Herbarium LE - Specimens

Peliosanthes triandra Aver. et N.Tanaka sp.nov. Type Southern Cambodia, to the N of Sianukwille town, Kaoh Rong Samoloem Khong Island in Siam Gulf. Low hills around point 10°37’246’N 103°18’598’’. Dry evergreen lowland forest at elev. about 78 m a.s.l. 27 November 2012. Coll. T.Maisak, M.Telepova, L.Osinovets, I.Kutuzova. No 996. Flowered under cultivation in April 2012. Flowers odorless, uniform light green to yellowish-green. 8 April 2013. Maisak et al., 996. Photos.

FLORA OF VIETNAM

FLORA OF LAOS Convallariaceae Peliosanthes hirsuta Aver. et N.Tanaka, sp.nov. Vientiane Prov., Kasi Distr., Namken village, Phachao Mt., around point 19°18’45.5”N 102°22’31.4”E. Primary broad-leaved evergreen forest on very steep rocky slopes of remnant mountain composed with highly eroded solid crystalline limestone at elevations 1500-1750 m. Terrestrial herb on rocky shady mountain slope at elev. 700-1000 m. Leaves uniform dark glossy green. Common. LA-VN 840 24 March 2013 Coll.: L.Averyanov, N.S.Khang, S.Lorphengsy Fl. in cult. 21 Oct. 2016 & 1 Nov. Photos.

FLORA OF VIETNAM

FLORA OF VIETNAM Peliosanthes kenhillii Aver., N.Tanaka et K.S.Nguyen Epitype Ninh Binh prov., Nho Quan district, Cuc Phuong national park, around point 20°15'03.6"N 105°41'58.7"E (N20 15 03.6 E105 41 58.7), at elev. about 165 m a.s.l., scattered in closed evergreen broad-leaved lowland secondary forests at the base of limestone hill. Perennial herb with rhizome, inflorescence about 15-20 cm, flower pale white-green. Common 9 December 2014, Nguyen Sinh Khang NSK 760. d-EXSICCATES OF VIETNAMESE FLORA 0222/NSK 760 (LE) © L.Averyanov & K.S.Nguyen

FLORA OF VIETNAM Peliosanthes hexagona Aver., N.Tanaka et K.S.Nguyen Epitype Hoa Binh prov., Lac Thuy district, Phu Thanh commune, Tan Lam village, around point 20°34'55.48"N 105°40'56.20"E, at elev. about 120 m a.s.l., scattered under secondary closed evergreen broad-leaved lowland forests on limestone. Prennial herb with rhizome, inflorescence about 20 cm, flower pale green, not common. 7 May 2014, Nguyen Sinh Khang, NSK 674 d-EXSICCATES OF VIETNAMESE FLORA 0218/NSK 674 (LE) © L.Averyanov & N.S.Khang

FLORA OF VIETNAM

FLORA OF LAOS Convallariaceae Peliosanthes densiflora Aver. & N. Tanaka Vientiane Prov., Vang Vieng Distr., Nathong village, about 5 km to the W of Vang Vieng town, Tham Kang Mt., around point 18°55’51.4”N 102°23’50.6”E. Dry broad-leaved primary and secondary evergreen forest on very steep rocky slopes and on vertical cliffs of remnant highly eroded rocky limestone mountains composed by solid crystalline limestone at elevation 300-400 m. Clustering terrestrial and lithophytic herb with erect to suberect many branching rhizome on steep rocky slope. Leaves to 80 cm long, petiole 30-45 cm long, leaf blade glossy uniform green on both sides 25-35 cm long, 6-10 cm wide. Not rare. LA-VN 421 14 March 2013 Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, P.V.The, S.Lorphengsy

