Herbarium LE - Specimens

FLORA OF LAOS Fam. Begoniaceae Begonia viscosa Aver., sp.nov. Type Vientiane Prov., Vang Vieng Distr., vicinity of Vang Vieng town, around point 18°54’58”N 102°24’52”E. Secondary broadleaved evergreen dry forest on very steep rocky slopes and on the top of remnant mountain composed with highly eroded crystalline marble-like limestone at elevation 400-450 m a.s.l. Lithophytic undershrub with erect shoots to 2 m tall in crevices of open vertical cliffs. Flowers white, sometimes with light punk tint, anthers and stigmas yellow. Leaves above light green to dull gray or gray-brownish with palmate whitish marks, uniform dull purple below, petioles green to purple. Locally common. CPC 2438 24 April 2011 Photos. Coll.: L.Averyanov, P.V.The. d-EXSICCATES OF VIETNAMESE FLORA 0187/CPC 2438 (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VIETNAM Begoniaceae Begonia rugosula Aver., sp.nov. Type Bac Kan Prov., Cho Don Distr., Xuan Lac municipality, Lung Ly locality, around point 22°17'24"N, 105°30'10"E, at elev. about 950 m. In partially destroyed primary closed evergreen broad-leaved forests near top of limestone mountain. Lithophyte. CPC 1260a 6 Marchch 2011 Coll.: N.Q.Hieu, N.T.Hiep, P.K.Loc, P.V.The, N.T.Vinh Flowered under cultivation at May 2011. Male flowers white, tepals light olive-green at base, filaments and anthers yellow; female flowers white, sometimes with light pink tint, tepals light olive-green at base, stiles and stigmas yellow. Photos. d-EXSICCATES OF VIETNAMESE FLORA 0185/CPC 1260a (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VIETNAM Plants of Phong Nha – Ke Bang National Park Fam. Begoniaceae Begonia gesneriifolia Aver. Epitype Vietnam, Quang Binh Prov., Bo Trach Distr.,Son Trach Municipality, around point 17º30’32’’N, 106º17’45’’E at elev. 300-400 m a.s.l. Secondary closed broadleaved evergreen forest and scrub on rocky steep slopes of remnant mountains composed with solid highly eroded crystalline deep gray limestone. Lithophytic herb on vertical shady cliff. Flowers white, anthers yellow. Locally common. No HAL 6354 2 February 2005. Collectors: L.Averyanov, P.K.Loc, P.V.The, A.Averyanova, N.T.Vinh, N.Q.Vinh, N.T.Binh. d-EXSICCATES OF VIETNAMESE FLORA 0011/HAL6354 (HN) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VIETNAM Begoniaceae Begonia sonlaensis Aver. Epitype. Son La Prov., Yen Chau Distr., Muong Lum Municipality, Na Hat village around point 21°00’35”N 104°29’06”E. Primary coniferous forest with Pinus kwangtungensis and Calocedrus rupestris on very steep rocky slopes of remnant mountain composed with highly eroded solid limestone at elevation 1000-1150 m a.s.l. Lithophytic herb on vertical shady cliff. Occasional. CPC 1876a 1 April 2011 Coll.: L.Averyanov, N.T.Hiep, T.B.Ngan. Flowered under cultivation in April-May 2012. Flowers light pink. Almost odorless. Coll.: L.Averyanov, 23 April 2012. CPC 1876aa. Photos. d-EXSICCATES OF VIETNAMESE FLORA 0192/CPC 1876aa (LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

Tupistra aurantiaca Wall. Darjieling 7000 f 25 March 1876 27463 F Legit CB Clarke

East Himalaya

Sikkim Himalaya

Sikkim Himalaya

Ex horto bot. Petropolitano. 55.5.

Orontium japonicum Thunb.⁣

Ex horto bot. Prtropolitano. 66. 5.

Orontium japonicum Thunb.⁣

Rohdea japonica (Thunb.) Roth⁣

FLORA VIETNAMICA

FLORA OF LAOS Fam. Begoniaceae Begonia viscosa Aver., sp.nov. Type Vientiane Prov., Vang Vieng Distr., vicinity of Vang Vieng town, around point 18°54’58”N 102°24’52”E. Secondary broadleaved evergreen dry forest on very steep rocky slopes and on the top of remnant mountain composed with highly eroded crystalline marble-like limestone at elevation 400-450 m a.s.l. Lithophytic undershrub with erect shoots to 2 m tall in crevices of open vertical cliffs. Flowers white, sometimes with light punk tint, anthers and stigmas yellow. Leaves above light green to dull gray or gray-brownish with palmate whitish marks, uniform dull purple below, petioles green to purple. Locally common. CPC 2438 24 April 2011 Photos. Coll.: L.Averyanov, P.V.The. d-EXSICCATES OF VIETNAMESE FLORA 0187/CPC 2438 (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VIETNAM

