Herbarium LE - Specimens

Asteraceae

Aspidistra sp.

Khanh Hoa Provinces, road to Khanh Le Pass, east slope of Hon Giao Ridge, dry evergreen broad-leaved forest on granite at elevation 600-700 m a.s.l. Lithophytic herb on steep rocky slope of small stream. Locally common. Flower buds dark violet. 21 April 2025, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3902.

Flowered in cultivation on 17 June 2025, Nguyen Van Canh, AL3902

Euphorbiaceae

Phyllanthus reticulatus Poir.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone and limestone slope at elevation about 80-150 m a.s.l. along forest trail. Shrub or treelet to 5 m tall on open rocky slope. Fruits reddish-purple. Not common.

17 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4363 (NHC163).

Ebenaceae

Diospyros sp. (Maba sp.)

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone and limestone slope at elevation about 80-150 m a.s.l. along forest trail. Treelet to 2 m tall in shady rocky slope. Flowers white. Not common.

17 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4362 (NHC162).

Convallariaceae

Polygonatum punctatum Royle ex Kunth?

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone and limestone slope at elevation about 80-150 m a.s.l. along forest trail. Terrestrial and lithophytic rhizomatous herb to 1m talon rocky slope. Flowers white. Not common.

17 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4361 (NHC161).

Euphorbiaceae

Suregada multiflora (A.Juss.) Baill.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone and limestone slope at elevation about 80-150 m a.s.l. along forest trail. Tree 6-7 m tall on limestone slope. Flower buds greenish Not common.

17 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4360 (NHC160).

Cyperaceae

Gahnia sp.?

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone and limestone slope at elevation about 80-150 m a.s.l. along forest trail. Terrestrial sedge in wet open place. Not common.

17 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4359 (NHC159).

Moraceae

Ficus sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone and limestone slope at elevation about 80-150 m a.s.l. along forest trail. Tree 6-7 m tall on limestone slope. Syconia green to yellowish-green. Common.

17 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4358 (NHC158).

Fabaceae

Baphia capparidifolia Baker

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone and limestone slope at elevation about 80-150 m a.s.l. along forest trail. Scandent shrub with shoots to 3 m long in open place. Flowers white. Not common.

17 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4357 (NHC157).

Euphorbiaceae

Sapium discolor (Champ. ex Benth.) Müll.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone and limestone slope at elevation about 80-150 m a.s.l. along forest trail. Tree 6-7 m tall on limestone slope. Flowers greenish. Common.

17 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4356 (NHC156).

Polypodiaceae s.l.

Lygodium japonicum (Thunb.) Sw.?

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone and limestone slope at elevation about 80-150 m a.s.l. along forest trail. Herbaceous vine to 3 m long in open place along trail. Very common.

17 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4355 (NHC155).

Arecaceae

Phoenix sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone and limestone slope at elevation about 80-150 m a.s.l. along forest trail. Palm to 2 m tall among limestone rocks. Not rare.

17 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4354 (NHC154).

Convallariaceae

Peliosanthes macrostegia Hance

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone and limestone slope at elevation about 80-150 m a.s.l. along forest trail. Terrestrial rhizomatous herb in shady place. Flowers dirty violet. Locally common.

17 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4353 (NHC153).

Rhizophoraceae

Rhizophora mucronata Poir.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Littoral mangrove shrub or treelet to 2.5 m tall in flooded littoral zone. Flower. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4352 (NHC152).

Euphorbiaceae

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Littoral mangrove shrub or treelet with milky juice to 3 m tall along flooded littoral zone. Flower buds yellowish green. Very common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4351 (NHC151).

Verbenaceae

Lumnitzera sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Littoral mangrove shrub or treelet to 2 m tall in flooded littoral zone. Flower buds yellowish green. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4350 (NHC150).

Capparidaceae

Capparis sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Secondary evergreen broad-leaved forest with bamboo on limestone cliffs along seashore. Scandent shrub or woody vine to 5-6 m long along seashore line among sandstone rocks. Flowers white, tepals with yellow or purple brown spot near base. Not common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4348 (NHC148).

Fabaceae

Derris sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Secondary evergreen broad-leaved forest with bamboo on limestone cliffs along seashore. Tree about 10 m tall along seashore line among sandstone rocks. Flowers purple. Occasional.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4347 (NHC147).

Anacardiaceae

Pistacia weinmanniifolia J.Poiss. ex Franch.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Secondary evergreen broad-leaved forest with bamboo on limestone cliffs along seashore. Treelet about 5 m tall on vertical limestone cliff along seashore line. Occasional.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4346 (NHC146).

Vitaceae

Ampelopsis sp.?

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Secondary evergreen broad-leaved forest with bamboo on limestone cliffs along seashore. Herbaceous to semiwoody vine to 2-4 m long on vertical limestone cliff along seashore line. Flower buds light green. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4345 (NHC145).

Asclepiadaceae

Dischidia sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Secondary evergreen broad-leaved forest with bamboo on limestone cliffs along seashore. Herbaceous vine to 3 m long with milky juice on vertical limestone cliff along seashore line. Flowers white to light yellowish. Very common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4344 (NHC144).

Arecaceae

Livistona chinensis (Jacq.) R.Br. ex Mart.?

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4377. Photo only.

Arecaceae

Korthalsia sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone and limestone slope at elevation about 50 m a.s.l. along forest trail. Woody vine to 5 m long with ascending apex. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4376. Photo only.

Rubiaceae

Ixora sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Undershrub to shrum 0.5-1.5 m tall in open place along trail. Flowers bright red. Very common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4375. Photo only.

Pandanaceae

Pandanus tectorius Parkinson

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Sublittoral palm-like shrub to 3 m tall along rocky seashore line among open limestone and sandstone boulders. Fruits orange to orange-red. Very common

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4374. Photo only.

Myrsinaceae

Aegiceras corniculatum (L.) Blanco

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Littoral mangrove shrub to 1.5 m tall in flooded littoral zone. Flowers white. Very common. Main dominant of mangrove plant community.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4349 (NHC149).

Theaceae

Camellia oleifera C.Abel

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Treelet 7-8 m tall and about 20 cm DBH along trail. Flower buds light green, flowers white. Not common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4343 (NHC143).

Sapindaceae

Nephelium lappaceum L.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Tree to 15 m tall along trail. Fruits green. Not common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4342 (NHC142).

Verbenaceae

Clerodendrum japonicum (Thunb.) Sweet

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Undershrub to 2 m tall in open rocky place along trail. Flowers bright red. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4341 (NHC141).

Euphorbiaceae

Antidesma fordii Hemsl.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Treelet 4-6 m tall in open place along trail. Flowers light greenish. Not rare.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4340 (NHC140).

Myrsinaceae

Ardisia sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Treelet to 2.5 m tall in open place along trail. Flowers pink-purple. Occasional.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4339 (NHC139).

Vitaceae

Cayratia sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Herbaceous vine to 3 m long in open place along trail. Young fruits green. Not rare.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4338 (NHC138).

Fabaceae s.l. (Caesalpiniaceae)

Peltophorum pterocarpum (DC.) Backer ex K.Heyne

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Tree to 20 m tall and 0.8 m DBH. Flowers bright yellow. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4337 (NHC137).

Rubiaceae

Ixora finlaysoniana Wall.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Shrub to 1.5 m tall in open place along trail. Flowers white. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4336 (NHC136).

Sapindaceae

Xerospermum noronhianum (Blume) Blume

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Tree to 20 m tall. Flowers white to yellowish. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4335 (NHC135).

Pandaceae

Microdesmis caseariifolia Planch. ex Hook.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Treelet to 6-7 m tall in open place along trail. Flowers bright yellow. Not common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4334 (NHC134).

Rutaceae

Micromelum falcatum (Lour.) Tanaka

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Shrub or small treelet to 3 m tall in open place along trail. Fruits orange to red. Very common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4333 (NHC133).

Apocynaceae

Tabernaemontana bufalina Lour.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Shrub to 3 m tall in open place along trail. Flowers white, fruits orange. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4332 (NHC132).

Annonaceae

Uvaria littoralis (Blume) Blume

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Woody vine to 10 m long. Flowers bright red. Not common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4331 (NHC131).

Fabaceae

Baphia capparidifolia Baker

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Scandent shrub with shoots to 3 m long in open place. Flowers white. Not common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4330 (NHC130).

Fabaceae s.l. (Mimosaceae)

Entada phaseoloides (L.) Merr.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Woody vine to 25 m long on shady slope. Not rare.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4329 (NHC129).

Oleaceae

Myxopyrum smilacifolium (Wall.) Blume

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Woody vine to 5 m long in open places along trail. Medicinal plant. Locally common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4328 (NHC128).

Rosaceae

Rubus leucanthus Hance

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Climbing shrub with shoots to 2 m long in open places along trail. Flowers white. Very common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4327 (NHC127).

Sauraiaceae

Saurauia tristyla DC.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Treelet to 6 m tall in open wet rocky place. Flowers light pink. Occasional.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4326 (NHC126).

Fabaceae s.l. (Caesalpiniaceae)

Cassia sp.?

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Woody vine to 10 m long in open place. Flowers bright yellow. Not cmmon.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4325 (NHC125).

Moraceae

Streblus sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Tree to 10 m tall in shady place. Fruits dull green. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4324 (NHC124).

Polypodiaceae s.l.

Leptochilus sp.?

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Creeping lithophytic fern in shady place. Not rare.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4323 (NHC123).

Euphorbiaceae

Antidesma ghaesembilla Gaertn.?

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Treelet or shrub to 3 m tall in rather open place. Flowers greenish. Common.

Anacardiaceae

Allospondias lakonensis (Pierre) Stapf

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Tree about 15 m tall in rather open place. Flowers white. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4320 (NHC121).

Simaroubaceae

Eurycoma longifolia Jack

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Treelet to 4 m tall in rather open place. Flowers purplish. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4319 (NHC120).

Apocynaceae

Tabernaemontana pauciflora Blume

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Treelet to 4 m tall in rather open place. Flowers white. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4318 (NHC119).