FLORA OF LAOS Convallariaceae Peliosanthes choriandra Aver. et al. Type Luang Prabang prov., Pon Xay distr., Houay Man village, Phou Pak Sang Mountain, around point 19°59'03.8''N 102°25'09.0''E. Degraded primary evergreen and semideciduous forest on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. 1000-1150 m a.s.l. Terrestrial herb on steep shady rocky mountain slope. Flowers green. Occasional. LA-VN 2059 4 April 2017 Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.Maisak, Vichith Lamxay, Koudkeo Phommachanh, Phizar Sayalath, Keooudone Souvannakhoummane. d-EXSICCATES OF VIETNAMESE FLORA 0282/LA-VN 2059 © L.Averyanov

FLORA OF VIETNAM Ophiopogon tristylatus Aver., N. Tanaka et Luu Epitype Vietnam, Lam Dong Prov., Lac Duong Distr., Da Chais Municipality, near Giang Ly Forest Station around point 12°10’52’’N, 108°42’44’’E. Fertile and humid soil, near stream, under evergreen broad-leaved forest at elevation about 1700 m a.s.l. Luu 902 27 June 2012 Coll.: Luu Hong Truong, Pham Huu Nhan d-EXSICCATES OF VIETNAMESE FLORA 0197/Luu 902 (LE) © Luu Hong Truong, L.Averyanov

FLORA OF VIETNAM NEW Plants of Phong Nha - Ke Bang National Park Fam. Liliaceae s.l. Convallariaceae Peliosanthes argenteostriata Aver., sp.nov. Type Vietnam, Quang Binh Prov., Minh Hoa Distr., Thuong Hoa Municipality, about 1-1.5 km to SW of Ban On village (17º40'21''N, 105º58'00''E). Ca Xach mountain (17º39'20''N, 105º57'42''E). Primary closed evergreen broad-leaved gnarled stunted forest along rocky ridge composed with solid marble-like highly eroded crystalline limestone at elevation 600-800 m a.s.l. Terrestrial and lithophytic herb on very steep rocky slope. Leave deep green with broad median whitish stripe. Not rare. No HAL 5975 20 January 2005. Collectors: L.Averyanov, P.K.Loc, P.V.The, A.Averyanova, N.T.Vinh, N.Q.Vinh, N.T.Binh.

FLORA OF VIETNAM Convallariaceae Ophiopogon alatus Aver. et N.Tanaka Cao Bang Prov., Bao Lac Distr., Hong An municipality, Mi Lung village. Primary broad-leaved and mixed humid evergreen forest on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation 1500-1550 m a.s.l. around point 22°50’01.3’’N 105°50’05.7’’E. Terrestrial herb among mossy rocks on rocky steep slope near mountain top. Fruits glossy dark blue. Very common. 20 November 2014 CPC 7535a / TM 1193. Seeds collected only. Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz Plant grown from seeds. CPC 7535a / TM 1193 / 14334. Flowers white with purple-violet tint, darker inside. Photos. 20 Nov. 2016. L. Averyanov

Asclepiadaceae Hoya nutans Aver. et V.T. Pham Type Hoa Binh Prov., Lac Son Distr., Tu Do Municipality, Mon village, around point 20°25’29”N 105°19’36”E. Primary broad-leaved forest on steep rocky slopes of remnant mountain composed with solid crystalline highly eroded limestone at elevation about 500-700 m a.s.l. Semi-woody vine. Occasional. CPC 1592e / 11531 25 March 2011 Coll.: N.Q.Hieu, L.Averyanov, N.T.Hiep, P.K.Loc, P.V.The, N.T.Vinh, T.B.Ngan, N.Tap, D.D.Dao, V.T.Ha, T. Maisak, L. Osinovetz. Fl. et coll. under cultivation at 16 July 2016. Flowers with light sweet fragrance, white, corona pink. L.Averyanov. Photos. d-EXSICCATES OF VIETNAMESE FLORA 0258/CPC 1592e © L.Averyanov