FLORA OF LAOS

FLORA OF VIETNAM

FLORA OF VIETNAM

FLORA OF VIETNAM Convallariaceae Aspidistra lubae Aver. et Tillich var. lubae Type Hoa Binh Prov., Lac Son Distr., Tu Do Municipality, Mon village, around point 20°25’29”N 105°19’36”E. Primary broadleaved forest on steep rocky slopes of remnant mountain composed with solid crystalline highly eroded limestone at elevation about 500-700 m a.s.l. Terrestrial herb in shady place. Not rare. CPC 1566a/sp.1 25 March 2011. Coll.: L.Averyanov, T.Maisak, L.Osinovetz et al. Fl. under cultivation at December 2012. Perianth yellowish with dirty violet marking, dark dirty violet inside, stigma white to yellowish, filaments white, anthers light yellowish. Photos. Averyanov L.

FLORA OF VIETNAM Aspidistra heterocarpa Aver., Tillich & V.T. Pham var. echinata T.A.Le, Aver. & Tillich Thua Thien - Hue province, Nam Dong district, Thuong Quang commune, around point 16°07’28.5”N 107°35’52.9”E, primary and secondary evergreen broad-leaved lowland forest at elevation 500-550 m a.s.l., terrestrial herb on shady reclined slope, not common, 10 November 2017, Le Tuan Anh, Nguyen Thi Lien Thuong, LTA101. d-EXSICCATES OF VIETNAMESE FLORA 0295/LTA101 © Le Tuan Anh & L.Averyanov.

FLORA OF VIETNAM Convallariaceae Aspidistra coccigera Aver. et H.-J. Tillich sp.nov. Vietnam, Quang Binh Prov., Minh Hoa Distr., Dan Hoa municipality, Vietnam-Laotian border, around Cha Lo border entrance, point 17º40’52’’N 105º46’00’’E. Broad-leaved primary forest on very steep slopes and along shale ridge at elevation 500-750 m a.s.l. Terrestrial herb on very steep rocky shady slope. Flowers white, tips of petals dark violet-black, stigma brightly red. Locally common. HAL 11817 22 April 2008 photo Coll.: L.Averyanov, P.K.Loc, N.T.Vinh, N.S.Khang

Aspidistra atrata Aver., Tillich & Q.H.Bui⁣

FLORA OF LAOS Convallariaceae Tupistra albiflora K.Larsen Vientiane Prov., Vang Vieng Distr., Phol Xai village, about 8 km to the W of Vang Vieng town, Phar Poon Mt., around point 18°56’11.5”N 102°20’05.0”E. Dry broad-leaved primary and secondary evergreen forest on very steep rocky slopes and on vertical cliffs of remnant highly eroded rocky limestone mountains composed by solid crystalline limestone at elevation 300-900 m. Terrestrial herb with erect stem to 0.3 m tall in shady place. Very common at elevation 700-800 m. LA-VN 431 15 March 2013 Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, P.V.The, S.Lorphengsy. (Fl. 14 Nov. 2016, LA-VN 431 / garden no 13009. Flowers white, tepal apices violet or lilac, anthers pale light yellowish, inflorescence rachis pinkish. Flowers with strange strong smell like chlorophorm). Flowered 6 Dec. 2017 LA-VN 431 / garden no 13009 Photos L. Averyanov.

FLORA OF VIETNAM Convallariaceae Tupistra wattii Dien Bien Prov., Tua Chua Distr., Sin Chai Municipality, around point 22°03’38”N 103°19’56”E. Primary humid evergreen broad-leaved forest on very steep rocky slopes and on tops of remnant mountain composed with highly eroded marble-like solid crystalline limestone at elevation 1350-1500 m a.s.l. Terrestrial and lithophytic herb to 0.5 m tall on shady rocky slope. Flowers light green. Very common. CPC 900 13 December 2010 Coll.: L.Averyanov, P.K.Loc, P.V.The, N.T.Vinh.

