tall on very steep slope of small rocky stream. Not rare.

21 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4427 (NHC220).

Convallariaceae

Aspidistra sp.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Secondary broad-leaved evergreen forest with bamboo on rocky sandstone slopes at elevation 10–50 m a.s.l. Terrestrial and lithophytic rhizomatous herb on steep slopes of small shady rocky stream. Leaves to 0.6 m long, leaf blade linear, uniform grassy green. Fruit green globular, 5-7 mm in diameter. Locally very common.

21 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4426 (NHC219).

Polypodiaceae s.l.

Tectaria sp.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Secondary broad-leaved evergreen forest with bamboo on rocky sandstone slopes at elevation 10–50 m a.s.l. Terrestrial fern in open place along forest trail. Common.

21 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4425 (NHC218).

Polypodiaceae s.l.

Tectaria sp.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Secondary broad-leaved evergreen forest with bamboo on rocky sandstone slopes at elevation 10–50 m a.s.l. Creeping lithophytic fern on wet rocky slope along small shady forest stream. Common.

21 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4424 (NHC217).

Rubiaceae

Hedyotis sp.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Secondary broad-leaved evergreen forest with bamboo on rocky sandstone slopes at elevation 10–50 m a.s.l. Erect terrestrial herb to 0.5 m tall in open place along forest trail. Flowers white. Common.

21 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4423 (NHC216).

Araceae

Acorus gramineus?

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Secondary broad-leaved evergreen forest with bamboo on rocky sandstone slopes at elevation 10–50 m a.s.l. Lithophytic herb on wet mossy rocks along small rocky stream. Locally common.

21 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4422 (NHC215).

Chloranthaceae

Sarcandra sp.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Secondary broad-leaved evergreen forest with bamboo on rocky sandstone slopes at elevation 10–50 m a.s.l. Erect terrestrial herb to 0.6 m tall in open wet place along small rocky stream. Flowers bright yellow. Occasional.

21 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4421 (NHC214).

Polypodiaceae s.l.

Leptochilus sp.?

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Secondary broad-leaved evergreen forest with bamboo on rocky sandstone slopes at elevation 40–100 m a.s.l. Terrestrial or lithophytic fern on shady rocky slope. Not common.

21 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4419 (NHC212).

Gnetaceae

Gnetum sp.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Secondary broad-leaved evergreen forest with bamboo on rocky sandstone slopes at elevation 40–100 m a.s.l. Woody vine about 6-8 m tall. Male strobiles light green. Not common.

21 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4418 (NHC211).

Anacardiaceae

Spondias sp.?

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Secondary broad-leaved evergreen forest with bamboo on rocky sandstone slopes at elevation 40–100 m a.s.l. Tree to 6-8 m tall in rather open place. Flowers white very fragrant. Not common.

21 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4417 (NHC210).

Capparidaceae??

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Secondary broad-leaved evergreen forest with bamboo on rocky sandstone slopes at elevation 40–100 m a.s.l. Woody vine about 3 m long in rather open place. Fruits dull green. Not common.

21 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4416 (NHC209).

Rubiaceae

Ophiorrhiza sp.?

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Secondary broad-leaved evergreen forest with bamboo on rocky sandstone slopes at elevation 40–100 m a.s.l. Erect terrestrial herb or undershrub to 0.8 m tall in rather open place. Flower buds whitish. Occasional.

21 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4415 (NHC208).

Euphorbiaceae

Mallotus sp.?

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Secondary broad-leaved evergreen forest with bamboo on rocky sandstone slopes at elevation 40–100 m a.s.l. Shrub 2-3 m tall in rather open place. Fruits dull brownish-purple. Common.

21 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4414 (NHC207).

Fabaceae

Baphia capparidifolia Baker

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Secondary broad-leaved evergreen forest with bamboo on rocky sandstone slopes at elevation 40–100 m a.s.l. Woody vine to 3-4 m long in rather open place. Flowers white. Not common.

21 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4413 (NHC206).

Flacourtiaceae

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Secondary broad-leaved evergreen forest with bamboo on rocky sandstone slopes at elevation 40–100 m a.s.l. Woody vine to 6 m long in rather open place. Fruits bright orange. Not common.

21 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4412 (NHC205).

Euphorbiaceae

Mallotus hookerianus (Seem.) Müll.Arg.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Secondary broad-leaved evergreen forest with bamboo on rocky sandstone slopes at elevation 40–100 m a.s.l. Shrub or small treelet to 3-4 m tall. Fruits light green. Not common.

