Herbarium LE - Specimens

FLORA OF LAOS

FLORA OF VIETNAM Fam. Liliaceae s.l. Convallariaceae Ophiopogon fruticulosus Aver., N.Tanaka et S.K.Nguyen Type Vietnam, Ha Giang Prov., Bac Me Distr., Phieng Luong Municipality, around Phieng Luong village about 4 km to SW of Phieng Luong Municipality headquarter 22º39’29’’N, 105º19’35’’E. Slightly logged primary closed evergreen broad-leaved forest on slopes of remnant mountains composed with solid highly eroded crystalline and partially shale limestone at elev. 1000-1200 m a.s.l. Undershrub 0.5-1.5 m tall on rocky top of ridge. Fruits deep blue. Locally very common. No HAL 6489 5 March 2005. Collectors: L.Averyanov, P.K.Loc, N.T.Vinh, A.Averyanova.

FLORA OF VIETNAM.

FLORA OF VIETNAM.

FLORA OF VIETNAM Convallariaceae Ophiopogon petraeus Aver. et N.Tanaka Type Lang Son Prov., Huu Lung Distr., Huu Lien Commune, Huu Lien Nature Reserve, around point 21°40’28”N, 106°22’34”E. Primary closed evergreen seasonal tropical mixed forests on rocky limestone at elevation 400-600 m. Lithophytic rosulate herb. Not rare. HAL 11220b 24 September 2007 Coll.: P.K.Loc, N.T.Vinh, H.V.Luat Flowered under cultivation at September-October 2013. Flowers white, anthers light yellow-green, pedicel white with violet tint. Leaves fleshy, at the base subsucculent, juicy, transversely-oval in section. Photos. L. Averyanov, HAL 11220b/1. d-EXSICCATES OF VIETNAMESE FLORA 0207/HAL 11220b (LE) © L.Averyanov

FLORA OF VIETNAM Fam. Liliaceae s.l. Convallariaceae Ophiopogon fruticulosus Aver., N.Tanaka et S.K.Nguyen Type Vietnam, Ha Giang Prov., Bac Me Distr., Phieng Luong Municipality, around Phieng Luong village about 4 km to SW of Phieng Luong Municipality headquarter 22º39’29’’N, 105º19’35’’E. Slightly logged primary closed evergreen broad-leaved forest on slopes of remnant mountains composed with solid highly eroded crystalline and partially shale limestone at elev. 1000-1200 m a.s.l. Undershrub 0.5-1.5 m tall on rocky top of ridge. Fruits deep blue. Locally very common. No HAL 6489 5 March 2005. Collectors: L.Averyanov, P.K.Loc, N.T.Vinh, A.Averyanova. Fl. In cult in November – December 2015. Flowers white with light violet tint. Photos. Averyanov L. Collected from cultivated plant grown from seeds. L. Averyanov 15 July 2014.

FLORA OF CAMBODIA

FLORA OF LAOS Convallariaceae Peliosanthes choriandra Aver. et al. Type Luang Prabang prov., Pon Xay distr., Houay Man village, Phou Pak Sang Mountain, around point 19°59'03.8''N 102°25'09.0''E. Degraded primary evergreen and semideciduous forest on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. 1000-1150 m a.s.l. Terrestrial herb on steep shady rocky mountain slope. Flowers green. Occasional. LA-VN 2059 4 April 2017 Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.Maisak, Vichith Lamxay, Koudkeo Phommachanh, Phizar Sayalath, Keooudone Souvannakhoummane. d-EXSICCATES OF VIETNAMESE FLORA 0282/LA-VN 2059 © L.Averyanov

FLORA OF VIETNAM

FLORA OF LAOS

FLORA OF LAOS Convallariaceae Peliosanthes densiflora Aver. et N.Tanaka sp.nov. Laos, Vientiane prov., Kasi. Берег реки Nam Pong Gnai, новая дорога на Луанг Пробанг, 13 км от Каси. 827 м над ур. м., 19°21.978N, 102°09.282E. Е.Константинов K140. сбор живым корневищем, гербарий/фото 22.10.2013.

FLORA OF VIETNAM Liliaceae s.l. (Convallariaceae) Peliosanthes sp.nov. Lai Chau Prov., Phong Tho Distr., Sin Suoi Ho Municipality, Sa Ma Pho village, around point 22°33’21’’N 103°34’29’’E. Secondary evergreen broad-leaved wet closed forest with bamboo on very steep mountain slopes and along ridges composed predominantly with weathered granite at elev. about 1900-2050 m a.s.l. Terrestrial herb on shady rocky slope. Flowers light dull yellowish. Fruits blue, juicy. Locally very common. HAL 10767 14 Dec. 2006 Photo. Coll.: Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc, Pham Van The, N.T.Vinh. d-EXSICCATES OF VIETNAMESE FLORA 0089/HAL10767 (HN, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF LAOS Convallariaceae Peliosanthes tatianae Aver et al., sp.nov. Type Vientiane prov., Vang Vieng distr., Patang village, around point 19°04'15.6''N 102°24'33.2''E. Secondary broad-leaved semi-deciduous dry forest on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. about 670 m a.s.l. Terrestrial herb in shady steep slope. Flower buds white. Not rare. LA-VN 1607 23 March 2017 Photos Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.Maisak, Khamla Inkhavilay, Hongthong Southyvohanh, Pinmany Sisavath, Chanthala Cheuaysamphanh.

FLORA OF VIETNAM

FLORA OF VIETNAM Liliaceae s.l. (Convallariaceae) Peliosanthes retroflexa Aver. et N.Tanaka Type Quang Binh Prov., Minh Hoa Distr., Dan Hoa Municipality, Giang Man Ridge (Vietnam-Laos border) near Cha Lo Vietnam-Lao border gate around point 17°41’50”N 105°45’54”E. Primary evergreen broad-leaved and coniferous (with Dacrycarpus and Fokienia) wet forest along ridge edge composed with shale and sandstone at elevation about 900-1100 m a.s.l. Terrestrial herb on shady slope. Fruits blue. Common. HAL 12374 8 February 2009 Coll.: L.Averyanov, P.K.Loc, N.T.Vinh, L.T.Son.

FLORA OF LAOS Convallariaceae Rohdea dracaenoides Aver. et N.Tanaka, sp.nov. Type. Vientiane Prov., Kasi Distr., Namken village, Phachao Mt., around point 19°18’45.5”N 102°22’31.4”E. Primary broad-leaved evergreen forest on very steep rocky slopes of remnant mountain composed with highly eroded solid crystalline limestone at elevations 1500-1750 m. Lithophyte on steep rocky slopes and shady cliffs. Flowers light green to light yellow-orange. Very common from elev. 1600 to mountain top. LA-VN 785 24 March 2013 Coll.: L.Averyanov, N.S.Khang, S.Lorphengsy

FLORA OF LAOS Convallariaceae Rohdea dracaenoides Aver. et N.Tanaka, sp.nov. Type. Vientiane Prov., Kasi Distr., Namken village, Phachao Mt., around point 19°18’45.5”N 102°22’31.4”E. Primary broad-leaved evergreen forest on very steep rocky slopes of remnant mountain composed with highly eroded solid crystalline limestone at elevations 1500-1750 m. Lithophyte on steep rocky slopes and shady cliffs. Flowers light green to light yellow-orange. Very common from elev. 1600 to mountain top. LA-VN 785 24 March 2013 Coll.: L.Averyanov, N.S.Khang, S.Lorphengsy

FLORA OF VIETNAM Convallariaceae Tupistra khangii sp.nov. Son La Prov., Van Ho Distr., Tan Xuan Municipality, Cot Moc village, territory of Xuan Nha natural reserve, eastern slopes of Pha Luong Mountain. Remnants of primary and secondary broad-leaved evergreen humid forest on very steep mountain slopes composed by shale and sandstone at elevation about 1000 m a.s.l. around point 20°40’33.3’’N 104°39’00.3’’E. Large lithophytic clustering herb on large mossy stream boulders along narrow shady rocky stream canyon. Flowers white, anthers light yellowish. Occasional. 15 November 2013 CPC 7158 Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, N.D.Thang, L.D.Qui Photos

FLORA OF VIETNAM Fam. Convallariaceae Tupistra theana Aver. et N.Tanaka, sp.nov. Epitype Quang Binh Prov., Minh Hoa Distr., Hoa Tien municipality, La Van village, around point 17°51’45”N 105°49’45”E. Highly degraded primary and secondary evergreen broad-leaved forest on very steep rocky slopes of remnant limestone hills composed with highly eroded marble-like crystalline limestone at elev. 200-450 m a.s.l. Lithophytic herb on shelves and in crevices of tall shady vertical cliff near to top of mountain. Not common. CPC 2581 30 April 2011. Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, P.V.The, N.T.Vinh. d-EXSICCATES OF VIETNAMESE FLORA 0182/HAL 2581 (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VIETNAM Convallariaceae Tupistrapatula Aver., N. Tanaka et K.S. Nhuyen Isotype Ha Giang Prov., Bac Me Distr., Thuong Tan municipality. Primary broad-leaved humid evergreen forest on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation 1200-1285 m a.s.l. around point 22°38’19.0’’N 105°17’06.5’’E. Terrestrial herb on mountain shady slope in very wet place. Old flowers dull yellowish, then entirely black. Not common. 16 November 2014 CPC 7492 Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz (holotype: Herbarium of the Center for Plant Conservation, Hanoi)

FLORA OF VIETNAM Fam. Convallariaceae Tupistra theana Aver. et N.Tanaka, sp.nov. Epitype Quang Binh Prov., Minh Hoa Distr., Hoa Tien municipality, La Van village, around point 17°51’45”N 105°49’45”E. Highly degraded primary and secondary evergreen broad-leaved forest on very steep rocky slopes of remnant limestone hills composed with highly eroded marble-like crystalline limestone at elev. 200-450 m a.s.l. Lithophytic herb on shelves and in crevices of tall shady vertical cliff near to top of mountain. Not common. CPC 2581 30 April 2011. Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, P.V.The, N.T.Vinh. d-EXSICCATES OF VIETNAMESE FLORA 0182/HAL 2581 (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF LAOS Convallariaceae Tupistra orlovii Aver., N. Tanaka & K. S. Nguyen, sp.nov. Bolikhamsai prov., Thaphabat distr., Nam Bon village, Phou Khao Khouay national park, Tad Xai Waterfall, around point 18°27'25.9''N 103°08'17.5''E. Degraded primary and secondary evergreen dry forest along river on eroded sandstone at elev. About 340 m a.s.l. Lithophytic and occasionally terrestrial large herb on shady mossy boulders and river cliffs. Leaves uniform green, fruits glossy green. Rare, but locally abundant. LA-VN 2191 11 April 2017 Photos Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.Maisak. d-EXSICCATES OF VIETNAMESE FLORA 0283/Orlov s.n. & LA-VN 2191 © L.Averyanov

FLORA OF VIETNAM

FLORA OF VIETNAM Convallariaceae Ophiopogon petraeus Aver. et N.Tanaka Type Lang Son Prov., Huu Lung Distr., Huu Lien Commune, Huu Lien Nature Reserve, around point 21°40’28”N, 106°22’34”E. Primary closed evergreen seasonal tropical mixed forests on rocky limestone at elevation 400-600 m. Lithophytic rosulate herb. Not rare. HAL 11220b 24 September 2007 Coll.: P.K.Loc, N.T.Vinh, H.V.Luat Flowered under cultivation at September-October 2013. Flowers white, anthers light yellow-green, pedicel white with violet tint. Leaves fleshy, at the base subsucculent, juicy, transversely-oval in section. Photos. L. Averyanov, HAL 11220b/1. d-EXSICCATES OF VIETNAMESE FLORA 0207/HAL 11220b (LE) © L.Averyanov

FLORA OF VIETNAM Convallariaceae Ophiopogon fruticulosa Aver., N.Tanaka et S.K.Nguyen, sp.nov. Type Tuyen Quang Prov., Na Hang Distr., Sinh Long Municipality, Khuoi Phin village, around point 22°38'21"N, 105°20'28"E. Remnants of primary forest along stream, on wet very steep shady humid travertine river valley. Erect shrub to 1.2 m tall in shady wet place. Flowers white with light violet tint. Locally abundant. 30 September 2013 CPC 5345 L.Averyanov, N.T.Hiep, L.M.Tuan, N.S.Khang, D.V.Cuong, N.V.Truong.T.Maisak, L.Osinovets

FLORA OF LAOS Convallariaceae Peliosanthes choriandra Aver. et al. Type Luang Prabang prov., Pon Xay distr., Houay Man village, Phou Pak Sang Mountain, around point 19°59'03.8''N 102°25'09.0''E. Degraded primary evergreen and semideciduous forest on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. 1000-1150 m a.s.l. Terrestrial herb on steep shady rocky mountain slope. Flowers green. Occasional. LA-VN 2059 4 April 2017 Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.Maisak, Vichith Lamxay, Koudkeo Phommachanh, Phizar Sayalath, Keooudone Souvannakhoummane. d-EXSICCATES OF VIETNAMESE FLORA 0282/LA-VN 2059 © L.Averyanov

FLORA OF LAOS Convallariaceae Peliosanthes densiflora Aver. et N.Tanaka sp.nov. Laos, Vientiane prov., Kasi. Берег реки Nam Pong Gnai, новая дорога на Луанг Пробанг, 13 км от Каси. 827 м над ур. м., 19°21.978N, 102°09.282E. Е.Константинов K141. сбор/фото 22.10.2013. К142 Amorphophallus sp. сбор фото 22.10.2013 Берег реки Nam Pong Gnai, новая дорога на Луанг Пробанг, 13 км от Каси 2 живых клубня 827 м над ур. м. N 19 21.978 E 102 09.282

FLORA OF VIETNAM Liliaceae s.l. (Convallariaceae) Peliosanthes sp.nov. Lai Chau Prov., Phong Tho Distr., Sin Suoi Ho Municipality, Sa Ma Pho village, around point 22°33’21’’N 103°34’29’’E. Secondary evergreen broad-leaved wet closed forest with bamboo on very steep mountain slopes and along ridges composed predominantly with weathered granite at elev. about 1900-2050 m a.s.l. Terrestrial herb on shady rocky slope. Flowers light dull yellowish. Fruits blue, juicy. Locally very common. HAL 10767 14 Dec. 2006 Photo. Coll.: Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc, Pham Van The, N.T.Vinh. d-EXSICCATES OF VIETNAMESE FLORA 0089/HAL10767 (HN, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF LAOS

FLORA OF LAOS Convallariaceae Peliosanthes densiflora Aver. et N.Tanaka sp.nov. Laos, Vientiane prov., Kasi. Окрестности пещеры Khuang Lang Cave берег лесной реки Древостой включает: Tetrameles nudiflora, Pometia eximia, Croton argyratus, Arenga westerhoutii, Ficus religlosa. По табличкам нац. парка. 631 м над ур. м.,N 19°18.198N, 102°09.031E. Е.Константинов K103. сбор живым корневищем, гербарий/фото 21.10.2013.

FLORA OF LAOS Convallariaceae Peliosanthes argenteostriata Laos, Vientiane prov., окрестности Kasi, elev. 700 m a.s.l. старая дорога на Луангпробанг 40 км от Каси на север, облесенный берег реки Nam Kean, подножие западного склона горы Ph. Phaday 30.11.2014 E.Konstantinov K 305. Сбор. Фото

FLORA OF LAOS Convallariaceae Peliosanthes tatianae Aver et al., sp.nov. Type Vientiane prov., Vang Vieng distr., Patang village, around point 19°04'15.6''N 102°24'33.2''E. Secondary broad-leaved semi-deciduous dry forest on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. about 670 m a.s.l. Terrestrial herb in shady steep slope. Flower buds white. Not rare. LA-VN 1607 23 March 2017 Photos Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.Maisak, Khamla Inkhavilay, Hongthong Southyvohanh, Pinmany Sisavath, Chanthala Cheuaysamphanh.

FLORA OF VIETNAM

FLORA OF VIETNAM

FLORA OF VIETNAM Liliaceae s.l. (Convallariaceae) Peliosanthes retroflexa Aver. et N.Tanaka Type Quang Binh Prov., Minh Hoa Distr., Dan Hoa Municipality, Giang Man Ridge (Vietnam-Laos border) near Cha Lo Vietnam-Lao border gate around point 17°41’50”N 105°45’54”E. Primary evergreen broad-leaved and coniferous (with Dacrycarpus and Fokienia) wet forest along ridge edge composed with shale and sandstone at elevation about 900-1100 m a.s.l. Terrestrial herb on shady slope. Fruits blue. Common. HAL 12374 8 February 2009 Coll.: L.Averyanov, P.K.Loc, N.T.Vinh, L.T.Son.

FLORA OF LAOS Convallariaceae Rohdea dracaenoides Aver. et N.Tanaka, sp.nov. Type. Vientiane Prov., Kasi Distr., Namken village, Phachao Mt., around point 19°18’45.5”N 102°22’31.4”E. Primary broad-leaved evergreen forest on very steep rocky slopes of remnant mountain composed with highly eroded solid crystalline limestone at elevations 1500-1750 m. Lithophyte on steep rocky slopes and shady cliffs. Flowers light green to light yellow-orange. Very common from elev. 1600 to mountain top. LA-VN 785 24 March 2013 Coll.: L.Averyanov, N.S.Khang, S.Lorphengsy

FLORA OF LAOS Convallariaceae Tupistra orlovii Aver., N. Tanaka & K. S. Nguyen, sp.nov. Bolikhamsai prov., Thaphabat distr., Nam Bon village, Phou Khao Khouay national park, Tad Xai Waterfall, around point 18°27'25.9''N 103°08'17.5''E. Degraded primary and secondary evergreen dry forest along river on eroded sandstone at elev. About 340 m a.s.l. Lithophytic and occasionally terrestrial large herb on shady mossy boulders and river cliffs. Leaves uniform green, fruits glossy green. Rare, but locally abundant. LA-VN 2191 11 April 2017 Photos Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.Maisak. d-EXSICCATES OF VIETNAMESE FLORA 0283/Orlov s.n. & LA-VN 2191 © L.Averyanov

FLORA OF LAOS Convallariaceae Tupistra theana Aver. et N.Tanaka Laos, Khammouane prov., Thakhek. На известняковых отломах останца в подлеске. Растение с длинными жесткими листьями. 161 м над ур. м., 17°27.195N, 104°57.790E. Е.Константинов K248. Гербарный сбор/фото 03.11.2013.

FLORA OF VIETNAM Fam. Convallariaceae Tupistra theana Aver. et N.Tanaka, sp.nov. Epitype Quang Binh Prov., Minh Hoa Distr., Hoa Tien municipality, La Van village, around point 17°51’45”N 105°49’45”E. Highly degraded primary and secondary evergreen broad-leaved forest on very steep rocky slopes of remnant limestone hills composed with highly eroded marble-like crystalline limestone at elev. 200-450 m a.s.l. Lithophytic herb on shelves and in crevices of tall shady vertical cliff near to top of mountain. Not common. CPC 2581 30 April 2011. Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, P.V.The, N.T.Vinh. d-EXSICCATES OF VIETNAMESE FLORA 0182/HAL 2581 (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VIETNAM Fam. Convallariaceae Tupistra theana Aver. et N.Tanaka, sp.nov. Epitype Quang Binh Prov., Minh Hoa Distr., Hoa Tien municipality, La Van village, around point 17°51’45”N 105°49’45”E. Highly degraded primary and secondary evergreen broad-leaved forest on very steep rocky slopes of remnant limestone hills composed with highly eroded marble-like crystalline limestone at elev. 200-450 m a.s.l. Lithophytic herb on shelves and in crevices of tall shady vertical cliff near to top of mountain. Not common. CPC 2581 30 April 2011. Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, P.V.The, N.T.Vinh. d-EXSICCATES OF VIETNAMESE FLORA 0182/HAL 2581 (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc