Herbarium LE - Specimens

FLORA OF LAOS Convallariaceae Aspidistra sp. Vientiane prov., Kasi distr., between San and Tham Tai villages, around point 19°17'47.9''N 102°08'58.9''E. Secondary broad-leaved evergreen dry forest on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. 900-1000 m a.s.l. Creeping terrestrial herb with horizontal rhizome on shady rocky slope. Fruits green to dirty violet. Locally common. LA-VN 1682 25 March 2017 Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.Maisak, Khamla Inkhavilay, Hongthong Southyvohanh, Pinmany Sisavath, Chanthala Cheuaysamphanh.

FLORA OF LAOS Convallariaceae Aspidistra sp. Kham Mouan Prov., Bounlapha Distr., Thong Sam village, Hinnam No Protected Area, Pu Pha Song Mt. composed by sandstone, around point 17°35’13.2”N 105°47’49.8”E. Primary evergreen broad-leaved and mixed forest on W macroslope at elevation from 500 m to the mountain top elevated to 1425 m. Terrestrial herb on shady slopes at elevation 600-1000 m. Leaves uniform green or with unclear yellowish spotting. Fruit green, globose 1 cm in diam. Very common. LA-VN 286 09 March 2013 Coll.: L.Averyanov, N.T.Hiep, V.Lamxay, N.S.Khang, P.V.The, S.Lorphengsy

FLORA OF VIETNAM Convallariaceae Aspidistra subrotata Son La Prov., Van Ho Distr., Chieng Xuan Municipality, Co Hong village, territory of Xuan Nha natural reserve, Pha Luong Mountain. Remnants of primary and secondary broad-leaved evergreen forest on very steep mountain slopes composed by brown sandstone at elevation 700-1000 m a.s.l. around point 20°42’14.2’’N 104°43’53.9’’E. Terrestrial herb with erect stem to 10 cm tall on steep humid slope. Flowers white with finest dirty purple marks. Locally very common. 12 November 2013 CPC 6963 Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, N.D.Thang, L.D.Qui

FLORA OF LAOS Convallariaceae Aspidistra subrotata Y.Wan & C.C.Huang Kham Mouan Prov., Bounlapha Distr., Thong Sam village, Hinnam No Protected Area, Pu Pha Song Mt. composed by sandstone, around point 17°35’13.2”N 105°47’49.8”E. Primary evergreen broad-leaved and mixed forest on W macroslope at elevation from 500 m to the mountain top elevated to 1425 m. Terrestrial herb on shady slopes at elevation 600-1000 m. Leaves uniform green or with few unclear yellowish spotting. Very common. LA-VN 287 09 March 2013 Coll.: L.Averyanov, N.T.Hiep, V.Lamxay, N.S.Khang, P.V.The, S.Lorphengsy Fl. In cult. 13 November 2015 (LA-VN 287/1303), tepals dirty brown-purplish, stigma white with small purple speckles. Photos. Averyanov L.

Cao Bang Prov., Bao Lac Distr., Dinh Phung municipality, Ban Meo village. Secondary broad-leaved evergreen forest on steep stream slope composed by shale stratified limestone at elevation 800-900 m a.s.l. around point 22°44’32.0’’N 105°46’29.1’’E. Terrestrial creeping herb in shady rocky stream valley. Leaves uniform green or with dull yellow spots. Locally common. 24 November 2014 CPC 7654 Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz

FLORA OF VIETNAM Aspidistra subrotata Y.Wan & C.C.Huang Hoa Binh Prov., Lac Son Distr., Tu Do Municipality, Mon village, around point 20°25’29”N 105°19’36”E. Primary broadleaved forest on steep rocky slopes of remnant mountain composed with solid crystalline highly eroded limestone at elevation about 500-700 m a.s.l. Terrestrial herb in shady place. Not rare. CPC 1566c (“big leaf, big flower”) 25 March 2011. Coll.: L.Averyanov, T.Maisak, L.Osinovetz et al. Fl. under cultivation at November 2015. Photos.

