Herbarium LE - Specimens

Lamiaceae

Orthosiphon sp.?

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Erect lithophytic herb to 0.5 m tall among open stream rocks. Flowers light violet. Not common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4607.

All photos © - L. Averyanov

Oleaceae

Jasminum sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Scandent shrub of semi-woody vine to 3 m long in open place along road. Flowers white. Common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4606.

All photos © - L. Averyanov

Potamogetonaceae

Aponogeton robinsonii A.Camus?

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Aquatic herb with immersed linear leaves in shallow slow water of rocky forest stream. Flowers white. Locally common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4605.

All photos © - L. Averyanov

Acanthaceae

Acanthus sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Creeping ascending herb on shady wet stream slope. Flowers white. Locally very common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4604.

All photos © - L. Averyanov

Rubiaceae

Psychotria langbianensis Wernham

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial herb to 1 m tall in rather open place along trail. Flowers white. Not rare.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4603.

All photos © - L. Averyanov

Selaginellaceae

Selaginella sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Wetland thicket in marshland depression with domination of Pandanus and Livistona along stream. Terrestrial scandent herb to 1.5 m long in wet open place. Common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4602.

All photos © - L. Averyanov

Rubiaceae

Brachytome walichii Hook.f?

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Treelet or shrub to 2 m tall in rather open place along trail. Flowers yellow to yellow orange. Occasional.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4601.

All photos © - L. Averyanov

Selaginellaceae

Selaginella sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Wetland thicket in marshland depression with domination of Pandanus and Livistona along stream. Terrestrial scandent herb to 2 m long in wet open place. Common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4600.

All photos © - L. Averyanov

Orchidaceae

Tainia cordifolia Hook.f.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial prostrate herb in shady place. Flowers yellowish brown, lip white with yellow apex. Not common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4599.

All photos © - L. Averyanov

Selaginellaceae

Selaginella sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial herb to 0.5 m tall on shady stream slope. Common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4598.

All photos © - L. Averyanov

Orchidaceae

Pholidota imbricata Hook

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Clustering epiphyte on tall tree in open place along stream rocks. Occasional.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4597.

All photos © - L. Averyanov

Convallariaceae

Aspidistra cylindrica?

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial rhizomatous herb with thick hypogeous rhizome on shady stream slopy. Occasional.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4596.

All photos © - L. Averyanov

Selaginellaceae

Selaginella sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Wetland thicket in marshland depression with domination of Pandanus and Livistona along stream. Terrestrial sedge to 1 m tall on shady slope. Common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4595.

All photos © - L. Averyanov

Cyperaceae

Carex sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’00N 107º46’00E. Wetland thicket in marshland depression with domination of Pandanus and Livistona along stream. Terrestrial sedge in open place. Common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4594.

All photos © - L. Averyanov

Selaginellaceae

Selaginella sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial herb on shady stream slope. Not common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4593.

All photos © - L. Averyanov

Orchidaceae

Paphiopedilum pittchianum sp. nov.

Plant originated from Indonesia, Maluku Province, Greater Lai Island, without exact location.

Photos and herbarium specimen prepared from cultivated plants in Thailand, BioBeaM Garden,

10 February 2026, Pittch Suntichitrungruang, AL5034.

All photos © Pittch Suntichitrungruang

Orchidaceae

Paphiopedilum × lathamianum hybrid. nov.

Plant originated from northern Myanmar, without exact location.

Photos and herbarium specimen prepared from cultivated plants in Thailand, BioBeaM Garden,

10 February 2026, Pittch Suntichitrungruang, AL5036.

All photos © Pittch Suntichitrungruang

Orchidaceae

Paphiopedilum × grussianum hybrid. nov.

Plant originated from Vietnam, without exact location.

Photos and herbarium specimen prepared from cultivated plants in Thailand, BioBeaM Garden,

10 February 2026, Pittch Suntichitrungruang, AL5035.

All photos © Pittch Suntichitrungruang

Orchidaceae

Aerides odorata Lour.

Thanh Hoa Province, plant obtained from local collectors and cultivated in garden. Collected by Nguyen Van Canh on July 2025, AL4555.

All photos © - L. Averyanov & Truong Ba Vuong

Acanthaceae

Phlogacanthus pulcherrimus T.Anderson⁣

Dak Lac Province, Yok Don National Park, Dipterocarp forest at elevation of about 200 m a.s.l., flowers dirty purple. Plant cultivated in Canh Garden in Buon Ma Thuot Town. Collected by on 26 March 2026, Nguyen Van Canh, L. Averyanov, Truong Ba Vuong, T. Maisak, AL4554.

All photos © - L. Averyanov

Acanthaceae

Phlogacanthus turgidus (Fua ex Hook.f.) Lindau⁣

Plant obtained from local collectors in Son La Province in broad-leaved evergreen forest at about 700-800 m a.s.l. Flowers white with tube yellow inside.

Collected in Nguyen Van Canh Garden on 26 March 2026, Nguyen Van Canh, L. Averyanov, Truong Ba Vuong, T. Maisak, AL4553.

All photos © - L. Averyanov

Euphorbiaceae

Breynia phuongiana N.K.T.Tram, Vuong, Aver. & V.C.Nguyen

Originally collected in Dak Lak Province, Ea H'leo District, evergreen forest at elevation of about 700 m a.s.l., 15 Feb. 2023, Nguyen Van Canh, AL 2456

Clonotype. Collected in Nguyen Van Canh Garden on 26 March 2026, Nguyen Van Canh, L. Averyanov, Truong Ba Vuong, T. Maisak, AL4552.

All photos © - L. Averyanov

Convallariaceae

Peliosanthes graminea Ridl.?

Dac Lak Province, Lac District, Evergreen broad-leaved forest on very steep slopes along stream. Not common. Flowers dark violet.

Collected in Nguyen Van Canh Garden on 26 March 2026, Nguyen Van Canh, L. Averyanov, Truong Ba Vuong, T. Maisak, AL4551.

All photos © - L. Averyanov

Convallariaceae

Aspidistra opaca Tillich

Dac Lak Province, Lac District, Evergreen broad-leaved forest on very steep slopes along stream. Locally very common. Flowers dark violet, stigma white.

Collected in Nguyen Van Canh Garden on 26 March 2026, Nguyen Van Canh, L. Averyanov, Truong Ba Vuong, T. Maisak, AL4550.

All photos © - L. Averyanov

Convallariaceae

Peliosanthes griffithii Baker?

Cultivated plant originated from Dak Nong Province, Nam Nung Nature Reserve, evergreen broad-leaved forest at elevation of about 800 m a.s.l.

All photos © - L. Averyanov

Convallariaceae

Peliosanthes “subsessiliflora” sp. new.? (P. micrantha aff.)

Cultivated plant originated from Binh Dinh Province, Van Canh District, lowland forest along stream, at elevation of about 300 m a.s.l., locally common.

Collected in Nguyen Van Canh Garden on 26 March 2026, Nguyen Van Canh, L. Averyanov, Truong Ba Vuong, T. Maisak, AL4548. Flowers dark violet, fruits glossy blue.

All photos © - L. Averyanov & Truong Ba Vuong

Convallariaceae

Peliosanthes weberi (L.Rodr.) N.Tanaka

Cultivated plant originated from Dac Nong Province, Dak Mil District, evergreen forest mixed with dipterocarp forest at elevation of about 300 m a.s.l., common,

Collected in Nguyen Van Canh Garden on 26 March 2026, Nguyen Van Canh, L. Averyanov, Truong Ba Vuong, T. Maisak, AL4547. Flowers light green.

All photos © - L. Averyanov & Truong Ba Vuong

Ericaceae (Monotropaceae)

Monotropa callistoma Ze Wei, Li J.Liu & Bing Liu

Plant obtained from local collectors in Lao Cai Province, Bat Xat District, evergreen broad-leaved forest at elevation of about 1800 m a.s.l. Not common. Flowers white with yellow mouth.

26 March 2026, Nguyen Van Canh, AL4546.

All photos © - L. Averyanov & Truong Ba Vuong

Convallariaceae

Ophiopogon muongnhensis K.S.Nguyen, N.Tanaka & Aver.⁣

Dien Bien Prov., Muong Nhe Distr., Muong Nhe Natural Reserve, Sin Thau Municipality, A Pa Chai Village around point about N22º22’51’’ E102º14’26”. Old humid secondary evergreen broad-leaved submontane forest of steep mountain slopes composed with sandstone at elevations 700–800 m a.s.l. along small stream. Terrestrial herb with stem to 0.8 m long on shady slope. Very common.

15 May 2022, L. Averyanov, Tran Huy Thai, Khang Sinh Nguyen, Cuong Huu Nguyen, T. Maisak, Bac Cam Khoa, VR1641a.

Flowered in culture on 12 Mat 2026, L. Averyanov, VR1641a, garden number GN 22174.

All photos © - L. Averyanov

Convallariaceae

Ophiopogon muongnhensis K.S.Nguyen, N.Tanaka & Aver.

