Herbarium LE - Specimens

Arecaceae

Calamus sp.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Secondary broad-leaved evergreen forest with bamboo on rocky sandstone slopes at elevation 10–50 m a.s.l.

Climbing and ascending woody vine to 5-6 m long on very steep slope of small rocky stream. Fruits greenish. Common.

21 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4428 (NHC221).

Sapotaceae

Planchonella obovata (R.Br.) Pierre?

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Young secondary broad-leaved evergreen forest on rocky sandstone slope along seashore at 0–5 m a.s.l.

Treelet or shrub to 4 m tall in open place in sublittoral not flooded zone along rocky seashore. Flower buds greenish. Not common.

20 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4406 (NHC199).

Rhizophoraceae

Ceriops tagal (Perr.) C.B.Rob.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Young secondary broad-leaved evergreen forest on rocky sandstone slope along seashore at 0–5 m a.s.l.

Treelet or shrub to 2 m tall among mangrove shrubs along rocky seashore. Flower buds yellowish-green. Common.

20 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4405 (NHC198).

Verbenacae

Lumnitzera sp.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Young secondary broad-leaved evergreen forest on rocky sandstone slope along seashore at 0–5 m a.s.l.

Treelet or shrub to 2 m tall in open place in sublittoral not flooded zone along rocky seashore. Flower buds greenish. Common.

20 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4404 (NHC197).

Loranthaceae

Scurrula ferruginea (Roxb. ex Jack) Danser?

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Young secondary broad-leaved evergreen forest on rocky sandstone slope along seashore at 0–5 m a.s.l.

Parasitic shrub with erect shoot to 1 m long in canopies of tall tree (Litsea). Flowers outside brownish-yellow, inside dark violet. Not common.

20 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4403 (NHC196).

Smilacaceae

Smilax sp.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Young secondary broad-leaved evergreen forest on rocky sandstone slope along seashore at 0–5 m a.s.l.

Semi-woody vine to 2-3 m long among dense shrubs in sublittoral not flooded zone along rocky seashore. Flowers light green. Not common.

20 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4402 (NHC195).

Rubiaceae

Mussaenda sp.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Young secondary broad-leaved evergreen forest on rocky sandstone slope along seashore at 0–5 m a.s.l.

Woody vine to 5 m long in dense shrubs in sublittoral not flooded zone along rocky seashore. Fruits dark green. Not common.

20 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4401 (NHC194).

Verbenacae

Lumnitzera sp.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Young secondary broad-leaved evergreen forest on rocky sandstone slope along seashore at 0–5 m a.s.l.

Treelet or shrub to 2 m tall in open place in sublittoral not flooded zone along rocky seashore. Flower buds green. Common.

20 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4400 (NHC193).

Arecaceae

Korthalsia sp.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Young secondary broad-leaved evergreen forest on rocky sandstone slope along seashore at 0–5 m a.s.l.

Woody vine to 5 m long in sublittoral not flooded zone along rocky seashore. Flowers dull yellow. Very common.

20 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4399 (NHC192).

Euphorbiaceae

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Young secondary broad-leaved evergreen forest on rocky sandstone slope along seashore at 0–5 m a.s.l.

Undershrub to 1.5 m in shady place in sublittoral not flooded zone along rocky seashore. Flowers dull yellowish. Not rare.

20 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4398 (NHC191).

Asclepiadaceae

Cerbera manghas L.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Young secondary broad-leaved evergreen forest on rocky sandstone slope along seashore at 0–5 m a.s.l.

Treelet 4-6 m tall in sublittoral not flooded zone along rocky seashore. Flowers white, in center reddish. Common.

20 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4397 (NHC190).

Convolvulaceae

Ipomoea sp.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Young secondary broad-leaved evergreen forest on rocky sandstone slope along seashore at 0–5 m a.s.l.

Herbaceous or semi-woody vine to 8 m long in sublittoral not flooded zone along rocky seashore. Flower buds yellowish. Common.

20 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4396 (NHC189).

Euphorbiaceae

(Dark green leaves, male only)

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Young secondary broad-leaved evergreen forest on rocky sandstone slope along seashore at 0–5 m a.s.l.

Sublittoral evergreen treelet or shrub to 2 m tall along rocky seashore in not flooded zone with dark green leaves. Male flowers yellowish-green. Very common.

19 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4395 (NHC188).

Euphorbiaceae

(Red leaves, male, female)

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Young secondary broad-leaved evergreen forest on rocky sandstone slope along seashore at 0–5 m a.s.l.

Sublittoral semideciduous treelet or shrub to 2 m tall along rocky seashore in not flooded zone with old leaves orange-red to bright red. Male young flower buds and fruits light green. Very common.

19 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4394 (NHC187).

Gnetaceae

Gnetum gnemon L.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Young secondary broad-leaved evergreen forest on rocky sandstone slope along seashore at 0–5 m a.s.l.

Tree to 15 m tall and 30 cm DBH. Occasional, commonly at elevation 50-150 m a.s.l.

