Herbarium LE - Specimens

Plants of Dong Chau - Khe Nuoc Trong Nature Reserve (DCKNTNR)

Orchidaceae

Tainia penangiana Hook.f.

Quang Binh Province, Le Thuy District, Kim Thuy Municipality. Secondary evergreen broad-leaved forest with bamboo on very steep river slopes composed by shaly limestone at elevation about 100 m a.s.l. around point 17º4’29’’N 106º43’5’’E. Terrestrial creeping herb on shady rocky stream slope. Not common.

22 November 2024, Nguyen Huu Cuong, Phan Thanh Quyet, Duong Van Nam, L. Averyanov, T. Maisak, AL3696. Living plant only.

Flowered in culture on 26 January 2026, L. Averyanov, T. Maisak, AL3696.

Convallariaceae

Peliosanthes griffithii Baker?

Lam Dong Province, Bao Lam District. Primary humid evergreen broad-leaved and mixed forest with Pinus kesiya on granite at elevation about 1100 m a.s.l. Terrestrial herb with large dark green leaves to 1 m long.

30 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3534.

Flowered in cultivation on 26 January 2026, flowers light greenish, corona dirty violet L. Averyanov, T. Maisak AL3534

Orchidaceae

Vanda sp.

Kon Tum Province, Tu Mo Rong District,

26 May 2025, Truong Ba Vuong, BV 1970

Note: the species has somewhat triangular oblong lip,

apex with retuse small ovate-rounded lobules.

All photos © Truong Ba Vuong

[manu Turczaninovii]

In viciniis acidularum Pogromezensium 1829

legit cl. Basnin

Turcz.

FLORA OF LAOS

Polypodiaceae s.l.

Leptochilus sp.

Houphan Prov., Hem Distr., Nacoc village, Pu He mountain around point 20°10'36.7''N 103°11'39.2''E. Secondary broad-leaved evergreen open dry forest with bamboo on very steep rocky mountain slopes composed by shaly limestone at elev. 700-900 m a.s.l. Short creeping lithophytic fern on wet mossy shady rocks along small rocky stream. Not rare.

LA-VN 1365 12 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

Polypodiaceae s.l.

Colysis digitata Ching?

Ha Giang Prov., Bac Me Distr., Phieng Luong municipality, around Phieng Luong village. Primary broad-leaved humid evergreen forest on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation 1100-1150 m a.s.l. around point 22°38’07.1’’N 105°19’22.1’’E. Lithophytic creeping fern on very steep shady rocky slope near and along ridge edge. Very common.

13 November 2014 CPC 7423

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz

FLORA OF VIETNAM

Polypodiaceae s.l.

Asplenium filiforme Kaulf.

Cao Bang Prov., Bao Lac Distr., Dinh Phung municipality, Ban O village. Primary mixed and coniferous evergreen humid forest with (Pinus sp., Pseudotsuga sinensis, Podocarpus neriifolius) on very steep slopes and on mountain tops composed by solid highly eroded limestone at elevation 1200-1300 m a.s.l. around point 22°45’24.8’’N 105°46’47.9’’E. Lithophyte on mossy rocks in shady places on mountain tops. Very common.

24 November 2014 CPC 7634

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz

FLORA OF LAOS

Polypodiaceae s.l.

Leptochilus decurrens Blume

Houphan Prov., Hem Distr., Khon Ngua village, around point 19°55'53.2''N 103°16'24.2''E. Secondary broad-leaved evergreen open dry forest with bamboo on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. 700-950 m a.s.l. Short creeping lithophyte in shady places. Common.

LA-VN 1478 15 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham, Somchanh Bounphanmy.

FLORA OF VIETNAM

Polypodiaceae s.l.

Adiantum capillus-veneris L.

Cao Bang Prov., Bao Lam Distr., Yen Tho municipality, Cha Hai village, Phia Hai Mt. Secondary highly degraded broad-leaved evergreen forest on steep mountain slopes near mountain top at elevation 1300-1350 m a.s.l. around point 22°41’22.3’’N 105°34’43.5’’E. Lithophytic fern on vertical shady wet cliff around cave entrance. Not common.

27 November 2014 CPC 7665

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz

FLORA OF LAOS

Polypodiaceae s.l.

Pyrrosia nummulariifolia (Sw.) Ching.

Houphan Prov., Viengxay Distr., Xieng Luong village, around point 20°22'06.1''N 104°17'18.6''E. Primary and secondary broad-leaved evergreen dry forest on very steep rocky slopes of highly eroded mountains composed by solid marble-like limestone at elev. 600-700 m a.s.l.

Creeping lithophyte on shady mossy cliffs. Very common.

LA-VN 1158 8 April 2015 photos

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Somneuk Bouthamounty.

FLORA OF LAOS

Polypodiaceae s.l.

Aglaomorpha coronans (Wall. ex Mett.) Copel.

Houphan Prov., Hem Distr., Nacoc village, Puphadeng mountain, around point 20°09'52.6''N 103°10'19.4''E. Primary and secondary broad-leaved evergreen open dry forest on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. 1000-1200 m a.s.l. Epiphyte and lithophyte on rocky slope near mountain top. Very common.

LA-VN 1440 12 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

FLORA OF LAOS

Polypodiaceae s.l.

Asplenium nidus L.

Houphan Prov., Hem Distr., Khon Ngua village, around point 19°55'53.2''N 103°16'24.2''E. Secondary broad-leaved evergreen open dry forest with bamboo on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. 700-950 m a.s.l.

Epiphyte on tall old trees. Common.

LA-VN 1497 15 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham, Somchanh Bounphanmy.

FLORA OF LAOS

Polypodiaceae s.l.

Asplenium antrophyoides Christ

Houphan Prov., Viengxay Distr., Xieng Luong village, around point 20°22'06.1''N 104°17'18.6''E. Primary and secondary broad-leaved evergreen dry forest on very steep rocky slopes of highly eroded mountains composed by solid marble-like limestone at elev. 600-700 m a.s.l.

Rosulate lithophyte on shady cliff near mountain top. Very common.

LA-VN 1193 8 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Somneuk Bouthamounty.

FLORA OF LAOS

Polypodiaceae s.l.

Pyrrosia nummulariifolia (Sw.) Ching

Houphan Prov., Hem Distr., Nacoc village, Puphadeng mountain, around point 20°09'52.6''N 103°10'19.4''E. Primary and secondary broad-leaved evergreen open dry forest on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. 1000-1200 m a.s.l. Creeping epiphyte and lithophyte on mountain top. Very common.

LA-VN 1436 12 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

Polypodiaceae s.l.

Pyrrosia nummulariifolia (Sw.) Ching.

Houphan Prov., Viengxay Distr., Vieng Xai village, around point 20°23'41.3''N 104°13'44.5''E. Primary and secondary broad-leaved evergreen dry forest on very steep rocky slopes of highly eroded mountains composed by solid marble-like limestone at elev. 800-950 m a.s.l.

Creeping lithophyte on shady mossy cliffs. Very common.

LA-VN 1139 7 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Somneuk Bouthamounty.

FLORA OF LAOS

Polypodiaceae s.l.

Asplenium antrophyoides Christ

Houphan Prov., Viengxay Distr., Vieng Xai village, around point 20°23'41.3''N 104°13'44.5''E. Primary and secondary broad-leaved evergreen dry forest on very steep rocky slopes of highly eroded mountains composed by solid marble-like limestone at elev. 800-950 m a.s.l.

Lithophytic rosulate fern on vertical shady cliffs. Very common.

LA-VN 1129 7 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Somneuk Bouthamounty.

FLORA OF LAOS

Polypodiaceae s.l.

Colysis digitata Ching

Xiangkhouang Prov., Peak Distr., Oran village, around point 19°38’53”N 103°23’12”E. Primary and secondary broad-leaved evergreen dry forest on shale sandstone slopes and along ridge edge at elev. about 1750-1850 m a.s.l. Short creeping lithophyte in shady place. Very common.

LA-VN 890 2 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

FLORA OF VIETNAM

Polypodiaceae s.l.

Ha Giang Prov., Bac Me Distr., Phieng Luong municipality, around Phieng Luong village. Primary broad-leaved humid evergreen forest on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation 1100-1150 m a.s.l. around point 22°38’07.1’’N 105°19’22.1’’E. Epiphytic fern on mossy trees on very steep shady rocky slope near and along ridge edge. Common.

13 November 2014 CPC 7426

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz

FLORA OF VIETNAM

Polypodiaceae s.l.

Lindsaea orbiculata (Lam.) Mett. ex Kuhn

Cao Bang Prov., Bao Lam Distr., Yen Tho municipality, Cha Hai village, Phia Hai Mt. Secondary highly degraded broad-leaved evergreen forest on steep mountain slopes near mountain top at elevation 1300-1350 m a.s.l. around point 22°41’22.3’’N 105°34’43.5’’E. Terrestrial fern on steep clayey slope along rtail. Common.

