Herbarium LE - Specimens

DRYOPTERIDACEAE

Polystichum

Son La

Yen Chau District; Muong Lum

Municipality; On Oc Village; collection

ca. 1 km from the village of On Oc on

road between On Oc and Lum village.

20°59'44"N 104°30'03"E 1080 m

Occasional, terrestrial, in moist shade.

Rachis light green with brown hairs;

short stem "woody", hard; sporangia

brown.

4 March 2001

D.K. Harder, P.K. Loc & N.T. Huong 7243

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

DRYOPTERIDACEAE

Pteridrys

Son La

Yen Chau District; Muong Lum

Municipality; On Oc Village; collection

ca. 1 km from the village of On Oc on

road between On Oc and Lum village.

20°59'44"N 104°30'03"E 1080 m

Occasional, terrestrial from short stem;

in shady, moist place. Rachis fluted and

brown at base; pinnae dark green;

sporangia brown; hairs at base of rachis

dark brown-black.

4 March 2001

D.K. Harder, P.K. Loc & N.T. Huong 7248

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

POLYPODIACEAE

Polypodium

Son La

Moc Chau District; Van Ho Municipality,

Hua Tat Village. Collections on

limestone ridge; degraded forest to

cultivation on lower slope to primary

evergreen, closed forest on limestone

cliff/ridge; northern exposure.

20°46'35"N 104°46'31"E 1180 - 1250 m

Lithophytic creeping fern on vertical

shady cliff. Common.

5 March 2001

D.K. Harder, P.K. Loc, N.X. Tam & L.V.

Averyanov 7318

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

DRYOPTERIDACEAE

Polystichum

Son La

Moc Chau District; Van Ho Municipality,

Hua Tat Village. Collections on

limestone ridge; degraded forest to

cultivation on lower slope to primary

evergreen, closed forest on limestone

cliff/ridge; northern exposure.

20°46'35"N 104°46'31"E 1180 - 1250 m

Lithophytic fern. Very common.

5 March 2001

D.K. Harder, P.K. Loc, N.X. Tam & L.V.

Averyanov 7313

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

DRYOPTERIDACEAE

Polystichum

Lao Cai

Van Ban District; Nam Xe Municipality;

Nam Xi Tan Village. Collection in forest

on steep W-facing slope along stream;

just above village; on N side of road

between Nam Xe and Khau Co Pass at near

40 km to Than Uyen mark.

22°02'31"N 103°58'32"E 700 - 1113 m

Occasional, on lower slope near stream

and higher up. Rachis green, shiny;

26 February 2001

D.K. Harder, P.K. Loc, N.T. Hiep, G.E.

Schatz, S. Bodine & N.Q. Hieu 6881

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

Pteridophyta J.Y. Bergen & B.M. Davis

Son La

Moc Chau District; Long Luong

Municipality; Sang Cai Village.

Collections in primary, wet,

broad-leaved, evergreen and

semi-deciduous, mossy forest along

limestone ridge situated between

1400-1500 m elevation.

20°45'51"N 104°51'26"E 1300 - 1500 m

Terrestrial and lithophytic fern. Very

8 March 2001

D.K. Harder, P.K. Loc, L.V. Averyanov &

N.X. Tam 7482

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

POLYPODIACEAE

Pyrrosia

Ha Giang

Vi Xuyen District; Minh Tan Municipality

Ban Ngay Village. Collection on E

facing ridge above river; on exposed

limestone rocks; collections in

disturbed agricultural land leading into

primary forest with some deforestation

of Burretiodendron (Tiliaceae) trees.

23°00'09"N 104°55'21"E 225 - 621 m

23°00'37"N 104°54'46"E

Occasional, on rocks. Thick tomentum on

17 February 2001

D.K. Harder, P.K. Loc, N.T. Hiep, G.E.

Schatz & S. Bodine 6474

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

Orchidaceae

Bulbophyllum umbellatum Lindl.

Lam Dong Prov., Dalat City area sine loc.

18.06.2026, L. Averyanov et al., AL 5059

Orchidaceae

Liparis cespitosa (Lam.) Lindl.

Son La Prov., Thuan Chau District.

14 August 2025, Truong Ba Vuong, BV 2004

Orchidaceae

Bulbophyllum protractum Hook.f.

Cao Bang Province, Trung Khanh District.

2 July 2025, Truong Ba Vuong, Numer: BV1993.

Inflorescence as long as or shorter than pseudobulbs, from base of pseudobulbs or along the rhizome nodes, carry 2 to 5 flowers, petals 3 veins, lip triangular, with 2 rounded lateral keels fused at apical part of lip, stelidia subulate, slender, seta like.

Orchidaceae

Bulbophyllum strigosum (Garay) Sieder & Kiehn

Kon Tum Province, Tu Mo Rong District.

23.07.2025, Truong Ba Vuong, BV2005.

Bulbophyllum section Rhytionanthos, lip remarkable hairy for B. strigosum.

Orchidaceae

Malaxis sp.

Dak Lak Province, Yok Don National Park, open dipterocarp forest at elevation 100-200 m a.s.l. Terrestrial herb. Not common.

September 2025, Nguyen Van Canh, AL4576.

Orchidaceae

Acriopsis indica Wright

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Clustering epiphyte on tall trees. Not rare.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4617.

Orchidaceae

Apostasia wallichii

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 800-900 m a.s.l. around the point 12º10’N 107º46’E. Erect terrestrial herb to 0.5 m tall in shady place. Occasional.

30 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4646.

Orchidaceae

Calanthe lyroglossa aff.

Dak Lak Province, M’Drak District, W foothills of Chu Mu Mt., dry secondary and remnants of primary evergreen broad-leaved forest on granite on steep slopes along rocky river at elevation 600-650 m a.s.l. Terrestrial herb to 1 m tall in shady wet stream slope along trail. Flowers yellow. Not rare.

14 February 2026, Nguyen Van Canh, AL4540.

Orchidaceae

Podochilus sp.

Khanh Hoa Province, Khanh Vinh District, Khanh Le Pass. Primary evergreen broad-leaved forest on very steep rocky mountain slopes on granite and gneiss at elevation of about 1000 m a.s.l., around point 12º13’N 108º44’E. Clustering lithophyte on big mossy boulders along rocky stream. Common.

17 April 2026, L. Averyanov, Nguyen Van Canh, T. Maisak, АL4926 (living plant also).

Orchidaceae

Taeniophyllum sp.

Binh Dinh Province, Van Canh District. Secondary evergreen broad-leaved forest with bamboo on granite at elevation 250-350 m a.s.l., around point 13º38’N 108º57’E. Leafless epiphyte on small tree along rocky river in shady place. Rare.

12 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4892.

Flowered in culture on 26 May 2026, L. Averyanov, AL4892.

Orchidaceae

Bulbophyllum sp. (sect. Desmosanthes)

Gia Lai Province, K'Bang District. Secondary and remnants of primary evergreen broad-leaved dry forest with Dacrydium on lava rocks (red soils) at elevation about 1000 m a.s.l., around point 14º15N 108º35E. Clustering and pendulous epiphyte on tall tree. Common.

6 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4800 (living plant only).

Orchidaceae

Bulbophyllum semiteretifolium Gagnep.

Lam Dong Province, Lac Duong District. Humid evergreen broad-leaved forest with Ducampopinus on granite along tops of ridge at elevation of about 1750 m a.s.l. around point 12º06’N 108º22’E. Epiphyte on mossy tree. Flowers white. Common.

18 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, T. Maisak, АL4957.

Orchidaceae

Bulbophyllum salmoneum Aver. & J.J.Verm.

Gia Lai Province, K'Bang District. Highly degraded secondary evergreen broad-leaved dry forest at elevation 600-800 m a.s.l., around point 14º26N 108º22E. Creeping epiphyte on tall tree. Locally common.

7 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4769.

Orchidaceae

Bulbophyllum retusiusculum

Gia Lai Province, Mang Yang District, Ayune Commune, Kon Ka Kinh National Park. Secondary and primary dry evergreen broad-leaved forest on shale and lava rocks at elevation of 900-1000 m a.s.l. around point 14º13N 108º20E. Clustering epiphyte on tall tree. Locally common.

2 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4700.

Orchidaceae

Bulbophyllum manhdatii Vuong, Aver., V.S.Dang & Q.T.Truong

Lam Dong Province, Lac Duong District. Humid evergreen broad-leaved forest with Ducampopinus on granite along tops of ridge at elevation of about 1750 m a.s.l. around point 12º06’N 108º22’E. Epiphyte on mossy tree. Flowers yellowish. Common.

18 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, T. Maisak, АL4958.

Orchidaceae

Bulbophyllum lemniscatoides Rolfe

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 800-900 m a.s.l. around the point 12º10’N 107º46’E. Appressed clustering epiphyte on mossy tree along stream. Not common.

30 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4652.

Orchidaceae

Bulbophyllum depressum King & Pantl.?

Gia Lai Province, K'Bang District, Kon Bong Village, Kon Bong Waterfall. Highly degraded secondary evergreen broad-leaved dry forest on shale and lava rocks along river valley at elevation 800-900 m a.s.l., around point 14º30.5N 108º23E. Lithophyte on mossy boulders along stream. Occasional.

8 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4840.

Orchidaceae

Bulbophyllum clandestinum Lindl.

Khanh Hoa Province, Khanh Vinh District, Khanh Le Pass. Primary evergreen broad-leaved forest on very steep rocky mountain slopes on granite and gneiss at elevation of 1000-1200 m a.s.l., around point 12º13’N 108º44’E. Epiphyte on tall mossy trees. Common.

19 April 2026, L. Averyanov, Nguyen Van Canh, T. Maisak, АL4979.

Orchidaceae

Bulbophyllum clandestinum Lindl.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Pendulous epiphyte on mossy trees along rocky stream in shady places. Occasional.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4615.

Orchidaceae

Bulbophyllum clandestinum Lindl.

Gia Lai Province, Mang Yang District, Ayune Commune, Kon Ka Kinh National Park. Secondary and primary dry evergreen broad-leaved forest on shale and lava rocks at elevation of 900-1000 m a.s.l. around point 14º13N 108º20E. Clustering epiphyte on tall tree. Occasional.

3 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4723.

Orchidaceae

Bulbophyllum blaoense Aver. & Tich

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 1000-1200 m a.s.l. around the point 12º10’N 107º46’E. Creeping epiphyte on tall trees. Flowers yellowish. Occasional.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4630.

Orchidaceae

Biermannia calcarata Aver.

Quang Binh Province, Tuyen Hoa District, evergreen broad-leaved forest at elevation of about 600 m a.s.l.

September 2025, Nguyen Van Canh, AL4587.

Orchidaceae

Appendicula sp. “gracilis”

Gia Lai Province, K'Bang District, Kon Bong Village, Kon Bong Waterfall. Highly degraded secondary evergreen broad-leaved dry forest on shale and lava rocks along river valley at elevation 800-900 m a.s.l., around point 14º30.5N 108º23E. Lithophyte on mossy boulders along stream. Occasional.

8 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4839 (living plant only).

Orchidaceae

Bidoupia micrantha Aver., V.C.Nguyen & Duy

Kon Tum Province, evergreen broad-leaved forest at elevation of 800-900 m a.s.l.

12 September 2025, Pham Thi Thanh Dat, Nguyen Van Canh, AL4586 (PTT0221).

Orchidaceae

Anoectochilus roxburghii (Wall.) Lindl.

Lai Chau Province, Muong Te District, Mu Ca Commune. Old rich secondary evergreen broad-leaved forest on stratified shale at elevation of about 540 m a.s.l. around point 22º29’40’’N 102º24’15’’E. Subprostrate terrestrial and lithophytic herb on shady rocky slope. Leaves dark green or brown with network of yellow or pink nerves. Not common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5022.

Orchidaceae

Aerides odorata

Thanh Hoa Province, plant obtained from local collectors and cultivated in garden. Collected by Nguyen Van Canh on July 2025, AL4555.

Orchidaceae

Acriopsis indica

Binh Dinh Province, Vinh Thanh District. Primary and secondary evergreen broad-leaved forest on granite at elevation of about 800 m a.s.l., around point 14º10’N 108º43’E. Clustering epiphyte on tall tree. Flowers pinkish. Common.

10 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4876.

Orchidaceae

Acanthophippium striatum Lindl.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 1000-1200 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial herb to 0.4 m tall in shady place. Occasional.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4628.

Orchidaceae

Bulbophyllum affine

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 1000-1200 m a.s.l. around the point 12º10’N 107º46’E. Creeping epiphyte on tall trees. Common.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4632.

Orchidaceae

Bulbophyllum frostii Summerh.

Binh Dinh Province, Vinh Thanh District. Primary and secondary evergreen broad-leaved forest on granite at elevation of about 800 m a.s.l., around point 14º10’N 108º43’E. Creeping epiphyte on tall tree. Common.

10 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4872.

Orchidaceae

Eria tomentosa (J.Koenig) Hook.f.

Lai Chau Province, Muong Te District, Mu Ca Commune. Highly degraded secondary evergreen broad-leaved forest along rocky stream on stratified shale and granite on steep mountain slope at elevation of about 500 m a.s.l. around point 22º24’42’’N 102º24’35’’E. Epiphyte on tall tree. Plant flowered under cultivation.

23 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL4981.

Orchidaceae

Habenaria pantlingii?

Kon Tum Province, Kon Plong District, Hieu Municipality, remnants of primary evergreen broad-leaved forest on steep shady slopes or rocky river canyon composed with granite and silicate shale at elevation of about 1200 m a.s.l. Terrestrial erect tuberiferous herb to 1.2 m tall on rocky shady stream slope. Rare.

29 April 2025, L. Averyanov, N.V. Canh, T. Maisak, AL4060.

Orchidaceae

Grosourdya muscosa (Rolfe) Garay

Plant originated from local dealers in Kon Tum Province, Mang Den District.

Flowered in cultivation on 16 September 2025, Nguyen Van Canh, AL4486.

Orchidaceae

Grosourdya appendiculata (Blume) Rchb.f.?

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Epiphyte on small trees along stream in shady places. Locally common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4608.

Orchidaceae

Grosourdya appendiculata (Blume) Rchb.f.?

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Epiphyte on small trees along stream in shady places. Locally common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4608.

Orchidaceae

Goodyera procera (Ker Gawl.) Hook.

Gia Lai Province, K'Bang District, Kon Bong Village, Kon Bong Waterfall. Highly degraded secondary evergreen broad-leaved dry forest on shale and lava rocks along river valley at elevation 800-900 m a.s.l., around point 14º30.5N 108º23E. Erect lithophytic herb on wet river rocks. Flowers white. Not common.

8 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4824.

Orchidaceae

Flickingeria fimbriata

Gia Lai Province, Mang Yang District, Ayune Commune, Kon Ka Kinh National Park. Secondary and primary dry evergreen broad-leaved forest on shale and lava rocks at elevation of 900-1000 m a.s.l. around point 14º13N 108º20E. Clustering erect epiphytic undershrub on tall tree. Locally common.

2 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4702.

Orchidaceae

Erythrorchis altissima

Binh Dinh Province, Van Canh District. Secondary evergreen broad-leaved forest with bamboo on granite at elevation 250-350 m a.s.l., around point 13º38’N 108º57’E. Achlorophyllous creeping epiphytic vine to 5 m long. Flowers pale pinkish to almost white. Rare.

12 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4888.

Orchidaceae

Eria sp.

Gia Lai Province, Mang Yang District, Ayune Commune, Kon Ka Kinh National Park. Secondary and primary dry evergreen broad-leaved forest on shale and lava rocks at elevation of 900-1000 m a.s.l. around point 14º13N 108º20E. Clustering epiphyte on tall tree. Common.

3 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4724.

Orchidaceae

Eria tomentosa (J.Koenig) Hook.f.

Lai Chau Province, Muong Te District, Mu Ca Commune. Highly degraded secondary evergreen broad-leaved forest along rocky stream on stratified shale and granite on steep mountain slope at elevation of about 500 m a.s.l. around point 22º24’42’’N 102º24’35’’E. Epiphyte on tall tree. Plant flowered under cultivation.

23 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T.

Maisak, АL4981.

Orchidaceae

Eria scabrilinguis Lindl.

Gia Lai Province, K'Bang District. Highly degraded secondary evergreen broad-leaved forest on granite and basalt around large waterfall at elevation of about 700 m a.s.l., around point 14º20’N 108º40’E. Lithophytic herb on large mossy boulders along river. Not common.

10 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4865.

Orchidaceae

Eria perpusilla

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 800-900 m a.s.l. around the point 12º10’N 107º46’E. Appressed clustering epiphyte on mossy trees along rocky stream. Flowers white. Locally very common.

30 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4660.

Orchidaceae

Eria pannea Lindl.

Gia Lai Province, Mang Yang District, Ayune Commune, Kon Ka Kinh National Park. Secondary and primary dry evergreen broad-leaved forest on shale and lava rocks at elevation of 900-1000 m a.s.l. around point 14º13N 108º20E. Creeping epiphyte on tall tree. Locally very common.

2 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4695.