19 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4393 (NHC186).

Moraceae

Artocarpus sp.

Quang Ninh Province, Van Don District, Minh Chau Commune, Bai Tu Long National Park, Ba Mun protection sector|, Ba Mun Island, around point 21º04’33’’N 107º35’28’’E. Young secondary broad-leaved evergreen forest on rocky sandstone slope along seashore at 0–5 m a.s.l.

Tree about 10 m tall. Young fruits white to paly yellow, edible. Not rare.

19 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4392 (NHC185).

Rhizophoraceae

Bruguiera gymnorhiza (L.) Lam. ex Savigny

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Littoral mangrove shrub or treelet to 2.5 m tall in flooded littoral zone. Flower. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4352 (NHC152).

Gesneriaceae

Primulina drakei (B.L.Burtt) Mich.Möller & A.Weber

Quang Ninh Province, Van Don District, Minh Chuu Commune, Bai Tu Long National Park, Ba Mun protection sector, Mang Ha Island, around point 21º05’92’’N 107º34’99’’E. Open broad-leaved evergreen forest and scrub on steep slopes and cliffs composed by solid highly eroded limestone at elevation about 20 m a.s.l. Rosulate herb with succulent stem to 0.5 m tall on exposed cliff. Flowers dark purple-violet. Locally common.

18 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4378 (NHC171).

Apocynaceae??

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Littoral mangrove shrub or treelet with milky juice to 3 m tall along flooded littoral zone. Flower buds yellowish green. Very common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4351 (NHC151).

Verbenaceae

Lumnitzera sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Littoral mangrove shrub or treelet to 2 m tall in flooded littoral zone. Flower buds yellowish green. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4350 (NHC150).

Balsaminaceae

Impatiens sp. (“yellow”)

Kon Tum Province, Dak Gley District, W slopes of Ngoc Linh Mountain system. Remnants of primary and secondary evergreen broad-leaved forest on granite and shale at elevation of about 1100 m a.s.l. Erect terrestrial and lithophytic herb to 1 m tall among rocks along shady stream canyon. Flowers bright yellow. Occasional.

02 May 2025, L. Averyanov, N.V. Canh, T. Maisak, AL4111.

Gesneriaceae

Aeschynanthus sp. (“bright red”)

Kon Tum Province, Dak Gley District, W slopes of Ngoc Linh Mountain system. Remnants of primary and secondary evergreen broad-leaved forest on granite and shale at elevation of about 1800 m a.s.l. Undershrub or scandent semiwoody vine to 1.5 m long in rather open place among Betula alnoides saplings along road, on steep slope. Flowers bright red. Occasional.

02 May 2025, L. Averyanov, N.V. Canh, T. Maisak, AL4110.

Ericaceae

Rhododendron sp. (“white pinkish medium flower”)

Kon Tum Province, Dak Gley District, W slopes of Ngoc Linh Mountain system. Remnants of primary and secondary evergreen broad-leaved forest on granite and shale at elevation of about 1800 m a.s.l. Shrub to 1.2 m tall in rather open place among Betula alnoides saplings along road. Flowers white, with pinkish tint in center. Locally common.

02 May 2025, L. Averyanov, N.V. Canh, T. Maisak, AL4109.

Ericaceae

Rhododendron sp. (“white yellowish big flower”)

Kon Tum Province, Dak Gley District, W slopes of Ngoc Linh Mountain system. Remnants of primary and secondary evergreen broad-leaved forest on granite and shale at elevation of about 1800 m a.s.l. Shrub to 2 m tall in rather open place among Betula alnoides saplings along road. Flowers light yellowish at opening, later almost pure white. Locally common.

02 May 2025, L. Averyanov, N.V. Canh, T. Maisak, AL4108.

Ericaceae

Rhododendron sp. (“white yellowish big flower”)

Kon Tum Province, Dak Gley District, W slopes of Ngoc Linh Mountain system. Remnants of primary and secondary evergreen broad-leaved forest on granite and shale at elevation of about 1800 m a.s.l. Shrub to 2 m tall in rather open place among Betula alnoides saplings along road. Flowers light yellowish at opening, later almost pure white. Locally common.

02 May 2025, L. Averyanov, N.V. Canh, T. Maisak, AL4108.

Rosaceae

Docynia indica (Wall.) Decne.

Kon Tum Province, Dak Gley District, W slopes of Ngoc Linh Mountain system. Remnants of primary and secondary evergreen broad-leaved forest on granite and shale at elevation about 1200 m a.s.l. Tree about12 m tall in open place along rocky stream. Not common.

02 May 2025, L. Averyanov, N.V. Canh, T. Maisak, AL4106.

Balsaminaceae

Impatiens sp.

Kon Tum Province, Kon Plong District, Mang Den Town area, Pa Sy Waterfall, secondary dry evergreen broad-leaved forest on steep river slopes composed with lava rocks and silicate shale at elevation of about 1100 m a.s.l. Lithophilic herb to 0.5 m tall on wet waterfall cliffs. Flowers pink purple. Locally common.

