Herbarium LE - Specimens

All photos © L. Averyanov & K.S. Nguyen

KOMAROV BOTANICAL INSTITUTE PLANTS OF BOTANICAL GARDEN Tupistra tupistroides (Kunth) Dandy ( = Macrostigma tupistroides Kunth, 1848, Index seminum horto botanico Berolinensi anno 1848 collectorum. Species novae horti regii botanici Berolinensis (1848): 12) Plant of unknown origin obtained from Munich at around 2000–2005 year. Most probably, clonotype. Flowered in 30 November 2018. Flowers outside white, perigone lobes dark purple-brown, tube inside with 6 purple-brown longitudinal stripes, anthers white, style white, stigma dark purple-violet. 30 Nov. 2018, L. Averyanov, AL 463.

FLORA OF LAOS Hin Nam No Nature Protected Area Annonaceae Goniothalamus sp. 1 Laos, Khammouane prov., Boualapha distr., Nong Seng village, southern sandstone slopes of Phou Chuang (Poujeuang) Mt. around point 17°34'13.3"N 105°47'57.5"E. Primary evergreen broad-leaved dry forest at elev. 700-1000 m a.s.l. Caulifloric treelet 3-4 m tall. Flowers pale yellow-greenish to dull orange. Not common. 3 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 543.

FLORA OF LAOS Hin Nam No Nature Protected Area Annonaceae Anaxagorea sp. ? Laos, Khammoune prov., Boualapha distr., Vang Maner village, Palai mountain area, around point 17°30'13.6"N, 105°49'23.9"E, 520 m a.s.l. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 300-500 m a.s.l. Undershrub 1.5 m tall. Flowers white. Occasional. 8 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 761.

FLORA OF LAOS Hin Nam No Nature Protected Area Annonaceae Miliusa velutina (Dunal) Hook. f. & Thomson Laos, Khammoune prov., Boualapha distr., Nong Seng village, Chanh Daeng mountain, around point 17°31'11.1"N, 105°46'41.4"E. Secondary dry evergreen and semideciduous broad-leaved forest on sandy alluvial flat valley along river between karstic limestone hills at about 180 m a.s.l. Tree 15-20 m tall in open place. Fruits green. Occasional. 11 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 1048.

FLORA OF LAOS Hin Nam No Nature Protected Area Annonaceae Miliusa velutina (Dunal) Hook. f. & Thomson Laos, Khammoune prov., Boualapha distr., Nong Seng village, Chanh Daeng mountain, around point 17°31'11.1"N, 105°46'41.4"E. Secondary dry evergreen and semideciduous broad-leaved forest on sandy alluvial flat valley along river between karstic limestone hills at about 180 m a.s.l. Tree 15-20 m tall in open place. Fruits green. Occasional. 11 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 1048.

FLORA OF LAOS Nakai-Nam Theun Protected Area Annonaceaea Melodorum fruticosum Lour. ? Laos, Khammoune prov., Boualapa distr., Langkhang village, steep southern slopes of sandstone ridge allied to the road from the north between points 17°34'02.2"N 105°42'42.6"E, 200 m and 17°34'51.1"N 105°43'01.2"E, 535 m a.s.l. Primary lowland semideciduous and evergreen dry forest on very steep rocky slopes. Treelet about 6 m tall in rather open place along trail. Flowers dull yellow-greenish. Occasional. 15 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, AL 1074.

FLORA OF LAOS Hin Nam No Nature Protected Area Annonaceae Mitrella sp. ? Laos, Khammouane prov., Boualapha distr., Nong Seng village, southern sandstone slopes of Phou Chuang (Poujeuang) Mt. around point 17°34'13.3"N 105°47'57.5"E. Primary evergreen broad-leaved dry forest at elev. 700-1000 m a.s.l. Treelet. Flowers greenish to yellow. Occasional. 3 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 580.

FLORA OF LAOS Hin Nam No Nature Protected Area Annonaceae Goniothalamus macrophyllus (Blume) Hook.f. & Thomson ? Laos, Khammouane prov., Boualapha distr., Nong Seng village, southern sandstone slopes at summit of Phou Chuang (Poujeuang) Mt. around point 17°35'18.1"N, 105°47'47.5"E. Primary evergreen mixed and coniferous dense wind-formed forest at elev. 1000-1400 m a.s.l. Shrub about 2 m tall on mountain top. Flowers dull yellowish-orange. Common. 5 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 670a.