All photos © - L. Averyanov

FLORA OF VIETNAM

FLORA OF VIETNAM. Convallariaceae. Aspidistra subrotata Y.Wan & C.C.Huang?. Son La Prov., Van Ho Distr., Chieng Xuan Municipality, Co Hong village, territory of Xuan Nha natural reserve, Pha Luong Mountain. Remnants of primary and secondary broad-leaved evergreen forest on very steep mountain slopes composed by brown sandstone at elevation 700-1000 m a.s.l. around point 20°42'14.2"N 104°43'53.9"E. Terrestrial herb with erect stem to 10 cm tall on steep humid slope. Locally very common. Leaves sometimes with unclear yellowish spots. 12 November 2013 CPC 6962. Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, N.D.Thang, L.D.Qui

FLORA OF VIETNAM Fam. Convallariaceae Aspidistra nikolai Aver. et Tillich Type Thua Thien – Hue Prov., Nam Dong Distr., Thuong Lo Municipality, along Cha Mang stream, around point 16°07’52’’N 107°43’49’’E. Primary evergreen broad-leaved lowland forest on very steep rocky stream slope composed with stratified shale at elev. 400-450 m a.s.l. Terrestrial herb on steep shady rocky slope. Flowers black-violet to black. Locally very common. HAL 11041 9 Apr. 2007. Photo. Coll.: L.Averyanov, N.S.Khang, A.Averyanova, N.D.Phuong, L.V.Hung. d-EXSICCATES OF VIETNAMESE FLORA 0107/HAL 11041 (HN, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VIETNAM. Aspidistra sp. S. Vietnam, Prov. Lam Dong, 31 km to NE from Dalat city, distr. Lac. Duong, municipalite Da Chay (N 12° 08', E 108° 39'). Closed primary. broadleaved mt forest on NW macroslope of Bi Dup mt system at. 1550-1700 m alt. Common herb in shadow places. No VH 2769  17.03.1997. International Botanical Expedition of the U.S.A. National Geographic. Society (grant No 5803-96). . Collectors: L. Averyanov, N.Q. Binh, N.V. Duy, P.K. Loc

FLORA OF VIETNAM. Aspidistra sp. S. Vietnam, Prov. Lam Dong, 14 km to NNW from Dalat city (N 12° 05', E 108° 22'). Primary closed mixed forest with Pinus kesiya and. P. krempfii at 1740-1760 m alt. Terrestrial herb on steep slopes along stream. Rare. No VH 2498  11.03.1997. International Botanical Expedition of the U.S.A. National Geographic. Society (grant No 5803-96). . Collectors: L. Averyanov, N.Q. Binh, N.V. Duy, P.K. Loc

FLORA OF VIETNAM Aspidistra hainanensis W.Y. Chun et F.C. How Lam Dong province., Bao Loc provincial city area, about 18 km to NNW of Bao Loc City, Dasara waterfall, evergreen forest with bamboo at elevation 700–800 m a.s.l. along flat stream valley and on hill slopes around waterfall. 10 January 2017, Nguyen Danh Duc & Nguyen Hoang Tuan, 110. d-EXSICCATES OF VIETNAMESE FLORA 0276/N.D.Duc, N.H.Tuan s.n. © L.Averyanov

Cult. in h. b. Petropol 1834

Aspidistra lurida Ker Gawl.

FLORA OF VIETNAM. Convalariaceae. Aspidistra coccigera Aver. et Tillich. Quang Binh Prov., Minh Hoa Distr., Thuong Hoa municipality, environs of Mo O O O village, around point 17°38'00.4"N 105°55'57.9"E, Kalap Mountain. Primary mixed and coniferous forest (with Dacrydium elatum and Dacrycarpus imbricatus) on very steep rocky slope and on rocky top of remnant mountain composed with highly eroded solid crystalline limestone at elevation 750-800 m a.s.l. Terrestrial herb on steep shady rocky slope near to mountain top. Tepals dark violet, stigma red. Locally common. CPC 3748 25 July 2011. Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Vinh.