30 April 2025, L. Averyanov, N.V. Canh, T. Maisak, AL4069.

Theaceae

Pyrenaria jonquieriana Laness

Kon Tum Province, Kon Plong District, Mang Den Town area, Khu Ba Waterfall, secondary dry evergreen broad-leaved forest on steep river slopes composed with silicate shale at elevation of about 900 m a.s.l. Treelet to 4 m tall. Flowers white, anthers yellow. Not common.

30 April 2025, L. Averyanov, N.V. Canh, T. Maisak, AL4073.

Theaceae

Pyrenaria jonquieriana Laness

Kon Tum Province, Kon Plong District, Mang Den Town area, Khu Ba Waterfall, secondary dry evergreen broad-leaved forest on steep river slopes composed with silicate shale at elevation of about 900 m a.s.l. Treelet to 4 m tall. Flowers white, anthers yellow. Not common.

30 April 2025, L. Averyanov, N.V. Canh, T. Maisak, AL4073.

Urticaceae

Dendrocnide sp.

The border on Kon Tum and Quang Nam Provinces along the road Da Nang – Dak Gley. Secondary broad-leaved evergreen forest on very steep slopes composed by silicate shale at elevation of 800–1100 m a.s.l. Tree about 5-6 m tall on side of rocky stream among open rocks. Leaves strongly urticate. Fruits white juicy sweet edible. Not common.

03 May 2025, L. Averyanov, N.V. Canh, T. Maisak, AL4132.

Melastomataceae

Perilimnastes setipetiola J.H.Dai, T.V.Do & Ying Liu

The border of Lam Dong and Khanh Hoa Provinces, Khanh Le Pass (Hon Giao Pass) area, primary humid evergreen broad-leaved cloud forest on granite and gneiss at elevation 1750-1850 m a.s.l. Terrestrial herb or undershrub to 1.5 m tall on wet rocky slope. Flowers white with pink tint. Locally very common. 19 April 2026, Truong Quang Cuong, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3847.

Araliaceae

Schefflera langbianensis Q.C.Truong⁣ ⟨not published name⟩

det. Truong Quang Cưong at 19.04.2025

The border of Lam Dong and Khanh Hoa Provinces, Khanh Le Pass (Hon Giao Pass) area, primary humid evergreen broad-leaved cloud forest on granite and gneiss at elevation 1750-1850 m a.s.l. Tree to 6 m tall along ridge edge. Common. 19 April 2025, Truong Quang Cuong, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3839.

Clusiaceae

Calophyllum sp.

The border of Lam Dong and Khanh Hoa Provinces, Khanh Le Pass (Hon Giao Pass) area, primary humid evergreen broad-leaved cloud forest on granite and gneiss at elevation 1750-1850 m a.s.l. Tree to 10 m tall along ridge edge. Flowers white. Not common. 19 April 2025, Truong Quang Cuong, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3836.

Meliosmaceae

Meliosma sp.

The border of Lam Dong and Khanh Hoa Provinces, Khanh Le Pass (Hon Giao Pass) area, primary humid evergreen broad-leaved cloud forest on granite and gneiss at elevation 1750-1850 m a.s.l. Shrub to 2.5 m tall. Flowers light violet. Not common. 19 April 2025, Truong Quang Cuong, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3835.

Rubiaceae

Lasianthus sp.

The border of Lam Dong and Khanh Hoa Provinces, Khanh Le Pass (Hon Giao Pass) area, primary humid evergreen broad-leaved cloud forest on granite and gneiss at elevation 1750-1850 m a.s.l. Shrub to 1.5 m tall. Flowers white with violet tint, sweet fragrant. Occasional. 19 April 2025, Truong Quang Cuong, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3831.

Myricaceae

Myrica amplexicaulis Q.C.Truong⁣ ⟨not published name⟩

det. Truong Quang Cưong at 19.04.2025

The border of Lam Dong and Khanh Hoa Provinces, Khanh Le Pass (Hon Giao Pass) area, primary humid evergreen broad-leaved cloud forest on granite and gneiss at elevation 1750-1850 m a.s.l. Treelet to 4 m tall. Not rare. 19 April 2025, Truong Quang Cuong, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3830.

Poaceae

Arundinaria sp.?

The border of Lam Dong and Khanh Hoa Provinces, Khanh Le Pass (Hon Giao Pass) area, primary humid evergreen broad-leaved cloud forest on granite and gneiss at elevation 1750-1850 m a.s.l. Woody grass with slender stems to 3 m long along ridge edge. Common. 19 April 2025, Truong Quang Cuong, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3837.

Lorantaceae

Helixanthera sp.?

The border of Lam Dong and Khanh Hoa Provinces, Khanh Le Pass (Hon Giao Pass) area, primary humid evergreen broad-leaved cloud forest on granite and gneiss at elevation 1750-1850 m a.s.l. Semi parasitic shrub in canopies of tall trees. Flowers bright red. Not common. 19 April 2025, Truong Quang Cuong, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3834.