FLORA OF VIETNAM. Fam. Convallariaceae. Tupistra wattii (C.B. Clarke) Hook.f. ???. Northern Vietnam, Lao Cai Prov., Van Ban Distr., to the west of Nam. Xay village (around point 21°58'09"N, 104°00'55"E). Primary. broad-leaved evergreen montane forest with bamboo along tops of. granite/quartzite ridge at elev. 1800 m a.s.l. Creeping terrestrial. herb on open wet slope. Flowers greenish to dull orange. Not common. No HAL 2135   07 March 2002. . Collectors: L.Averyanov, P.K.Loc, D.T.Doan (FFI Project).

FLORA OF VIETNAM Fam. Convallariaceae Tupistra sp. N. Vietnam, Prov. Cao Bang, Distr. Tra Linh, Subdistr. Quoc Toan, evergreen forest on slopes of remnant limestone mountains near Thang Heng lake in environs of Thang Hen and Lung Tao villages at 600-700 m alt. Herb with leaves up to 1.5 m lg. Flowers black. Very common at 600-700 m alt. No VH 2463 04.01.1996 International Botanical Expedition of the U.S.A. National Geographic Society (grant No 5094-93) Participants: L. Averyanov, N.T. Ban, A. Budantzev, L. Budantzev, N.T. Hiep, D.D. Huyen, P.K. Loc, G. Yakovlev

FLORA OF VIETNAM. Fam. Convallariaceae. Tupistra sp. N. Vietnam, prov. Lai Chau, distr. Sin Ho, municipality Ta Ngau about. 30 km of Lai Chau town to the N (N 22°18' E 103°15'). Secondary open. wet mossy evergreen broadleaved forest with remnants of primary. elements on rocky steep slope of remnant karst limestone ridge at. elev. about 1650 m. Terrestrial and lithophyte forb up to 0.6 m hg on. shady steep slope. Very common. No NTH 2789   25 May 1999. International Botanical Expedition of the U.S.A. National Geographic. Society (grant No 6300-98). . Collectors: N.T.Hiep, P.H.Hoang, Averyanov L.

FLORA OF LAOS Convallariaceae Tupistra muricata(Gagnep.) N.Tanaka Vientiane Prov., Vang Vieng Distr., Nathong village, about 5 km to the W of Vang Vieng town, Tham Kang Mt., around point 18°55’51.4”N 102°23’50.6”E. Dry broad-leaved primary and secondary evergreen forest on very steep rocky slopes and on vertical cliffs of remnant highly eroded rocky limestone mountains composed by solid crystalline limestone at elevation 300-400 m. Terrestrial herb with erect stem to 5 cm tall on shady slope. Leaves uniform green, to 1 m long, petiole 30-50 cm long, leaf blade elliptic to 50 cm long 15 cm wide. Rare. LA-VN 410a 14 March 2013 (living only) Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, P.V.The, S.Lorphengsy Fl. in cult. in October – November 2015. Flowers with chloroform smell, white, tepals violet at apices.

FLORA OF LAOS Convallariaceae Tupistra muricata (Gagnep.) N.Tanaka Luang Prabang prov., Ngoy distr., Nong Khiew village, Pha Deng Mountain around point 20°34'14.3''N 102°37'31.8''E. Secondary semideciduous and evergreen dry forest on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. 700-825 m a.s.l. Terrestrial herb on steep rocky shady slope. Fruits dirty violet. Not common. LA-VN 1904 31 March 2017 Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.Maisak, Vichith Lamxay, Koudkeo Phommachanh, Phizar Sayalath, Keooudone Souvannakhoummane.

FLORA OF VIETNAM Convallariaceae Aspidistra lubae Aver. et Tillich var. lancifolia Aver. et Tillich Type Hoa Binh Prov., Lac Son Distr., Tu Do Municipality, Mon village, around point 20°25’29”N 105°19’36”E. Primary broadleaved forest on steep rocky slopes of remnant mountain composed with solid crystalline highly eroded limestone at elevation about 500-700 m a.s.l. Terrestrial herb in shady place. Not rare. CPC 1566a/sp.4 25 March 2011. Coll.: L.Averyanov, T.Maisak, L.Osinovetz et al. Fl. under cultivation at January 2013. Perianth white with dirty purple-violet marking, dark dirty violet inside, stigma white to light yellowish, filaments white, anthers light yellowish. Photos. Averyanov L.