FLORA OF VIETNAM Asclepiadaceae Hoya crassipetiolata Aver. Type Dien Bien Prov., Muong Cha Distr., Hua Ngai Municipality, Ha La Chu Village, around point 21°53’46”N 103°10’17”E. Highly degraded primary evergreen broadleaved humid forest on very steep slopes of remnant mountains composed with solid limestone at elevation 1100-1400 m a.s.l. Epiphyte on tall mossy tree. Common. CPC 2049a 07 April 2011 Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, N.T.Vinh. Flowered in cult. in 6 May 2015. Flowers white with pink tint. Photos. d-EXSICCATES OF VIETNAMESE FLORA 0257/CPC 2049a © L.Averyanov

FLORA OF VIETNAM

FLORA OF VIETNAM Fam. Convallariaceae Tupistra theana Aver. et N.Tanaka, sp.nov. Epitype Quang Binh Prov., Minh Hoa Distr., Hoa Tien municipality, La Van village, around point 17°51’45”N 105°49’45”E. Highly degraded primary and secondary evergreen broad-leaved forest on very steep rocky slopes of remnant limestone hills composed with highly eroded marble-like crystalline limestone at elev. 200-450 m a.s.l. Lithophytic herb on shelves and in crevices of tall shady vertical cliff near to top of mountain. Not common. CPC 2581 30 April 2011. Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, P.V.The, N.T.Vinh. d-EXSICCATES OF VIETNAMESE FLORA 0182/HAL 2581 (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VIETNAM Fam. Convallariaceae Tupistra theana Aver. et N.Tanaka, sp.nov. Epitype Quang Binh Prov., Minh Hoa Distr., Hoa Tien municipality, La Van village, around point 17°51’45”N 105°49’45”E. Highly degraded primary and secondary evergreen broad-leaved forest on very steep rocky slopes of remnant limestone hills composed with highly eroded marble-like crystalline limestone at elev. 200-450 m a.s.l. Lithophytic herb on shelves and in crevices of tall shady vertical cliff near to top of mountain. Not common. CPC 2581 30 April 2011. Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, P.V.The, N.T.Vinh. d-EXSICCATES OF VIETNAMESE FLORA 0182/HAL 2581 (CPC, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF VIETNAM Tupistra gracilis Aver. & N.Tanaka Epitype Thanh Hoa Prov., Thuong Xuan Distr., Bat Mot Municipality, Duc village, Xuan Lien Natural Reserve. Primary broad-leaved evergreen forest on rocky, highly eroded, crystalline limestone at elevation 800-1000 m a.s.l. around point 20°01’11.6’’N 104°57’47.5’’E. Terrestrial and lithophytic herb with leaves to 1.5 m long and thick creeping ascending rhizome in very steep shady rocky slope. Rare. 4 November 2013 CPC 6721 Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, N.D.Thang, L.V.Tien Fl. in 22 Nov. 2016. d-EXSICCATES OF VIETNAMESE FLORA 0269/CPC 6721 © L.Averyanov

FLORA OF LAOS Convallariaceae Tupistra theana Aver. et N.Tanaka Laos, Khammouane prov., Thakhek. На известняковых отломах останца в подлеске. Растение с длинными жесткими листьями. 161 м над ур. м., 17°27.195N, 104°57.790E. Е.Константинов K248. Гербарный сбор/фото 03.11.2013.

FLORA OF LAOS Convallariaceae Tupistra orlovii Aver., N. Tanaka & K. S. Nguyen, sp.nov. Bolikhamsai prov., Thaphabat distr., Nam Bon village, Phou Khao Khouay national park, Tad Xai Waterfall, around point 18°27'25.9''N 103°08'17.5''E. Degraded primary and secondary evergreen dry forest along river on eroded sandstone at elev. About 340 m a.s.l. Lithophytic and occasionally terrestrial large herb on shady mossy boulders and river cliffs. Leaves uniform green, fruits glossy green. Rare, but locally abundant. LA-VN 2191 11 April 2017 Photos Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.Maisak. d-EXSICCATES OF VIETNAMESE FLORA 0283/Orlov s.n. & LA-VN 2191 © L.Averyanov