27 November 2014 CPC 7676

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz

FLORA OF LAOS

Polypodiaceae s.l.

Leptochilus sp.

Houphan Prov., Hem Distr., Na Puok village, Pu Tham Nhon mountain around point 20°10'00.0''N 103°24'37.0''E. Primary and secondary broad-leaved evergreen dry forest on very steep rocky mountain slopes composed by solid marble-like highly eroded limestone at elev. 1050-1150 m a.s.l. Lithophytic creeping fern on mossy rocks. Occasional.

LA-VN 1345 11 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

FLORA OF LAOS

Polypodiaceae s.l.

Colysis sp.

Houphan Prov., Hem Distr., Nacoc village, Pu He mountain around point 20°10'36.7''N 103°11'39.2''E. Secondary broad-leaved evergreen open dry forest with bamboo on very steep rocky mountain slopes composed by shaly limestone at elev. 700-900 m a.s.l. Terrestrial fern along shady ridge edge. Locally very common.

LA-VN 1372 12 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

FLORA OF LAOS

Polypodiaceae s.l.

Asplenium antrophyoides Christ

Houphan Prov., Hem Distr., Khon Ngua village, around point 19°55'53.2''N 103°16'24.2''E. Secondary broad-leaved evergreen open dry forest with bamboo on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. 700-950 m a.s.l.

Rosulate lithophyte on very steep rocky slope and on shady cliffs. Very common.

LA-VN 1493 15 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham, Somchanh Bounphanmy.

FLORA OF VIETNAM

Polypodiaceae s.l.

Onichium sp.?

Cao Bang Prov., Bao Lac Distr., Dinh Phung municipality, Ban O village. Primary mixed and coniferous evergreen humid forest with (Pinus sp., Pseudotsuga sinensis, Podocarpus neriifolius) on very steep slopes and on mountain tops composed by solid highly eroded limestone at elevation 1200-1300 m a.s.l. around point 22°45’24.8’’N 105°46’47.9’’E. Lithophyte on mossy rocks in shady places on mountain tops. Common.

24 November 2014 CPC 7633

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz

FLORA OF LAOS

Polypodiaceae s.l.

Leptochilus sp.

Houphan Prov., Hem Distr., Khon Ngua village, around point 19°55'53.2''N 103°16'24.2''E. Secondary broad-leaved evergreen open dry forest on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. 700-950 m a.s.l. Short creeping epiphyte on shady mountain slope. Not common.

LA-VN 1470 15 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham, Somchanh Bounphanmy.

FLORA OF LAOS

Polypodiaceae s.l.

Lepisorus sp.?

Xiangkhouang Prov., Peak Distr., Oran village, around point 19°38’53”N 103°23’12”E. Primary and secondary broad-leaved evergreen dry forest on shale sandstone slopes and along ridge edge at elev. about 1750-1850 m a.s.l. Creeping epiphytic vine 3-5 m long in shady place. Common.

LA-VN 889 2 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

FLORA OF LAOS

Polypodiaceae s.l.

Lepisorus sp.?

Xiangkhouang Prov., Peak Distr., Oran village, around point 19°38’53”N 103°23’12”E. Primary and secondary broad-leaved evergreen dry forest on shale sandstone slopes and along ridge edge at elev. about 1750-1850 m a.s.l. Creeping epiphytic vine 2-4 m long in shady place. Common.

LA-VN 920 2 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

FLORA OF LAOS

Polypodiaceae s.l.

Adiantum sp.

Xiangkhouang Prov., Kham Distr., Huad village, Phou tat Vinh mt., around point 19°32'31.6'N 103°39'40.4'E. Primary and secondary broad-leaved evergreen and semideciduous dry forest on very steep rocky slopes of limestone highly eroded mountains composed by solid marble like limestone at elev. 1200-1400 m a.s.l. Terrestrial fern in shady place. Locally common.

LA-VN 1077 4 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

FLORA OF VIETNAM

Polypodiaceae

Gen. Sp.

Ha Giang Prov., Vi Xuyen Distr., Tung Ba municipality, Khau Ca NR. Primary broad-leaved humid evergreen forest on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation 700-1150 m a.s.l. around point 22°50’43.9’’N 105°07’18.5’’E. Epiphytic herb, common.

09 May 2015 CPC 7812

Coll.: N.Q. Hieu, N.T.Hiep, T.B. Ngan, C.Q. Ngan, D.K. Harder, C. P. Tuy, G.C.Pao

FLORA OF VIETNAM

Polypodiaceae

Gen. sp.

Ha Giang Prov., Vi Xuyen Distr., Tung Ba municipality, Khau Ca NR. Primary broad-leaved humid evergreen forest on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation 700-1150 m a.s.l. around point 22°50’43.9’’N 105°07’18.5’’E. ( Khau Ca forest station to travel b,a,c to Minh Son to Lung village to Khau Ca Foretbstation) Epiphytic herb. occasional.

10 May 2015 CPC 7879

Coll.: N.Q. Hieu, N.T.Hiep, T.B. Ngan, C.Q. Ngan, D.K. Harder, C. P. Tuy, G.C.Pao

FLORA OF LAOS

Polypodiaceae s.l.

Adiantum sp.

Houphan Prov., Hem Distr., Na Puok village, Pu Tham Nhon mountain around point 20°10'00.0''N 103°24'37.0''E. Primary and secondary broad-leaved evergreen dry forest on very steep rocky mountain slopes composed by solid marble-like highly eroded limestone at elev. 1050-1150 m a.s.l. Terrestrial fern on steep shady slope. Common.

LA-VN 1312 11 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

FLORA OF LAOS

Polypodiaceae s.l.

Adiantum sp.

Houphan Prov., Viengxay Distr., Xieng Luong village, around point 20°22'06.1''N 104°17'18.6''E. Primary and secondary broad-leaved evergreen dry forest on very steep rocky slopes of highly eroded mountains composed by solid marble-like limestone at elev. 600-700 m a.s.l.

Terrestrial rosulate fern on shady rocky slope. Locally common.

LA-VN 1205 8 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Somneuk Bouthamounty.

FLORA OF LAOS

Polypodiaceae s.l.

Davallia sp.

Houphan Prov., Xon Distr., Nong Hua Pa Noi village, around point 20°13'02.1''N 103°24'23.3''E. Primary and secondary broad-leaved evergreen forest on very steep rocky mountain slopes composed by shaly limestone at elev. 1100-1200 m a.s.l. Creeping epiphyte on tall mossy tree along ridge edge. Locally very common.

LA-VN 1275 11 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

FLORA OF LAOS

Polypodiaceae s.l.

Pseudodrynaria sp.

Houphan Prov., Xon Distr., Nong Hua Pa Noi village, around point 20°13'02.1''N 103°24'23.3''E. Primary and secondary broad-leaved evergreen forest on very steep rocky mountain slopes composed by shaly limestone at elev. 1100-1200 m a.s.l. Short creeping epiphyte on tall trees. Very common.

LA-VN 1228 11 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

FLORA OF LAOS

Polypodiaceae s.l.

Belvisia sp.

Houphan Prov., Hem Distr., Na Puok village, Pu Tham Nhon mountain around point 20°10'00.0''N 103°24'37.0''E. Primary and secondary broad-leaved evergreen dry forest on very steep rocky mountain slopes composed by solid marble-like highly eroded limestone at elev. 1050-1150 m a.s.l. Epiphyte on old mossy trees among shady mossy rocks on mountain top. Common.

LA-VN 1339 11 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

FLORA OF LAOS

Polypodiaceae s.l.

Asplenium rupestre Hope

Houphan Prov., Hem Distr., Na Puok village, Pu Tham Nhon mountain around point 20°10'00.0''N 103°24'37.0''E. Primary and secondary broad-leaved evergreen dry forest on very steep rocky mountain slopes composed by solid marble-like highly eroded limestone at elev. 1050-1150 m a.s.l. Terrestrial and epiphyte on old mossy trees among shady mossy rocks on mountain top. Common.

LA-VN 1338 11 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

Orchidaceae

Tainia penangiana Hook.f.

Dak Lak Province, Lak District, around Lieng Bok Sak Waterfall at elevation about 400 m a.s.l. Dry evergreen broad-leaved forest along rocky stream on granite. Terrestrial herb among rocks along stream. Occasional.

11 May 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3144.

Flowered in culture on 12 January 2026, L. Averyanov, T. Maisak, AL3144.

Acanthaceae

Gen. ?

Dak Lak Province, M’Drak District, W foothills of Chu Mu Mt., dry secondary and remnants of primary evergreen broad-leaved forest on granite on steep slopes along rocky river at elevation 600-650 m a.s.l. Terrestrial adpressed rosulate herb on soil cliff in shady place along stream. Flowers light blue violet. Not rare.

25 April 2025, L. Averyanov, N.V. Canh, T. Maisak, AL3944.