Dien Bien Prov., Muong Nhe Distr., Muong Nhe Natural Reserve, Sin Thau Municipality, A Pa Chai Village around point about N22º22’51’’ E102º14’26”. Old humid secondary evergreen broad-leaved submontane forest of steep mountain slopes composed with sandstone at elevations 700–800 m a.s.l. along small stream. Terrestrial herb with stem to 0.8 m long on shady slope. Very common.

15 May 2022, L. Averyanov, Tran Huy Thai, Khang Sinh Nguyen, Cuong Huu Nguyen, T. Maisak, Bac Cam Khoa, VR1641a.

Flowered in culture on 12 Mat 2026, L. Averyanov, VR1641a, garden number GN 22174.

All photos © - L. Averyanov

Orchidaceae

Thrixspermum flammeum Toolmal, Schuit. & Culham

Lam Dong Province, Bao Lam District. Secondary grassy coniferous woodlands and open scrub on thin degraded volcanic lateritic rocky soil at elevation about 900 m a.s.l. Epiphyte.

30 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3503.

Flowered in cultivation 12 May 2026, L. Averyanov, AL3503.

All photos © - L. Averyanov

Convallariaceae

Peliosanthes sp.

Ninh Thuan Province, Ninh Hai District, Vinh Hy Townlet. Very dry evergreen primary and secondary short-tall wind-formed coastal forest on very steep sea faced sandstone rocky slopes at elevation of 200–250 m a.s.l. Terrestrial herb in shady rocky places. Flowers dark violet. Not common.

16 May 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3235.

Flowered in culture on 12 May 2026, L. Averyanov, AL3235.

All photos © - L. Averyanov

Convallariaceae

Peliosanthes sp.

Gia Lai Province, G’Bang District, Kon Chu Rang Nature Reserve, around point 14º30.324´ N 108º29.762´ E. Primary evergreen broad-leaved forest with Dacrycarpus and Dacrydium at elevation about 1200 m a.s.l. Terrestrial clustering herb in shady place. Leaves grassy green. Not rare.

21 November 2023, L. Averyanov, Nguyen Van Canh, T. Maisak, AL2741 (living plant only).

Flowered in culture on 12 May 2026, L. Averyanov, AL2741 (garden number 2465).

All photos © - L. Averyanov

Convallariaceae

Peliosanthes sp.

Gia Lai Province, G’Bang District, Kon Chu Rang Nature Reserve, around point 14º30.324´ N 108º29.762´ E. Primary evergreen broad-leaved forest with Dacrycarpus and Dacrydium at elevation about 1200 m a.s.l. Terrestrial clustering herb in shady place. Leaves grassy green. Not rare.

21 November 2023, L. Averyanov, Nguyen Van Canh, T. Maisak, AL2741A (living plant only).

Flowered in culture on 12 May 2026, L. Averyanov, AL2741A (garden number 2465).

All photos © - L. Averyanov

Convallariaceae

Peliosanthes sp.

Lam Dong Province, Di Linh District, Brah Yang Mountain, primary and old secondary evergreen broad-leaved and mixed forest (with Pinus khasya) on granite around point 11º27.669ʹ N 108º03.543ʹ E at elevation of 1000-1100 m a.s.l. Terrestrial rhizomatous herb on shady rocky slope. Fruits ovoid to ellipsoid, glossy blue. Locally common.

14 November 2023, L. Averyanov, Nguyen Van Canh, T. Maisak, AL2685 (photo and seeds only).

Flowered in cultivation on 12 May 2026, L. Averyanov, AL2685.

All photos © - L. Averyanov

Orchidaceae

Cymbidium nanulum Y.S.Wu & S.C.Chen

Living plant obtained at 25 Nov. 2022, Nguyen Van Canh, L. Averyanov, T. Maisak, AL 1696.

Plant originated from Dak Lak Province, Ea Hleo District, Yok Don NP, Dipterocarp forest at elevation 200–300 m a.s.l., along stream, locally common, 2020, Nguyen Van Canh.

(also common in Gia Lai and Dak Nong provinces).

Flowered in culture on 12 May 2026, L. Averyanov, AL1696.

All photos © - L. Averyanov

Melastomataceae

Sonerila sp. (“purpurifiolia”?)

Lai Chau Province, Muong Te District, Mu Ca Commune. Highly degraded secondary evergreen broad-leaved forest with bamboo along rocky river on stratified shale and limestone at elevation of about 500 m a.s.l. around point 22º24’42’’N 102º24’35’’E. Terrestrial herb to 6 cm tall on shady mossy steep roadcut. Petals, filaments and style purple, anthers pale yellow. Not common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5007.

All photos © - L. Averyanov

Auf Ebenen