FLORA OF VIETNAM Rohdea filosa Aver. et N. Tanaka Tuyen Quang prov., Na Hang distr., Sinh Long municipality, Khuoi Phin village, around point 22°38'21.9"N, 105°20'43.5"E. Primary evergreen mixed humid forest on rocky limestone on highly eroded karstic hills composed with gray solid marble-like limestone at elevation 1100-1200 m a.s.l. Terrestrial and lithophytic rosulate herb with ascending rhizome to 0.5 m long. Fruits globular, dark green, 1-seeded. Locally very common. 29 September 2013 CPC 5299 L.Averyanov et al. Flowered 25 April 2017. Flowers green, sweet fragrant. d-EXSICCATES OF VIETNAMESE FLORA 0279/CPC 5299 © L.Averyanov

FLORA OF LAOS Convallariaceae Rohdea dracaenoides Aver. et N.Tanaka, sp.nov. Type. Vientiane Prov., Kasi Distr., Namken village, Phachao Mt., around point 19°18’45.5”N 102°22’31.4”E. Primary broad-leaved evergreen forest on very steep rocky slopes of remnant mountain composed with highly eroded solid crystalline limestone at elevations 1500-1750 m. Lithophyte on steep rocky slopes and shady cliffs. Flowers light green to light yellow-orange. Very common from elev. 1600 to mountain top. LA-VN 785 24 March 2013 Coll.: L.Averyanov, N.S.Khang, S.Lorphengsy

FLORA OF VIETNAM Liliaceae s.l. (Convallariaceae) Peliosanthes retroflexa Aver. et N.Tanaka Type Quang Binh Prov., Minh Hoa Distr., Dan Hoa Municipality, Giang Man Ridge (Vietnam-Laos border) near Cha Lo Vietnam-Lao border gate around point 17°41’50”N 105°45’54”E. Primary evergreen broad-leaved and coniferous (with Dacrycarpus and Fokienia) wet forest along ridge edge composed with shale and sandstone at elevation about 900-1100 m a.s.l. Terrestrial herb on shady slope. Fruits blue. Common. HAL 12374 8 February 2009 Coll.: L.Averyanov, P.K.Loc, N.T.Vinh, L.T.Son.

FLORA OF VIETNAM

FLORA OF VIETNAM

FLORA OF LAOS Convallariaceae Peliosanthes tatianae Aver et al., sp.nov. Type Vientiane prov., Vang Vieng distr., Patang village, around point 19°04'15.6''N 102°24'33.2''E. Secondary broad-leaved semi-deciduous dry forest on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. about 670 m a.s.l. Terrestrial herb in shady steep slope. Flower buds white. Not rare. LA-VN 1607 23 March 2017 Photos Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.Maisak, Khamla Inkhavilay, Hongthong Southyvohanh, Pinmany Sisavath, Chanthala Cheuaysamphanh.

FLORA OF VIETNAM Peliosanthes kenhillii Aver., N.Tanaka et K.S.Nguyen Epitype Ninh Binh prov., Nho Quan district, Cuc Phuong national park, around point 20°15'03.6"N 105°41'58.7"E (N20 15 03.6 E105 41 58.7), at elev. about 165 m a.s.l., scattered in closed evergreen broad-leaved lowland secondary forests at the base of limestone hill. Perennial herb with rhizome, inflorescence about 15-20 cm, flower pale white-green. Common 9 December 2014, Nguyen Sinh Khang NSK 760. d-EXSICCATES OF VIETNAMESE FLORA 0222/NSK 760 (LE) © L.Averyanov & K.S.Nguyen