21 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4411 (NHC204).

Tiliaceae (Sterculiaceae)?

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Secondary broad-leaved evergreen forest with bamboo on rocky sandstone slopes at elevation 40–100 m a.s.l. Woody vine to 8 m long in open place along trail. Flowers light yellowish, anthers yellow. Not common.

21 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4410 (NHC203).

Polygalaceae

Xanthophyllum hainanense Hu

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Secondary broad-leaved evergreen forest with bamboo on rocky sandstone slopes at elevation 40–100 m a.s.l. Tree to 5 m tall and 10 cm DBH in open place. Fruits dark green. Not common.

21 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4409 (NHC202).

Moraceae

Ficus sp.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Secondary broad-leaved evergreen forest with bamboo on rocky sandstone slopes at elevation 40–100 m a.s.l. Woody vine to 5 m long in open place. Syconia pale green. Not common.

21 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4408 (NHC201).

Lauraceae

Litsea sp.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Young secondary broad-leaved evergreen forest on rocky sandstone slope along seashore at 0–5 m a.s.l. Tree 10-12 m tall in open place in sublittoral not flooded zone along rocky seashore. Not common.

20 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4407 (NHC200).

Sapotaceae

Planchonella obovata (R.Br.) Pierre?

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Young secondary broad-leaved evergreen forest on rocky sandstone slope along seashore at 0–5 m a.s.l. Treelet or shrub to 4 m tall in open place in sublittoral not flooded zone along rocky seashore. Flower buds greenish. Not common.

20 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4406 (NHC199).

Rhizophoraceae

Ceriops tagal (Perr.) C.B.Rob.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Young secondary broad-leaved evergreen forest on rocky sandstone slope along seashore at 0–5 m a.s.l. Treelet or shrub to 2 m tall among mangrove shrubs along rocky seashore. Flower buds yellowish-green. Common.

20 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4405 (NHC198).

Verbenacae

Avicennia marina (Forssk.) Vierh.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Young secondary broad-leaved evergreen forest on rocky sandstone slope along seashore at 0–5 m a.s.l. Treelet or shrub to 2 m tall in open place in sublittoral not flooded zone along rocky seashore. Flower buds greenish. Common.

20 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4404 (NHC197).

Loranthaceae

Scurrula ferruginea (Roxb. ex Jack) Danser?

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Young secondary broad-leaved evergreen forest on rocky sandstone slope along seashore at 0–5 m a.s.l. Parasitic shrub with erect shoot to 1 m long in canopies of tall tree (Litsea). Flowers outside brownish-yellow, inside dark violet. Not common.

20 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4403 (NHC196).

Smilacaceae

Smilax sp.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Young secondary broad-leaved evergreen forest on rocky sandstone slope along seashore at 0–5 m a.s.l. Semi-woody vine to 2-3 m long among dense shrubs in sublittoral not flooded zone along rocky seashore. Flowers light green. Not common.

20 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4402 (NHC195).

Rubiaceae

Mussaenda sp.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Young secondary broad-leaved evergreen forest on rocky sandstone slope along seashore at 0–5 m a.s.l. Woody vine to 5 m long in dense shrubs in sublittoral not flooded zone along rocky seashore. Fruits dark green. Not common.

20 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4401 (NHC194).

Combretaceae

Lumnitzera racemosa Willd.⁣

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Young secondary broad-leaved evergreen forest on rocky sandstone slope along seashore at 0–5 m a.s.l. Treelet or shrub to 2 m tall in open place in sublittoral not flooded zone along rocky seashore. Flower buds green. Common.

20 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4400 (NHC193).

Arecaceae

Korthalsia sp.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Young secondary broad-leaved evergreen forest on rocky sandstone slope along seashore at 0–5 m a.s.l. Woody vine to 5 m long in sublittoral not flooded zone along rocky seashore. Flowers dull yellow. Very common.

20 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4399 (NHC192).

Euphorbiaceae

Antidesma ghaesembilla Gaertn.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Young secondary broad-leaved evergreen forest on rocky sandstone slope along seashore at 0–5 m a.s.l. Undershrub to 1.5 m in shady place in sublittoral not flooded zone along rocky seashore. Flowers dull yellowish. Not rare.

20 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4398 (NHC191).

Com. Fischer

a. 1817

Com. Stephan a. 1814.

Pro Gal. ruthenico...

Steven: qui ex Iberia habit

et cultum ... ex sem.

...