Herbarium LE - Specimens

Asclepiadaceae

Dischidia sp.

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Herbaceous creeping epiphytic wine to 1.5 m long on tall tree. Flowers white, Common.

09 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4204 (NHC015).

Lauraceae

Litsea baviensis Lecomte

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Treelet to 10 m tall on shady slope. Not common.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4230 (NHC041).

Zingiberaceae

Alpinia oblongifolia Hayata

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Terrestrial herb to 1 m tall in rather open place. Flowers white, median tepal with 2 red stripes. Locally common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4312 (NHC113).

Convallariaceae

Tupistra thangii N.Tanaka & D.D.Nguyen

Lam Dong Province, Bao Lam District, B 40 Pass, around point N11.4366 E107.4266. Fractionally logged primary evergreen broad-leaved and mixed forest with Dacrydium, Dacrycarpus and Pinus latteri on granite at elevation 800–900 m a.s.l. Terrestrial herb in dense forest. Locally common.

28 November 2022, L. Averyanov, Nguyen Van Canh, T. Maisak, Truong Ba Vuong, Nguyen Quoc Bao, AL 1862.

Araliaceae

Schefflera honbaensis Q.C.Truong⁣ ⟨not published name⟩

det. Truong Quang Cưong at 19.04.2025

The border of Lam Dong and Khanh Hoa Provinces, Khanh Le Pass (Hon Giao Pass) area, primary humid evergreen broad-leaved cloud forest on granite and gneiss at elevation 1750-1850 m a.s.l. Tree to 10 m tall along ridge edge. Common. 19 April 2026, Truong Quang Cuong, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3840.

Araliaceae

Schefflera honbaensis Q.C.Truong⁣ ⟨not published name⟩

det. Truong Quang Cưong at 19.04.2025

The border of Lam Dong and Khanh Hoa Provinces, Khanh Le Pass (Hon Giao Pass) area, primary humid evergreen broad-leaved cloud forest on granite and gneiss at elevation 1750-1850 m a.s.l. Tree to 10 m tall along ridge edge. Common. 19 April 2026, Truong Quang Cuong, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3840.

Melastomataceae

Perilimnastes setipetiola J.H.Dai, T.V.Do & Ying Liu

The border of Lam Dong and Khanh Hoa Provinces, Khanh Le Pass (Hon Giao Pass) area, primary humid evergreen broad-leaved cloud forest on granite and gneiss at elevation 1750-1850 m a.s.l. Terrestrial herb or undershrub to 1.5 m tall on wet rocky slope. Flowers white with pink tint. Locally very common. 19 April 2026, Truong Quang Cuong, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3847.

Smilacaceae

Smilax sp.

The border of Lam Dong and Khanh Hoa Provinces, Khanh Le Pass (Hon Giao Pass) area, primary humid evergreen broad-leaved cloud forest on granite and gneiss at elevation 1750-1850 m a.s.l. Herbaceous or semiwoody vine to 5 m long along ridge edge. Flowers greenish. Occasional 19 April 2026, Truong Quang Cuong, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3855.

Ericaceae

Rhododendron moulmainense Hook.

The border of Lam Dong and Khanh Hoa Provinces, Khanh Le Pass (Hon Giao Pass) area, primary humid evergreen broad-leaved and coniferous (with Pinis dalatensis and P. krempfii) cloud forest on granite and gneiss at elevation 1500-1600 m a.s.l. Treelet 4-5 m tall on steep slope along rocky stream. Not common. 20 April 2025, Truong Quang Cuong, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3880.

Zingiberaceae

Alpinia sp.

Dak Lak Province, Ea Hleo District, evergreen forest, at elevation of 900-1000 m a.s.l., herb to 1 m tall in forest. Common. Flowers wthite with purple.

February 2023, Nguyen Van Canh, AL3318.

Zingiberaceae

Alpinia sp.

Binh Dinh Province, Van Canh District, evergreen forest at elevation of about 1000 m a.s.l. Terrestrial herb to 1 m tall. Common. Flowers white with red.

2020, Nguyen Danh Duc, Nguyen Van Canh, AL3340.

Zingiberaceae

Newmania sp.

Binh Dinh Province, Van Canh District, evergreen forest at elevation of about 1000 m a.s.l. Terrestrial herb to 1 m tall. Common. Flowers white with purple.

2020, Nguyen Danh Duc, Nguyen Van Canh, AL3336.

Zingiberaceae

Amomum sp.? (new sp.?)

Gia Lai Province, Kbang District, Kon Chu Rang Nature Reserve, evergreen forest along stream, at elevation of about 700 m a.s.l., terrestrial herb to 1 m tall. Flowers white. Common.

May 2021, Nguyen Van Canh, AL3323.

Zingiberaceae

Alpinia hoangviet D.D.Nguyen & V.C.Nguyen

Dak Lak Province, M’Drak District, evergreen forest along stream at elevation of about 1300 m a.s.l., terrestrial herb to 1.5 m tall. Flowers white with yellow. Common.

June 2022, Nguyen Van Canh, AL3321.

Zingiberaceae

Alpinia sp. (new sp.?)

Gia Lai Province, Kbang District, Kon Chu Rang Nature Reserve, evergreen forest at elevation of about 700 m a.s.l., terrestrial herb to 1 m tall. Flowers white with red. Common.

May 2021, Nguyen Van Canh, AL3322.

Theaceae

Camellia sp.nov.?

Dak Lak Province, M’Drak District, Chu Mu Mountain, evergreen forest at elevation of about 1200 m a.s.l., treelet about 3 m tall, flowers yellow green.

2 September 2023 by Nguyen Van Canh, AL2451.

Pinaceae

Pinus dalatensis Ferre

Kon Tum Province, Kon Plong District, Mang Den Town area, remnants of humid primary evergreen broad-leaved and mixed forest (with Pinus dalatensis, Dacrydium and Dacrycarpus) on very steep mountain slopes composed with granite and silicate shale at elevation of about 1200 m a.s.l. Tree to 40 m tall and 2 m DBH on mountain top. Not common.

28 April 2025, L. Averyanov, N.V. Canh, T. Maisak, AL4031.

Orchidaceae

Cymbidium sp.

Ha Giang Province, Quan Ba District, wild collected plant from street market, 11 2022 Nguyen Van Canh, AL 2061. Living plant.

Russian-Vietnamese Tropical Centre

Orchidaceae

Dendrobium nobile Lindl. (“long”)

Wild collected cultivated plant originated from Kon Tum Province, Ngoc Linh Mountains. Cultivated in Kon Tum Province, Dak To Town by Mr. Nguyen Ta Ton.

01 May 2025, L. Averyanov, N.V. Canh, T. Maisak, AL4086.

Orchidaceae

Cymbidium sp. (ensifolium big leaves erect)

Dak Lak Province, Krong Bong District, Yang Hanh mt., evergreen broad-leaved forest at elevation of higher than 1700 m a.s.l.

2025. Nguyen Van Canh, AL3784. Living plant only and herbarium with flowers. Photos.

Orchidaceae

Cymbidium sp. (ensifolium big leaves erect)

Dak Lak Province, Krong Bong District, Yang Hanh mt., evergreen broad-leaved forest at elevation of higher than 1700 m a.s.l.

2025. Nguyen Van Canh, AL3784. Living plant only and herbarium with flowers. Photos.

Orchidaceae

Bulbophyllum pinicolum Gagnep.⁣

Lai Chau Prov. Sin Ho Distr.

Truong Ba Vuong, BV1942

03.11.2024

Det. Averyanov L.V. at 2025

Orchidaceae

Sunipia bicolor Lindl.

Lai Chau Prov. Sin Ho Distr.

Truong Ba Vuong, BV1939

02.12.2024

Det. Averyanov L.V. at 16.07.2025

Orchidaceae

Panisea yunnanensis S.C.Chen & Z.H.Tsi⁣

Lai Chau Prov. Sin Ho Distr.

Truong Ba Vuong, BV1940

02.12.2024

Det. Averyanov L.V. at 16.07.2025

Orchidaceae

Anoectochilus calcareus Aver.⁣

Son La Prov., Thuan Chau Distr.

Truong Ba Vuong, BV1206

27.07.2021

Det. Averyanov L.V. at 16.07.2025

Orchidaceae

Cylindrolobus truncatus (Lindl.) Rauschert

Quang Tri Prov.

Truong Ba Vuong, Le Tuan Anh

BV1942, 19.04.2025

Orchidaceae

Bulbophyllum bidoupense Aver. & Duy

The border of Lam Dong and Khanh Hoa Provinces, Khanh Le Pass (Hon Giao Pass) area, primary humid evergreen broad-leaved and coniferous (with Pinis dalatensis and P. krempfii) cloud forest on granite and gneiss at elevation 1500-1600 m a.s.l. Creeping epiphyte on tall tree. Common. 20 April 2025, Truong Quang Cuong, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3877.

Orchidaceae

Eria pannea Lindl.

The border of Lam Dong and Khanh Hoa Provinces, Khanh Le Pass (Hon Giao Pass) area, primary humid evergreen broad-leaved and coniferous (with Pinis dalatensis and P. krempfii) cloud forest on granite and gneiss at elevation 1500-1600 m a.s.l. Creeping epiphyte on tall tree. Common. 20 April 2025, Truong Quang Cuong, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3878.

Orchidaceae

Tainia longiscapa

Plant obtained from local collectos in Quang Binh Province.

December 2024, Nguyen Van Canh, AL3821.

Orchidaceae

Eria stricta Lindl.

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Clustering epiphyte on tall mossy tree. Very common.

09 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4195 (NHC006).

Orchidaceae

Eria javanica (Sw.) Blume

Bac Kan Province, Ba Be District, Ba Be National Park. Dry evergreen broad-leaved secondary forest on cliffs composed by white crystalline marble-like highly eroded limestone along lake shore on point 22º25’46’’N 105º36’58’E at elev. of about 170 m a.s.l. Lithophytic herb forming large clumps on very steep rocky slopes and cliffs. Flowers white. Locally common.

20 October 2023, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, Nguyen Van Ly, T. Maisak, AL2340.

Plants of Dong Chau - Khe Nuoc Trong Nature Reserve (DCKNTNR)

Orchidaceae

Appendicula hexandra (J.Koenig) J.J.Sm.?

Quang Binh Province, Le Thuy District, Kim Thuy Municipality. Secondary and primary fractionally logged evergreen broad-leaved forest on very steep hill slopes composed by gray and yellow solid marble-like limestone at elevation about 300–400 m a.s.l. around point 16º59.5 N 106º38.2 E. Tufted epiphyte or lithophyte in shady places. Not rare.

11 November 2024, Nguyen Huu Cuong, Nguyen Van Ly, Phan Thanh Quyet, Duong Van Nam, Le Thi Phuong Thao, L. Averyanov, T. Maisak, AL3661.

Convallariaceae

Aspidistra cylindrica Vislobokov & Nuraliev

Living plant received from N.V.Canh Garden on 26 November 2022, L. Averyanov, Nguyen Van Canh, T. Maisak, Truong Ba Vuong, et al., AL 1790.

Plant originated from Dak Lac Province, Chu Yang Sinh Mountain, evergreen broad-leaved forest, Nguyen Van Canh, 2022.

Flowered in culture on 13 August 2025, L. Averyanov, T. Maisak, AL1790a.

Orchidaceae

Dendrobium acerosum Lindl.⁣

Kon Tum Province, Dak Gley District, W slopes of Ngoc Linh Mountain system. Remnants of primary and secondary evergreen broad-leaved forest on granite and shale at elevation of about 1100 m a.s.l. Epiphyte on tall tree1100 m alt. Not common.

02 May 2025, L. Averyanov, N.V. Canh, T. Maisak, AL4121.

Flowered in culture 21 August 2025, L. Averyanov, AL4121.

Orchidaceae

Eria amica Rchb.f.?

Thanh Hoa Province, Quan Hoa District, Nam Dong Community, Ban Lo Village, Ke Stream, the territory of Nam Dong National Reserve around point 22º25’46’’N 105º36’58’E. Primary and old secondary broad-leaved evergreen humid forest on very steep rocky slopes on solid, marble-like highly eroded limestone at elevation 900–1200 m a.s.l. on steep rocky slope. Clustering epiphyte and lithophyte on rocky mossy mountain top. Not rare.

04 May 2024, Nguyen Huu Cuong, Nguyen Van Ly, L. Averyanov, T. Maisak, AL3052.

Orchidaceae

Appendicula hexandra (J.Koenig) J.J.Sm.⁣

Lam Dong Province, Bao Lam District. Secondary grassy coniferous woodlands and open scrub on thin degraded volcanic lateritic rocky soil at elevation about 900 m a.s.l.

30 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3498.

Flowered in cultivation 21 August 2025, L. Averyanov, AL3498.

FLORA OF VIETNAM

Convallariaceae

Aspidistra neglecta

Tuyen Quang prov., Na Hang distr., Sinh Long commune, Nam Duong vill., Phim Panh Mt. Evergreen broad-leaf forest in limestone. Around point 22°034'28.1"N, 105°22'04.1"E, 539 m a.s.l. herb, occasional. 6 Oct, 2017 CPC 8341 AL 396b N.T.Hiep, N.S.Khang, C.Q.Ngan, N.Q.Hieu.

Orchidaceae

Oberonia multidentata Aver.

The border of Lam Dong and Khanh Hoa Provinces, Khanh Le Pass (Hon Giao Pass) area, primary humid evergreen broad-leaved cloud forest on granite and gneiss at elevation 1750-1850 m a.s.l. Epiphyte. 19 April 2026, Truong Quang Cuong, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3859.

Fl. in culture on 27 August 2025, flowers orange, L.Averyanov, AL3859.

Pandanaceae

Pandanus (Mainstera) sp.

Dak Lak Province, M’Drak District, W foothills of Chu Mu Mt., dry secondary and remnants of primary evergreen broad-leaved forest on granite on steep slopes along rocky river at elevation 600-650 m a.s.l. Palm-like tree to 8 m tall in shady wet place on stream slope. Common.

25 April 2025, L. Averyanov, N.V. Canh, T. Maisak, AL3959.

Vitaceae

Tetrastigma sp.⁣

Lam Dong Province, Dalat Town Area, old secondary broad-leaved evergreen forest on granite at elevation about 1600 m a.s.l. April 2025, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3899.

Rafflesiaceae

Sapria himalayana Griff.

Lam Dong Province, Dalat Town Area, old secondary broad-leaved evergreen forest on granite at elevation about 1600 m a.s.l. Ground root achlorophyllous stemless parasite on Tetrastigma planicaulis. Flowers red-purple, tepals dotted with white. Rare. 21 April 2025, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3899.

Vitaceae

Tetrastigma sp.⁣

Lam Dong Province, Dalat Town Area, old secondary broad-leaved evergreen forest on granite at elevation about 1600 m a.s.l. 21 April 2025, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3899.

Rafflesiaceae

Sapria himalayana Griff.

Lam Dong Province, Dalat Town Area, old secondary broad-leaved evergreen forest on granite at elevation about 1600 m a.s.l. Ground root achlorophyllous stemless parasite on Tetrastigma planicaulis. Flowers red-purple, tepals dotted with white. Rare. 21 April 2025, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3899.

Orchidaceae

Bulbophyllum moniliforme?

Kon Tum Province, Kon Plong District, Mang Den Town area, remnants of humid primary evergreen broad-leaved and mixed forest (with Pinus dalatensis, Dacrydium and Dacrycarpus) on very steep mountain slopes composed with granite and silicate shale at elevation of about 1200 m a.s.l. Miniature creeping epiphyte on mossy trees. Rare.

28 April 2025, L. Averyanov, N.V. Canh, T. Maisak, AL4042.

Orchidaceae

Schoenorchis seidenfadenii Pradhan

Gia Lai Province, Dak Doa District, obtained from local collectors. 16 July 2024, Nguyen Van Canh, AL3274.

Orchidaceae

Didymoplexiopsis khiriwongensis Seidenf.

Quang Nam Province, Nong Sơn District, Elephant Species and Habitat Conservation Area. Secondary dry evergreen broad-leaved forest on step mountain slopes composed by sandstone at elevation of about 200 m a.s.l. Terrestrial achlorophyllous tuberiferous herb on stream slopes in open dry forest with bamboo. Flowers white. Rare

04 May 2025, L. Averyanov, N.V. Canh, T. Maisak, AL4146.

Convallariaceae

Aspidistra sp.

Dak Lak Province, Krong Bong District, Yang Hanh mt., evergreen broad-leaved forest at elevation of about 1000 m a.s.l.

02.2025. Nguyen Van Canh, AL3774. Living plant only and herbarium with fruits.

Polypodiaceae s.l.

Small 1

The border of Lam Dong and Khanh Hoa Provinces, Khanh Le Pass (Hon Giao Pass) area, primary humid evergreen broad-leaved cloud forest on granite and gneiss at elevation 1750-1850 m a.s.l. Epiphyte on mossy trees in very humid slope along stream. Common. 19 April 2026, Truong Quang Cuong, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3848.

Orchidaceae

Dendrobium parciflorum Lindl.

Lam Dong Province, Bao Lam District. Secondary grassy coniferous woodlands and open scrub on thin degraded volcanic lateritic rocky soil at elevation about 900 m a.s.l.

30 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3494.

Flowered in cultivation 25 July, L. Averyanov, T. Maisak, AL3494.