FLORA OF LAOS Hin Nam No Nature Protected Area Annonaceae Polyalthia simiarum (Hook. f. & Thomson) Benth. ( = Monoon simiarum (Hook. f. & Thomson) B. Xue & R. M. K. Saunders) Laos, Khammoune prov., Boualapha distr., Vang Maner village, around point 17°29'39.3"N, 105°50'24.6"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 350-430 m a.s.l. Shrub or treelet 3-4 m tall on shady rocky slope. Flowers pale yellow. Occasional. 9 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 845.

FLORA OF LAOS Hin Nam No Nature Protected Area Annonaceae Polyalthia cinnamomea Hook.f. & Thomson Laos, Khammoune prov., Boualapha distr., Vang Maner village, around point 17°29'39.3"N, 105°50'24.6"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 350-430 m a.s.l. Shrub or treelet 3-4 m tall. Flowers light pale yellow to almost white. Not rare. 9 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 843.

FLORA OF LAOS Hin Nam No Nature Protected Area Annonaceae Polyalthia sp. 3 Laos, Khammoune prov., Boualapha distr., Nong Seng village, Vang Phae Lay mountain, around point 17°31'02.7"N, 105°47'17.2"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 300-530 m a.s.l. Treelet or shrub on shady rocky slope. Flowers yellowish, internal tepals at apical half red striped. Occasional. 11 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 947.

FLORA OF LAOS Hin Nam No Nature Protected Area Annonaceae Goniothalamus sp. 2 Laos, Khammoune prov., Boualapha distr., Vang Maner village, around point 17°29'39.3"N, 105°50'24.6"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 350-430 m a.s.l. Shrub or treelet on shady rocky slope. Flowers dull yellow. Occasional. 9 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 837.

FLORA OF LAOS Hin Nam No Nature Protected Area Annonaceae Mitrephora keithii Ridl. ? Laos, Khammoune prov., Boualapha distr., Nong Seng village, Vang Phae Lay mountain, around point 17°31'02.7"N, 105°47'17.2"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 300-530 m a.s.l. Treelet 5-6 m tall on rocky mountain top. Flowers yellowish-brown. Common. 11 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 914.

FLORA OF LAOS Hin Nam No Nature Protected Area Annonaceae Goniothalamus sp. 1 Laos, Khammouane prov., Boualapha distr., Nong Seng village, southern sandstone slopes of Phou Chuang (Poujeuang) Mt. around point 17°34'13.3"N 105°47'57.5"E. Primary evergreen broad-leaved dry forest at elev. 700-1000 m a.s.l. Caulifloric treelet 3-4 m tall. Flowers pale yellow-greenish to dull orange. Not common. 3 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 543.

FLORA OF LAOS Hin Nam No Nature Protected Area Ancistrocladaceae Ancistrocladus tectorius (Lour.) Merr. Laos, Khammoune prov., Boualapha distr., Nong Seng village, Chanh Daeng mountain, around point 17°31'11.1"N, 105°46'41.4"E. Secondary dry evergreen and semideciduous broad-leaved forest on sandy alluvial flat valley along river between karstic limestone hills at about 180 m a.s.l. Scandent shrub or woody vine 4-6 m long. Very common. Usual co-dominant of shrub forest stratum. 11 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 1044.

FLORA OF LAOS Hin Nam No Nature Protected Area Anacardiaceae Mangifera sp. Laos, Khammouane prov., Boualapha distr., Nong Seng village, southern sandstone slopes of Phou Chuang (Poujeuang) Mt. around point 17°34'13.3"N, 105°47'57.5"E. Primary evergreen broad-leaved dry forest at elev. 650-750 m a.s.l. Tree 15-20 m tall on rocky cliff. Fruits light green. Occasional. 6 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 719.

FLORA OF LAOS Hin Nam No Nature Protected Area Anacardiaceae Lannea coromandelica (Houtt.) Merr. ? Laos, Khammoune prov., Boualapha distr., Vang Maner village, Palai mountain area, around point 17°30'13.6"N, 105°49'23.9"E, 520 m a.s.l. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble- like highly eroded limestone at 300-500 m a.s.l. Treelet 3-4 m tall on exposed rocks near mountain top. Not rare. 8 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 805.

FLORA OF LAOS Hin Nam No Nature Protected Area Anacardiaceae Lannea coromandelica (Houtt.) Merr. ? Laos, Khammoune prov., Boualapha distr., Nong Seng village, Chanh Daeng mountain, around point 17°31'11.1"N, 105°46'41.4"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 400-600 m a.s.l. Tree 10-15 m tall on open rocks near mountain top. Very common. 11 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 962.

FLORA OF LAOS Hin Nam No Nature Protected Area Anacardiaceae Allospondias lakonensis Stapf ? Laos, Khammoune prov., Boualapha distr., Vang Maner village, Palai mountain area, around point 17°30'13.6"N, 105°49'23.9"E, 520 m a.s.l. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 300-500 m a.s.l. Tree 20-30 m tall. Flowers white. Common. Usual co-dominant of the forest at middle part of hill slopes. 8 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 753.

FLORA OF LAOS Hin Nam No Nature Protected Area Anacardiaceae Lannea sp. ? Laos, Khammoune prov., Boualapha distr., Nong Seng village, Chanh Daeng mountain, around point 17°31'11.1"N, 105°46'41.4"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 400-600 m a.s.l. Tree 6-10 m tall trees on rocky slope near mountain top. Fruits green 1-seeded. Stem bark green. Common. 11 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 1017.

FLORA OF LAOS Hin Nam No Nature Protected Area Anacardiaceae Pistacia weinmanniifolia J. Poiss. ex Franch. Laos, Khammoune prov., Boualapha distr., Vang Maner village, Palai mountain area, around point 17°30'13.6"N, 105°49'23.9"E, 520 m a.s.l. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 300-500 m a.s.l. Tree 6-8 m tall on exposed rocks of mountain top. Common. Occasional co-dominant of deciduous forest on mountain tops. 8 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 781.

FLORA OF LAOS Hin Nam No Nature Protected Area Anacardiaceae Mangifera sp. Laos, Khammoune prov., Boualapha distr., Nong Seng village, Chanh Daeng mountain, around point 17°31'11.1"N, 105°46'41.4"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 400-600 m a.s.l. Tree about 20 m tall on shady rocky slope. Fruits dull glaucous green. Not common. 11 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 1021.

FLORA OF LAOS Hin Nam No Nature Protected Area Anacardiaceae Semecarpus cochinchinensis Engl. Laos, Khammoune prov., Boualapha distr., Nong Seng village, Chanh Daeng mountain, around point 17°31'11.1"N, 105°46'41.4"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 400-600 m a.s.l. Tree 5-7 m tall trees on rocky slope near mountain top. Fruit appendage fleshy, dull yellow. Very common. 11 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 1016.

FLORA OF LAOS Hin Nam No Nature Protected Area Anacardiaceae Lannea sp. ? Laos, Khammoune prov., Boualapha distr., Nong Seng village, Chanh Daeng mountain, around point 17°31'11.1"N, 105°46'41.4"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 400-600 m a.s.l. Tree 6-10 m tall trees on rocky slope near mountain top. Fruits green 1-seeded. Stem bark green. Common. 11 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 1017.

FLORA OF LAOS Hin Nam No Nature Protected Area Anacardiaceae Lannea coromandelica (Houtt.) Merr. ? Laos, Khammoune prov., Boualapha distr., Nong Seng village, Chanh Daeng mountain, around point 17°31'11.1"N, 105°46'41.4"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 400-600 m a.s.l. Treelet 3-7 m tall on open rocky slope near mountain top. Common. 11 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 1031.

FLORA OF LAOS Hin Nam No Nature Protected Area Amaranthaceae Deeringia polysperma (Roxb.) Moq. Laos, Khammoune prov., Boualapha distr., Nong Seng village, Vang Phae Lay mountain, around point 17°31'02.7"N, 105°47'17.2"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 300-530 m a.s.l. Shrub 2-3 m tall in open place. Fruits white. Occasional. 11 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 941.

FLORA OF LAOS Hin Nam No Nature Protected Area Agavaceae Dracaena cochinchinensis (Lour.) S.C. Chen Laos, Khammoune prov., Boualapha distr., Nong Seng village, Chanh Daeng mountain, around point 17°31'11.1"N, 105°46'41.4"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 400-600 m a.s.l. Palm-like treelet 3-4 m tall on exposed rocks near mountain top. Very common. 11 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 986.

FLORA OF LAOS Hin Nam No Nature Protected Area Agavaceae Dracaena angustifolia (Medik.) Roxb. ? Laos, Khammoune prov., Boualapha distr., Vang Maner village, Palai mountain area, around point 17°30'13.6"N, 105°49'23.9"E, 520 m a.s.l. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 300-500 m a.s.l. Shrub to 3 m tall. Fruits glossy olive green. Common. 8 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 780.

FLORA OF LAOS Hin Nam No Nature Protected Area Agavaceae Dracaena elliptica Thunb. Laos, Khammouane prov., Boualapha distr., Nong Seng village, southern sandstone slopes of Phou Chuang (Poujeuang) Mt. around point 17°34'13.3"N 105°47'57.5"E. Primary evergreen broad-leaved dry forest at elev. 700-1000 m a.s.l. Shrub 2-3 m tall, fruits red. Not rare. 3 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 575.

FLORA OF LAOS Hin Nam No Nature Protected Area Aceraceae Acer laurinum Hassk. Laos, Khammouane prov., Boualapha distr., Nong Seng village, southern sandstone slopes of Phou Chuang (Poujeuang) Mt. around point 17°34'13.3"N 105°47'57.5"E. Primary evergreen broad-leaved dry forest at elev. 700-1000 m a.s.l. Tree to 15 m tall. Fruits and fruit wing light greenish to yellowish. Occasional. 3 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 583.

FLORA OF LAOS Hin Nam No Nature Protected Area Acanthaceae Peristrophe bivalvis (L.) Merr. Laos, Khammouane prov., Boualapha distr., Nong Seng village, southern sandstone slopes at summit of Phou Chuang (Poujeuang) Mt. around point 17°35'18.1"N, 105°47'47.5"E. Primary evergreen mixed and coniferous dense wind-formed forest at elev. 1000-1400 m a.s.l. Terrestrial herb to small undershrub on open mossy rocks. Flowers purple-violet. Not common. 5 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 661a.

FLORA OF LAOS Hin Nam No Nature Protected Area Acanthaceae Xylacanthus laotica Aver. et K. S. Nguyen Laos, Khammoune prov., Boualapha distr., Nong Seng village, Chanh Daeng mountain, around point 17°31'11.1"N, 105°46'41.4"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 400-600 m a.s.l. Shrub or treelet 2-3 m tall on open rocky slope near mountain top. Very common. 11 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 1023.

FLORA OF LAOS Hin Nam No Nature Protected Area Acanthaceae Lepidagathis incurva D. Don Laos, Khammouane prov., Boualapha distr., Nong Seng village, southern sandstone slopes of Phou Chuang (Poujeuang) Mt. around point 17°34'13.3"N 105°47'57.5"E. Primary evergreen broad-leaved dry forest at elev. 700-1000 m a.s.l. Terrestrial and lithophytic herb on shady shelves of vertical cliff. Not common. 3 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 590.

FLORA OF VIETNAM

FLORA OF VIETNAM

FLORA OF VIETNAM

FLORA OF VIETNAM

FLORA OF VIETNAM

FLORA OF VIETNAM

FLORA OF VIETNAM Parnassiaceae Parnassia wightiana Wall. Ha Giang prov., Quan Ba distr., Tung Vai commune, Thang village, around point 23º03ʹ00.4ʺN 104º52ʹ02.2ʺE, steep rocky slopes of river valley composed with eroded limestone at elevation 1000–1060 m a.s.l. Remnants of primary evergreen broad-leaved very humid forest. Terrestrial and lithophytic herb in wet grassy places along river and on mossy open wet stream rocks. Flowers white, fruits green, later brown. Locally very common. 19 October 2018, L. Averyanov, Nguyen Sinh Khang, T. Maisak, Truong Quang Ngan, VR 1045.

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM Convallariaceae Ophiopogon caulescens (Blume) Backer Ha Giang prov., Quan Ba distr., Bat Dai Son Nature Reserve, Can Ty commune, Dau Cau 1 village, around point 23º05ʹ26.3ʺN 105º01ʹ03.0ʺE, 1100-1150 m a.s.l., karstic, highly eroded mountains composed of solid marble-like limestone, primary evergreen broad-leaved humid forest on steep rocky slopes near mountain tops. Terrestrial herb among rocks in shady place. Flowers white with violet tint. Not rare. 14 April 2018, L. Averyanov, Nguyen Sinh Khang, Chuong Quang Ngan, T. Maisak, VR 136.

All photos © L. Averyanov & K.S. Nguyen

All photos © L. Averyanov & K.S. Nguyen