FLORA OF LAOS Convallariaceae Peliosanthes argenteostriata Laos, Vientiane prov., окрестности Kasi, elev. 700 m a.s.l. старая дорога на Луангпробанг 40 км от Каси на север, облесенный берег реки Nam Kean, подножие западного склона горы Ph. Phaday 30.11.2014 E.Konstantinov K 305. Сбор. Фото

FLORA OF LAOS Convallariaceae Peliosanthes densiflora Aver. et N.Tanaka sp.nov. Laos, Vientiane prov., Kasi. Окрестности пещеры Khuang Lang Cave берег лесной реки Древостой включает: Tetrameles nudiflora, Pometia eximia, Croton argyratus, Arenga westerhoutii, Ficus religlosa. По табличкам нац. парка. 631 м над ур. м.,N 19°18.198N, 102°09.031E. Е.Константинов K103. сбор живым корневищем, гербарий/фото 21.10.2013.

FLORA OF VIETNAM Liliaceae s.l. (Convallariaceae) Peliosanthes sp.nov. Lai Chau Prov., Phong Tho Distr., Sin Suoi Ho Municipality, Sa Ma Pho village, around point 22°33’21’’N 103°34’29’’E. Secondary evergreen broad-leaved wet closed forest with bamboo on very steep mountain slopes and along ridges composed predominantly with weathered granite at elev. about 1900-2050 m a.s.l. Terrestrial herb on shady rocky slope. Flowers light dull yellowish. Fruits blue, juicy. Locally very common. HAL 10767 14 Dec. 2006 Photo. Coll.: Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc, Pham Van The, N.T.Vinh. d-EXSICCATES OF VIETNAMESE FLORA 0089/HAL10767 (HN, LE) © Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc

FLORA OF LAOS Convallariaceae Peliosanthes choriandra Aver. et al. Type Luang Prabang prov., Pon Xay distr., Houay Man village, Phou Pak Sang Mountain, around point 19°59'03.8''N 102°25'09.0''E. Degraded primary evergreen and semideciduous forest on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. 1000-1150 m a.s.l. Terrestrial herb on steep shady rocky mountain slope. Flowers green. Occasional. LA-VN 2059 4 April 2017 Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.Maisak, Vichith Lamxay, Koudkeo Phommachanh, Phizar Sayalath, Keooudone Souvannakhoummane. d-EXSICCATES OF VIETNAMESE FLORA 0282/LA-VN 2059 © L.Averyanov

FLORA OF VIETNAM Ophiopogon tristylatus Aver., N. Tanaka et Luu Epitype Vietnam, Lam Dong Prov., Lac Duong Distr., Da Chais Municipality, near Giang Ly Forest Station around point 12°10’52’’N, 108°42’44’’E. Fertile and humid soil, near stream, under evergreen broad-leaved forest at elevation about 1700 m a.s.l. Luu 902 27 June 2012 Coll.: Luu Hong Truong, Pham Huu Nhan d-EXSICCATES OF VIETNAMESE FLORA 0197/Luu 902 (LE) © Luu Hong Truong, L.Averyanov

FLORA OF VIETNAM Convallariaceae Ophiopogon fruticulosa Aver., N.Tanaka et S.K.Nguyen, sp.nov. Type Tuyen Quang Prov., Na Hang Distr., Sinh Long Municipality, Khuoi Phin village, around point 22°38'21"N, 105°20'28"E. Remnants of primary forest along stream, on wet very steep shady humid travertine river valley. Erect shrub to 1.2 m tall in shady wet place. Flowers white with light violet tint. Locally abundant. 30 September 2013 CPC 5345 L.Averyanov, N.T.Hiep, L.M.Tuan, N.S.Khang, D.V.Cuong, N.V.Truong.T.Maisak, L.Osinovets

FLORA OF LAOS NEW Asclepiadaceae Hoya uniflora Aver. Type Xiangkhouang Prov., Peak Distr., Khoang village, Pa Khao mt., around point 19°33’02.2’N 103°39’56.3’E. Primary and secondary broad-leaved evergreen dry forest on solid rocky highly eroded limestone at elev. 1200-1450 m a.s.l. Creeping lithophytic vine (10)15 25(35) cm long on open mossy rocks on cliff. Flower bud white. Locally abundant. LA-VN 961 3 April 2015 Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

FLORA OF HON BA NATURE RESERVE Peliosanthes aperta Aver. et N.Tanaka sp.nov. S. Vietnam, Khanh Hoa province, Hon Ba nature rerseve around point 12°06’531”N 108°59’409”E, elev. 407 m a.s.l. Terrestrial herb in evergreen forests. May 2014 Truong Ba Vuong, no 1.

FLORA OF VIETNAM Convallariaceae Ophiopogon petraeus Aver. et N.Tanaka Type Lang Son Prov., Huu Lung Distr., Huu Lien Commune, Huu Lien Nature Reserve, around point 21°40’28”N, 106°22’34”E. Primary closed evergreen seasonal tropical mixed forests on rocky limestone at elevation 400-600 m. Lithophytic rosulate herb. Not rare. HAL 11220b 24 September 2007 Coll.: P.K.Loc, N.T.Vinh, H.V.Luat Flowered under cultivation at September-October 2013. Flowers white, anthers light yellow-green, pedicel white with violet tint. Leaves fleshy, at the base subsucculent, juicy, transversely-oval in section. Photos. L. Averyanov, HAL 11220b/1. d-EXSICCATES OF VIETNAMESE FLORA 0207/HAL 11220b (LE) © L.Averyanov

FLORA OF VIETNAM

Asclepiadaceae Hoya nutans Aver. et V.T. Pham Type Hoa Binh Prov., Lac Son Distr., Tu Do Municipality, Mon village, around point 20°25’29”N 105°19’36”E. Primary broad-leaved forest on steep rocky slopes of remnant mountain composed with solid crystalline highly eroded limestone at elevation about 500-700 m a.s.l. Semi-woody vine. Occasional. CPC 1592e / 11531 25 March 2011 Coll.: N.Q.Hieu, L.Averyanov, N.T.Hiep, P.K.Loc, P.V.The, N.T.Vinh, T.B.Ngan, N.Tap, D.D.Dao, V.T.Ha, T. Maisak, L. Osinovetz. Fl. et coll. under cultivation at 16 July 2016. Flowers with light sweet fragrance, white, corona pink. L.Averyanov. Photos. d-EXSICCATES OF VIETNAMESE FLORA 0258/CPC 1592e © L.Averyanov

FLORA OF VIETNAM Asclepiadaceae Hoya sp.? Thua Thien – Hue Prov., Nam Dong Distr., Thuong Lo Municipality, Cha Mang river basin, around point 16°07’53’’N 107°43’27’’E. Primary evergreen broad-leaved lowland forest on very steep hill slopes composed with stratified shale at elev. 400-500 m a.s.l. Semi-woody vine 3-5 m long on very steep rocky shady river slope. Flowers yellow. Rare. HAL 11089 11 Apr. 2007. Photo. Coll.: L.Averyanov, N.S.Khang, A.Averyanova, N.D.Phuong, L.V.Hung.

FLORA OF VIETNAM Fam. Asclepiadaceae Hoya crassipetiolata Aver. et V.T. Pham Paratype N.Vietnam, Bac Kan Prov., Cho Don Distr., Ban Thi Municipality, village Phia Khao, Lung Li valley, around point 22º16'10''N, 105º31'01''E, Cot Moc mountain at elevation 800-900 m a.s.l. Primary broad-leaved evergreen closed forest on very steep slopes and cliffs near tops of rocky ridge composed with solid marble-like stratified limestone. Herbaceous or hemiwoody vine 4-6 m long. Flowers white, disc light pink. Not common. No HAL 4836 23 May 2004 Collectors: L.Averyanov, N.T.Hiep, P.V.The, N.T.Vinh.