no label

EAST BENGAL Herb Griffith. 5889

FLORA OF LAOS Convallariaceae Aspidistra subrotata Y.Wan & C.C.Huang = Aspidistra connata Tillich Saravan Prov., Sa Mouay Distr., to the SEE of Sa Mouay town, elevation from Ta Oy village (700 m elev.) to mountain ridge with highest point at 1350 m around 16°08’47.0”N 106°52’35.4”E. Primary evergreen humid mixed and coniferous forest on slopes composed with silicate rocks. Terrestrial herb in shady places along ridge slopes at elevation 1000-1200 m. Leaves elliptic, 30 cm long, 6 cm wide, stalk 50 cm long, finely yellowish spotted. Not common. LA-VN 199a 04 March 2013 (Living sample only) Coll.: L.Averyanov, N.T.Hiep, V.Lamxay, N.S.Khang, P.V.The, S.Lorphengsy Fl. 22 Nov. 2016. Tepals and stigma white with many purple speckles. Photos.

FLORA OF VIETNAM Fam. Convallariaceae Aspidistra brachystyla Aver. et Tillich Type Vietnam, Nghe An Prov., Tuong Duong Distr., Tam Quang Municipality, Tung Huong village, along Khe Mat stream around point 19º03’15’’N 104º36’19’’ at elevation about 500-560 m a.s.l. Lowland closed secondary and primary old evergreen forest on steep slopes along rocky limestone ridge and on rocky tops composed with solid limestone. Terrestrial herb along steep rocky shady stream slopes. Flowers light yellow. Fruits black. Not rare. HLF 6705 18 March 2007. Photo. Coll. Averyanov L., P.K.Loc, N.V.Trai, N.T.Vinh, N.D.Phuong, Averyanova A. d-EXSICCATES OF VIETNAMESE FLORA 0096/HLF6705 (HN, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

Cult. in hort. bot. Petropolit. 1834

Tupistra nutans Wall.⁣

All photos © L. Averyanov

Tupistra squalida Ker Gawl.