FLORA OF LAOS

FLORA OF VIETNAM Convallariaceae Aspidistra coccigera Aver. et H.-J. Tillich sp.nov. Vietnam, Quang Binh Prov., Minh Hoa Distr., Dan Hoa municipality, Vietnam-Laotian border, around Cha Lo border entrance, point 17º40’52’’N 105º46’00’’E. Broad-leaved primary forest on very steep slopes and along shale ridge at elevation 500-750 m a.s.l. Terrestrial herb on very steep rocky shady slope. Flowers white, tips of petals dark violet-black, stigma brightly red. Locally common. HAL 11817 22 April 2008 photo Coll.: L.Averyanov, P.K.Loc, N.T.Vinh, N.S.Khang

FLORA OF VIETNAM

All drawings © L. Averyanov

FLORA OF VIETNAM. Convallariaceae. Tupistra sp. Dien Bien Prov., Tua Chua Distr., Tua Thang Municipality, Ta Xi Lang village, around point 21°57'17"N 103°25'31"E. Highly degraded primary and secondary evergreen broadleaved forest on very steep slopes and on tops of remnant mountains composed with solid limestone at elevation 850-950 m a.s.l. Lithophytic herb to 20 cm tall on shady mossy rocks. Flowers green to yellow. Occasional. CPC 2284 13 April 2011. Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, N.T.Vinh.

FLORA OF VIETNAM. Fam. Convallariaceae. Tupistra sp. N. Vietnam, prov. Cao Bang, distr. Bao Lac, municipality Dinh Phung,. about 5 km to NW from Yen Lac village (N 22°44' E 105°50'), about 50. km to NWW from Cao Bang town. Old secondary closed broadleaved wet. cloud forest on steep slopes and cliffs along top of Nam Linh. limestone ridge at 1500-1580 m alt. Epilite and terrestrial herb up to. 0.6 m hg. Flowers green. Common. No CBL 468   19 Nov. 1998. International Botanical Expedition of the U.S.A. National Geographic. Society (grant No 6300-98). . Collectors: Averyanov L., P.K.Loc, N.X.Tam.

Tupistra sp.

FLORA OF VIETNAM. Convallariaceae. Campylandra theana Aver., sp.nov. Paratype. Quang Binh Prov., Tuyen Hoa Distr., Lam Hoa municipality, Hung village, around point 17°55'41"N 105°49'43"E. Highly degraded primary and secondary evergreen broadleaved forest on very steep rocky slopes of remnant limestone hills composed with highly eroded marble-like crystalline limestone at elev. 200-300 m a.s.l. Lithophytic herb on vertical shady cliff. Flowers dark dirty purple-violet, anthers dull yellow. Not common. CPC 2616 1 May 2011 Photos. Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, P.V.The, N.T.Vinh.

Текст этикетки FLORA OF LAOS Convallariaceae Tupistra muricata (Gagnep.) N.Tanaka Vientiane Prov., Kasi Distr., Tam Tai village, about 10 km to the NW of Kasi town (19°12'57"N 102°14'59"E). Remnants of primary and secondary broad-leaved evergreen forest on very steep rocky slopes of remnant mountain composed with highly eroded solid crystalline limestone at elevation 500-650 m. Terrestrial herb with erect thick stem to 0.5 m tall on shady rocky slope. Old flowers almost black. Not rare. LA-VN 756 24 March 2013 Coll.: L.Averyanov, N.S.Khang, S.Lorphengsy Flowered in October 2014. Flowers pure white. Photos.

FLORA OF LAOS Convallariaceae Tupistra muricata(Gagnep.) N.Tanaka Vientiane Prov., Vang Vieng Distr., Nathong village, about 5 km to the W of Vang Vieng town, Tham Kang Mt., around point 18°55°51.4°N 102°23°50.6°E. Dry broad-leaved primary and secondary evergreen forest on very steep rocky slopes and on vertical cliffs of remnant highly eroded rocky limestone mountains composed by solid crystalline limestone at elevation 300-400 m. Terrestrial herb with erect stem to 5 cm tall on shady slope. Leaves uniform green, to 1 m long, petiole 30-50 cm long, leaf blade elliptic to 50 cm long 15 cm wide. Rare. LA-VN 410a 14 March 2013 (living only) Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, P.V.The, S.Lorphengsy Fl. in cult. in October ° November 2015. Flowers with chloroform smell, white, tepals violet at apices.

FLORA OF LAOS. Convallariaceae. Tupistra muricata (Gagnep.) N.Tanaka. Vientiane prov., Kasi distr., Si Sang Vone village, around point 19°06'10.9"N 102°10'46.1"E. Secondary broad-leaved evergreen dry forest on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. 600-700 m a.s.l. Terrestrial and lithophytic herb on shady steep rocky slope. Locally common. Fruits globular, dirty dark green. Common. LA-VN 1786 28 March 2017. Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.Maisak, Khamla Inkhavilay, Hongthong Southyvohanh, Pinmany Sisavath, Chanthala Cheuaysamphanh.