FLORA OF LAOS Fam. Begoniaceae Begonia viscosa Aver., sp.nov. Type Vientiane Prov., Vang Vieng Distr., vicinity of Vang Vieng town, around point 18°54’58”N 102°24’52”E. Secondary broadleaved evergreen dry forest on very steep rocky slopes and on the top of remnant mountain composed with highly eroded crystalline marble-like limestone at elevation 400-450 m a.s.l. Lithophytic undershrub with erect shoots to 2 m tall in crevices of open vertical cliffs. Flowers white, sometimes with light punk tint, anthers and stigmas yellow. Leaves above light green to dull gray or gray-brownish with palmate whitish marks, uniform dull purple below, petioles green to purple. Locally common. CPC 2438 24 April 2011 Photos. Coll.: L.Averyanov, P.V.The. d-EXSICCATES OF VIETNAMESE FLORA 0187/CPC 2438 (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VIETNAM Begoniaceae Begonia rigidifolia Aver., sp.nov. Type Quang Binh Prov., Tuyen Hoa Distr., Lam Hoa municipality, Chuoi village, around point 17°56’51”N 105°49’18”E. Primary broadleaved evergreen dry forest on very steep slopes and on tops of rocky ridge composed with highly eroded crystalline limestone at elev. 200-250 m a.s.l. Lithophytic rosulate herb on shady vertical cliffs. Flowers light pinkish, anther and stigmas yellow, leaves dark to light green with irregular dirty brownish or dark green irregular palmate markings above, light green, nearly white below, with unclear dirty purple marks. Petioles dull brownish-purple. Not rare. CPC 2733 3 May 2011 Photos. Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, P.V.The, N.T.Vinh. d-EXSICCATES OF VIETNAMESE FLORA 0186/CPC 2733 (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VIETNAM Begoniaceae Begonia crassula Aver. sp.nov. Paratype Quang Binh Prov., Minh Hoa Distr., Thuong Hoa municipality, environs of Mo O O O village, around point 17°39’36.7”N 105°54’55.7”E. Remnants of short tall broad-leaved primary forest on rocky top of remnant mountain composed with highly eroded solid crystalline limestone at elevation about 612 m a.s.l. Lithophytic herb or undershrub to 1 m tall on exposed open rocks on rocky mountain top. Flowers light pink, anther and stigma yellow. Locally common. CPC 3775 26 July 2011 Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Vinh. d-EXSICCATES OF VIETNAMESE FLORA 0189/CPC 3775 (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VIETNAM Fam. Begoniaceae Begonia rubrosetosa Aver. Epitype Thua Thien – Hue Prov., Nam Dong Distr., Thuong Lo Municipality, along Cha Mang stream, around point 16°07’35’’N 107°44’53’’E. Remnants of primary evergreen broad-leaved lowland forest on very steep hill slopes composed with stratified shale and sandstone at elev. about 150 m a.s.l. Terrestrial and lithophytic herb on very steep rocky slope. Leaves uniform green with reddish nerves. Flowers red, anther and stigmas yellow. Locally very common. HAL 10883 4 Apr. 2007. Photo. Coll.: L.Averyanov, Phan Ke Loc, A.Averyanova, N.T.Vinh, N.D.Phuong, L.V.Hung. Det. N.Q.Hieu d-EXSICCATES OF VIETNAMESE FLORA 0104/HAL 10883 (HN, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VIETNAM Fam. Begoniaceae Begonia minuscula Aver., sp.nov. Type Quang Tri Prov., Huong Hoa Distr., Huong Viet municipality around point 16°51’06”N 106°34’38”E. Partially destroyed primary broadleaved evergreen forest on very steep rocky slopes of remnant hills composed with solid highly eroded crystalline limestone at elev. about 600-700 m a.s.l. Lithophytic herb on vertical tall cliff. Leaves uniform green, pinkish or brown, sometimes with whitish spots. Not common. CPC 2838 7 May 2011 Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, P.V.The, N.T.Vinh. d-EXSICCATES OF VIETNAMESE FLORA 0184/HAL 2838 (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VIETNAM Begoniaceae Begonia minuscula Aver. Paratype Vietnam, Quang Tri Prov., Huong Hoa Distr., Huong Viet Municipality, to N of Sa Mui pass, around point 16º51’14’’N 106º34’13’’E at elevation 550-650 m a.s.l. Remnants of primary broad-leaved evergreen lowland forest on rocky remnant highly eroded solid marble-like limestone hills and low mountains with very steep slopes and vertical cliffs. Lithophytic creeping herb on vertical shady cliffs. Leaves pure green to pink-brown. Very common. HLF 5910 31 March 2006. Coll. N.T.Hiep, L.Averyanov, P.K.Loc, P.V.The, N.S.Khang, P.H.Duc, L.T.Son, S.Bounphanmy, T.K.Ratana, O.Sonliya, T.Q.Dang, N.V.Hien, H.T.N.Trang, L.T.Chau. Grant from Henry Luce Foundation.

FLORA OF VIETNAM Begoniaceae Begonia crassula Aver. sp.nov. Paratype Quang Binh Prov., Minh Hoa Distr., Thuong Hoa municipality, environs of Mo O O O village, around point 17°39’36.7”N 105°54’55.7”E. Remnants of short tall broad-leaved primary forest on rocky top of remnant mountain composed with highly eroded solid crystalline limestone at elevation about 612 m a.s.l. Lithophytic herb or undershrub to 1 m tall on exposed open rocks on rocky mountain top. Flowers light pink, anther and stigma yellow. Locally common. CPC 3775 26 July 2011 Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Vinh. d-EXSICCATES OF VIETNAMESE FLORA 0189/CPC 3775 (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VIETNAM Fam. Begoniaceae Begonia sp. N. Vietnam, Bac Can prov., Na Ri distr., Kim Hy municipality, vicinities of Lung Hin Con village (N 22º17' E 106º03'), 27 km to 55º of Bac Can city. Primary evergreen mixed mossy forest with Pseudotsuga brevifolia and bamboo along tops of karst remnant limestone ridge at elev. 700 m a.s.l. Adpressed lithophytic forb in crevices of shady verical cliffs. Leaves below violet, above deep green with irregular palmate white spot. Very common. Usual codominanthe of lithophytic forb communities. No NTH 3725 20 Oct. 1999 Collectors: N.T. Hiep, P.K. Loc, L. Averyanov.

FLORA OF VIETNAM Fam. Begoniaceae Begonia minuscula Aver., sp.nov. Type Quang Tri Prov., Huong Hoa Distr., Huong Viet municipality around point 16°51’06”N 106°34’38”E. Partially destroyed primary broadleaved evergreen forest on very steep rocky slopes of remnant hills composed with solid highly eroded crystalline limestone at elev. about 600-700 m a.s.l. Lithophytic herb on vertical tall cliff. Leaves uniform green, pinkish or brown, sometimes with whitish spots. Not common. CPC 2838 7 May 2011 Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, P.V.The, N.T.Vinh. d-EXSICCATES OF VIETNAMESE FLORA 0184/HAL 2838 (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VETNAM. Dregea sp.nov.?. Kon Tum province, Kon Pl°ng district, Hieu commune, around point 14°65'24"N 108°43'01"E, 1200°1210 m a.s.l., secondary evergreen broad-leaved forest, on basaltic soils. Terrestrial liana, 3 m long with white latex, flower light green, sometimes with dull white tint. Not common. 2 May 2015. Le Tuan Anh, Pham Doan Quoc Vuong, PVT 713.

FLORA OF VIETNAM Fam. Begoniaceae Begonia rubrosetosa Aver. Epitype Thua Thien – Hue Prov., Nam Dong Distr., Thuong Lo Municipality, along Cha Mang stream, around point 16°07’35’’N 107°44’53’’E. Remnants of primary evergreen broad-leaved lowland forest on very steep hill slopes composed with stratified shale and sandstone at elev. about 150 m a.s.l. Terrestrial and lithophytic herb on very steep rocky slope. Leaves uniform green with reddish nerves. Flowers red, anther and stigmas yellow. Locally very common. HAL 10883 4 Apr. 2007. Photo. Coll.: L.Averyanov, Phan Ke Loc, A.Averyanova, N.T.Vinh, N.D.Phuong, L.V.Hung. Det. N.Q.Hieu d-EXSICCATES OF VIETNAMESE FLORA 0104/HAL 10883 (HN, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VIETNAM Fam. Begoniaceae Begonia minuscula Aver., sp.nov. Type Quang Tri Prov., Huong Hoa Distr., Huong Viet municipality around point 16°51’06”N 106°34’38”E. Partially destroyed primary broadleaved evergreen forest on very steep rocky slopes of remnant hills composed with solid highly eroded crystalline limestone at elev. about 600-700 m a.s.l. Lithophytic herb on vertical tall cliff. Leaves uniform green, pinkish or brown, sometimes with whitish spots. Not common. CPC 2838 7 May 2011 Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, P.V.The, N.T.Vinh. d-EXSICCATES OF VIETNAMESE FLORA 0184/HAL 2838 (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VIETNAM Begoniaceae Begonia crassula Aver. sp.nov. Paratype Quang Binh Prov., Minh Hoa Distr., Thuong Hoa municipality, environs of Mo O O O village, around point 17°39’36.7”N 105°54’55.7”E. Remnants of short tall broad-leaved primary forest on rocky top of remnant mountain composed with highly eroded solid crystalline limestone at elevation about 612 m a.s.l. Lithophytic herb or undershrub to 1 m tall on exposed open rocks on rocky mountain top. Flowers light pink, anther and stigma yellow. Locally common. CPC 3775 26 July 2011 Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Vinh. d-EXSICCATES OF VIETNAMESE FLORA 0189/CPC 3775 (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc