Herbarium LE - Specimens

FLORA OF VIETNAM
Orchidaceae
Liparis distans C.B. Clarke

N. Vietnam, prov. Cao Bang, distr. Bao Lac, municipality Yen Lac,
vicinities of Yen Lac village (N 22°44′ E 105°50′), about 44 km to NWW
from Cao Bang town. Primary open coniferous mossy forest along tops of
limestone remnant karst ridge at 1500 m alt. Epilite on steep slopes
and cliffs. Flowers: lip green, tepals yellowish-green. Very common.

CBL 324 15 Nov. 1998

International Botanical Expedition of the U.S.A. National Geographic
Society (grant No 6300-98)

Collectors: Averyanov L., N.T.Hiep, P.K.Loc, N.X.Tam.

Orchidaceae

Liparis distans C.B. Clarke

Vietnam. Prov. Ha Nam Ninh, Cuc Phuong, No CP 1168, 22 11 1971, Coll. Lac Thinh.

Det. Averyanov L. 1987

FLORA OF VIETNAM
Orchidaceae
Liparis distans C.B.Clarke

Son La Prov., Van Ho Distr., Xuan Nha Municipality, Muong An village, territory of Xuan Nha natural reserve.

Primary broad-leaved evergreen humid forest on very steep slopes and cliffs of remnant mountain composed by solid highly eroded crystalline limestone at elevation 800-1000 m a.s.l. around point 20°45′06.5″N 104°46′37.8″E.

Lithophytic clustering herb on very steep rocky slope. Flowers olive-orange. Abundant.

13 November 2013 CPC 7057
Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, N.D.Thang, L.D.Qui

FLORA OF VIETNAM
Orchidaceae
Liparis distans C.B. Clarke

N. Vietnam, prov. Cao Bang, distr. Thach An, about 25 km to SSE from
Cao Bang town (N 22°27′ E 106°22). Primary evergreen closed
semideciduous forest on tops of remnant limestone ridge at 500 m alt.
Terrestrial and epilite herb on open rocks and cliffs. Flowers
olive-green-brown. Common.

CBL 919 05 Dec. 1998

International Botanical Expedition of the U.S.A. National Geographic
Society (grant No 6300-98)

Collectors: Averyanov L., N.Q.Binh, N.X.Tam.

FLORA OF VIETNAM

Fam. Orchidaceae

Liparis distans C.B. Clarke

N. Vietnam, prov. Cao Bang, distr. Thach An, municipality Van Trinh, (N 22ш21' E 106ш27'), about 39 km to SSE from Cao Bang town. Secondary open semideciduous broadleaved forest on steep slopes near tops of remnant limestone ridge at 600 m alt. Epilite herb on steep slopes. Flowers olive-green. Not rare. No CBL 849.4 03 Dec. 1998

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 6300-98)

Collectors: Averyanov L., N.Q.Binh, P.K.Loc, N.X.Tam.

FLORA OF VIETNAM

Orchidaceae Juss.

Liparis distans C.B.Clarke ?

N. Vietnam, Prov. Cao Bang, Distr. Tra Linh, Subdistr. Quoc Toan, open dry evergreen eteleeria davidianaﰠforest on tops of remnant limestone mountains at 800-900 m alt. near Thang Heng lake in environs of Thang Hen and Lung Tao villages.

Epilite. Flowers greenish. Not rare at 700-800 m alt.

VH 2480 04.01.1996

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 5094-93)

Participants: L. Averyanov, N.T. Ban, A. Budantzev, L. Budantzev, N.T. Hiep, D.D. Huyen, P.K. Loc, G. Yakovlev

FLORA OF VIETNAM

Orchidaceae

Liparis distans C.B.Clarke

Thanh Hoa Prov., Thuong Xuan Distr., Bat Mot Municipality, Vin village, Xuan Lien Natural Reserve.

Remnants of primary broad-leaved evergreen humid forest on rocky, highly eroded, crystalline limestone at elevation 750-800 m a.s.l. around point 19°59’11.4’’N 104°58’31.8’’E.

Lithophytic herb on vertical shady cliffs. Flowers olive-yellowish. Abundant

1 November 2013 CPC 6421

L.Averyanov, N.T.Hiep, N.S.Khang

FLORA OF VIETNAM

Orchidaceae

Liparis distans C.B.Clarke

Thanh Hoa Prov., Thuong Xuan Distr., Bat Mot Municipality, Vin village, Xuan Lien Natural Reserve.

Primary broad-leaved evergreen humid forest on rocky limestone at elevation about 850 m a.s.l. around point 19°58’51.6’’N 104°59’38.08’’E.

Clustering lithophytic herb. Flowers yellow-brown. Very common.

5 November 2013 CPC 6798

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, N.D.Thang, L.V.Tien

FLORA OF VIETNAM
Orchidaceae
Liparis distans C.B.Clarke

Son La Prov., Van Ho Distr., Van Ho comm., Hua Tat village, around point 20°46′21.7N 104°47′47.5E.

Remnants of primary evergreen broad-leaved and mixed forest (with Pinus wangii) on very steep rocky slopes and on mountain tops composed with highly eroded karstic light gray marble-like limestone at elevation 1300-1350 m a.s.l.

Lithophyte on exposed mossy rocks. Flowers olive-brownish. Very common.

2 October 2016 CPC 8194
Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.V.Maisak, N.T.Son.

FLORA OF VIETNAM

Orchidaceae

Liparis distans C.B.Clarke

Thanh Hoa Prov., Thuong Xuan Distr., Bat Mot Municipality, Vin village, Xuan Lien Natural Reserve.

Primary broad-leaved evergreen humid forest on rocky limestone at elevation about 850 m a.s.l. around point 19°58’51.6’’N 104°59’38.08’’E.

Clustering lithophytic herb. Flowers yellow-brown. Very common.

5 November 2013 CPC 6798

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, N.D.Thang, L.V.Tien

FLORA OF VIETNAM
Orchidaceae
Liparis distans C.B.Clarke

Thanh Hoa Prov., Thuong Xuan Distr., Bat Mot Municipality, Duc village, Xuan Lien Natural Reserve.

Primary broad-leaved evergreen forest on rocky limestone at elevation 800-1000 m a.s.l. around point 20°01′11.6″N 104°57′47.5″E.

Clustering lithophytic herb on shady steep slope. Flowers dull yellow to yellow-olive-brownish. Abundant.

3 November 2013 CPC 6635
Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, N.D.Thang, L.V.Tien

FLORA OF LAOS
Orchidaceae
Liparis distans C.B.Clarke?

Houphan Prov., Viengxay Distr., Vieng Xai village, around point 20°23′41.3′'N 104°13′44.5′'E.

Primary and secondary broad-leaved evergreen dry forest on very steep rocky slopes of highly eroded mountains composed by solid marble-like limestone at elev. 800-950 m a.s.l.

Lithophyte on shady rocks near mountain top. Common.

LA-VN 1132 7 April 2015
Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Somneuk Bouthamounty.

FLORA OF VIETNAM
Orchidaceae
Liparis balansae Gagnep.

Cao Bang Prov., Bao Lac Distr., Dinh Phung municipality, Ban O village.

Primary mixed and coniferous evergreen humid forest with (Pinus sp., Pseudotsuga sinensis, Podocarpus neriifolius) on very steep slopes and on mountain tops composed by solid highly eroded limestone at elevation 1200-1300 m a.s.l. around point 22°45′24.8″N 105°46′47.9″E.

Clustering lithophyte on mountain tops. Flowers yellowish. Very common.

24 November 2014 CPC 7614
Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz

FLORA OF VIETNAM
Orchidaceae
Liparis distans C.B.Clarke

Cao Bang Prov., Nguyen Binh Distr., Ca Thanh Municipality, Cao Lu village, around point 22°42′47.7″N, 105°51′50.8″E.

Primary coniferous forest with Pseudotsuga sinensis along highly eroded rocky limestone ridge at elevation about 1300 m a.s.l.

Clustering lithophyte on shady mossy rocks. Flowers dull orange. Occasional.

4 October 2013 CPC 5401
L.Averyanov, N.T.Hiep, L.M.Tuan, N.S.Khang, T.Maisak, L.Osinovets

Begoniaceae

Ha Giang Prov., Quan Ba Distr., Can Ty Commune, Sin Suoi Ho Village, around point 23.09566ºN 105.02142ºE. Very steep mountain slopes composed of solid highly eroded karstic limestone at elevation 800–1000 m a.s.l. Remnants of primary evergreen broad-leaved and coniferous (with Calocedrus rupestris and Xanthocyparis vietnamensis) humid forest. Lithophytic and terrestrial ephemeroid herb in. Not common. 11 October 2019, L. Averyanov, Nguyen Sinh Khang, T. Maisak, VR 1575a.

Herbarium and photos prepared in 28 August 2020, L. Averyanov, T. Maisak, VR 1575.1. Rhizome very short fleshy white, petioles light pale pink, leaves above dark green with many irregular silvery marks, below pale greenish to almost white irregularly marked with purple, tepals white or white with very light purple tint, anther and stigma bright yellow, fruits grassy green with 3 triangular wings, median twice larger than two others.

Herbarium and photos prepared in cult. 08 09 2020, L. Averyanov, T. Maisak, HAL 3506.2

Hin Nam No Nature Protected Area

Orchidaceae

Paphiopedilum gratrixianum Rolfe

Laos, Khammouane prov., Boualapha distr., Nong Seng village, southern sandstone slopes at summit of Phou Chuang (Poujeuang) Mt. around point 17°35'00.3"N, 105°47'35.6"E. Primary evergreen mixed and coniferous dense wind-formed forest at elev. 1000-1400 m a.s.l. Lithophytic rosulate herb on vertical mossy cliffs. Leaves uniform green. Locally common, particularly at elevations 1200-1400 m. 4 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 677.

Flowered in 17 Sept 2020. Median sepal white, heavily purple spotted in apical part, dark purple in basal half, lateral sepals light brown with darker nerves, lip yellow-brownish, staminode yellow and synsepalum pale yellow. Photos L. Averyanov, AL 677.2

Hin Nam No Nature Protected Area

Orchidaceae

Uncifera dalatensis (Guillaumin) Seidenf. & Smitinand

Laos, Khammouane prov., Boualapha distr., Nong Seng village, southern sandstone slopes at summit of Phou Chuang (Poujeuang) Mt. around point 17°35'00.3"N, 105°47'35.6"E. Primary evergreen mixed and coniferous dense wind-formed forest at elev. 1000-1400 m a.s.l. Epiphyte on tall trees on mountain top. Common.

4 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 675.2

Herbarium specimen and photos prepared in 08 09 2020, L. Averyanov, T. Maisak, AL 675.2. Flowers white, apical half of all tepals purple.

Orchidaceae

Kon Tum Province, Ngoc Linh Mt., primary broad-leaved evergreen forest at elev. 1800-2000 m a.s.l. Epiphyte on tall mossy tree. Not common. September 2019, Nguyen Van Canh, AL 1107.

Herbarium and photos are prepared in 08 09 2020, L. Averyanov, T. Maisak, AL 1107.1

Orchideaceae

Liparis dendrochiloides Seidenf. ex Aver.⁣?

N.Vietnam , prov. Bai Ha, 19-4-62, 6268.

Det. Averyanov L. 1991

Type of Liparis dendrochilum Rchb.f.⁣

Naturhistorisches Museum Wien s.n.

FLORA OF VIETNAM

Plants of Phong Nha - Ke Bang National Park

Fam. Orchidaceae

Liparis dendrochiloides Aver. ?

Vietnam, Quang Binh Prov., Bo Trach Distr., Hung Trach Municipality, Khe Ba Tanh locality, around point 17є27'24''N, 106є23'14''E at elev. 600-700 m a.s.l. Slightly logged primary closed broad-leaved evergreen forest on slopes of mountains composed with shale, sandstone and occasionally granite. Epiphyte (on Dracontomelon duperreanum, Bischofia javanica, Michelia sp., Chisocheton paniculatus, Cinnamomum sp., Pometia pinnata, Pterospermum spp., Duabanga grandiflora) and lithophyte on shady steep slope of stream valley along stream. Very common.

No HAL 6274 30 January 2005.

Collectors: L.Averyanov, P.K.Loc, P.V.The, A.Averyanova, N.T.Vinh, N.Q.Vinh, N.T.Binh.

FLORA OF VIETNAM

Plants of Phong Nha - Ke Bang National Park

Fam. Orchidaceae

Liparis dendrochiloides Aver. ?

Vietnam, Quang Binh Prov., Bo Trach Distr., Hung Trach Municipality, Khe Ba Tanh locality, around point 17є27'24''N, 106є23'14''E at elev. 600-700 m a.s.l. Slightly logged primary closed broad-leaved evergreen forest on slopes of mountains composed with shale, sandstone and occasionally granite. Epiphyte (on Dracontomelon duperreanum, Bischofia javanica, Michelia sp., Chisocheton paniculatus, Cinnamomum sp., Pometia pinnata, Pterospermum spp., Duabanga grandiflora) and lithophyte on shady steep slope of stream valley along stream. Very common.

No HAL 6274 30 January 2005.

Collectors: L.Averyanov, P.K.Loc, P.V.The, A.Averyanova, N.T.Vinh, N.Q.Vinh, N.T.Binh.

Flowered under cultivation in Jan 2011. Flowers odorless; sepals and petals light green; lip yellowish-green; column white, apex green; operculum green with yellow margin. Photos.

Orchidaceae

Liparis sp.

Vietnam. Prov. Hoang Lien Son. Sapa. Takhtajan A. 038, 13.03.1972.

Det. Averyanov L. 1989

FLORA OF VIETNAM
Orchidaceae
Liparis dendrochiloides Aver.

Son La Prov., Van Ho Distr., Xuan Nha Municipality, Muong An village, territory of Xuan Nha natural reserve.

Primary broad-leaved evergreen humid forest on very steep slopes and cliffs of remnant mountain composed by solid highly eroded crystalline limestone at elevation 800-1000 m a.s.l. around point 20°45′06.5″N 104°46′37.8″E.

Creeping lithopyte on open mossy rocks on very steep rocky slope near mountain top. Very common.

13 November 2013 CPC 7083
Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, N.D.Thang, L.D.Qui

FLORA OF VIETNAM
Orchidaceae
Liparis dendrochiloides Aver.

Vietnam, Quang Binh Prov., Le Thuy Distr., Kim Thuy municipality, around Bai Dan village at 16°57′48″N 106°35′34″E.

Primary and old secondary broad-leaved closed forest along ridge composed with shale at elevation 750-1000 m a.s.l.

Epiphyte on mossy tree along streamlet. Locally very common.

HAL 11451 10 April 2008
Coll.: L.Averyanov, P.K.Loc, N.T.Vinh, N.S.Khang

Flowered under cultivation in Jan 2011. Flowers odorless; sepals and petals light green; lip yellowish-green; column white, apex green; operculum green with yellow margin. Photos.

Orchideaceae

Liparis delicatula Hook.f.⁣

Khanh Hoa prov., Khanh Son distr., Son Trung municipality. Coll. Nguyen Thien Tich , T. Gioi, Hong Truong Luu 19.07.2013

Det. Averyanov L. 15.02.2015

FLORA OF VIETNAM
Orchidaceae
Liparis cordifolia Hook.f.

Phu Tho Prov., Tan Son Distr., Xuan Son Municipality, Du village, around point 21°06′57”N 104°57′17”E.

Primary broad-leaved closed wet forest on very steep rocky slopes and cliffs of low remnant mountains composed with highly eroded solid marble-like limestone at elevation 400-500 m a.s.l.

Lithophytic herb on shady mossy boulders in rather wet place. Leaves uniform green. Rare.

HAL 12673 16 February 2009
Coll.: L.Averyanov, P.K.Loc, N.T.Vinh, L.T.Son.

FLORA OF VIETNAM

Orchidaceae

Liparis dendrochiloides Aver.

Vietnam, Quang Binh Prov., Quang Ninh Distr., Truong Son municipality, around point 17º27’43’’N 106º22’50’’E. Primary and secondary broad-leaved forest on steep shale mountain slopes at elevation 500-700 m a.s.l. Lithophyte on mossy shady stream boulders. Locally very common.

HAL 11579 16 April 2008

Coll.: L.Averyanov, P.K.Loc, N.T.Vinh, N.S.Khang

Soviet - Vietnamese Expedition

Orchidaceae

Liparis argenteopunctata m.

Vietnam. Prov. Ha Nam Ninh, Cuc Phuong, May bac. Terrestrial herb in rock crevices. Leaves with silvery spots. 02.05.1985, L. Averyanov et al., LX-VN 1818.

Det. Averyanov L. 1985

FLORA OF VIETNAM
Orchidaceae
Liparis stricklandiana Rchb.f. ?

S.Vietnam, Thua Thien Hue Prov., Xuan Loc Distr., Bach Ma National
Park, steep N slopes of Dong Truoi ridge around point 16°14′11′'N
107°44′28′'E at elev. 300 m a.s.l.

Broad-leaved evergreen secondary closed forest. Lithophytic herb on
vertical shady stream bluffs. Common. In large colonies.

HLF 1819 9 May 2003.

Coll. N.T.Hiep, L.Averyanov, N.T.Vinh et al.
Grant from Henry Luce Foundation.

FLORA OF VIETNAM
Orchidaceae
Liparis dendrochiloides Seidenf.

S.Vietnam, Thua Thien Hue Prov., Phu Loc Distr., Bach Ma National
Park, N slope of Bach Ma mt system, to the N-NW of point 16°12′11′'N
107°50′23′'E at elev. 400-600 m a.s.l.

Broad-leaved evergreen wet primary closed forest on very steep
sandstone, shale and granite slopes along rocky stream canyon.
Lithophytic herb on shady granite stream cliff. Occasional.

HLF 1513 30 April - 1 May 2003.

Coll. N.T.Hiep, L.Averyanov, N.T.Vinh et al.
Grant from Henry Luce Foundation.

FLORA OF VIETNAM

Plants of Phong Nha - Ke Bang National Park

Fam. Orchidaceae

Liparis dendrochiloides Aver. ?

Vietnam, Quang Binh Prov., Bo Trach Distr., Hung Trach Municipality, Khe Ba Tanh locality, around point 17є27'24''N, 106є23'14''E at elev. 600-700 m a.s.l. Slightly logged primary closed broad-leaved evergreen forest on slopes of mountains composed with shale, sandstone and occasionally granite. Epiphyte (on Dracontomelon duperreanum, Bischofia javanica, Michelia sp., Chisocheton paniculatus, Cinnamomum sp., Pometia pinnata, Pterospermum spp., Duabanga grandiflora) and lithophyte on shady steep slope of stream valley along stream. Very common.

No HAL 6274 30 January 2005.

Collectors: L.Averyanov, P.K.Loc, P.V.The, A.Averyanova, N.T.Vinh, N.Q.Vinh, N.T.Binh.

FLORA OF VIETNAM
Orchidaceae
Liparis dendrochiloides Aver.

Quang Binh Prov., Tuyen Hoa Distr., Lam Hoa municipality, Hung village, around point 17°55′41”N 105°49′43”E.

Highly degraded primary and secondary evergreen broadleaved forest on very steep rocky slopes of remnant limestone hills composed with highly eroded marble-like crystalline limestone at elev. 200-300 m a.s.l.

Lithophytic herb in humid wet depression between limestone rocky hills. Not rare.

CPC 2626 1 May 2011
Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, P.V.The, N.T.Vinh.

Soviet - Vietnamese Expedition

Orchidaceae

Liparis argenteopunctata m.

Vietnam. Prov. Ha Nam Ninh, Cuc Phuong, May bac. Terrestrial herb in rock crevices. Leaves with silvery spots. 02.05.1985, L. Averyanov et al., LX-VN 1818.

Fl. et coll in horto 28.08.1985

Det. Averyanov L. 1985

FLORA OF VIETNAM
Orchidaceae
Liparis cordifolia Hook.f.

Lai Chau Prov., Tam Duong Distr., Ho Thau Community, Ho Thau village, around point 22°22′31″N 103°35′15″E.

Secondary and primary evergreen closed forest on very steep mountain slopes composed with granite and quartzite at elevation from 1100 to 1900 m a.s.l.

Lithophyte on vertical mossy shady bluffs along rocky shady wet stream valley. Leaves uniform green. Not common.

HAL 10167 29 Nov. 2006
Coll.: Nguyen Tien Hiep, L.Averyanov, Pham Van The.

FLORA OF VIETNAM
Orchidaceae
Liparis cordifolia Hook.f.

Lai Chau Prov., Phong Tho Distr., Sin Suoi Ho Municipality, Sa Ma Pho village, around point 22°32′53″N 103°33′40″E.

Secondary open evergreen broad-leaved forest and secondary woodlands on very steep mountain slopes composed predominantly with weathered granite at elev. about 1600-1800 m a.s.l.

Lithophytic herb in crevices of shady boulders. Rare.

HAL 10490 11 Dec. 2006
Coll.: Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc, Pham Van The, N.T.Vinh.

FLORA VIETNAMICA

Orchidaceae

Liparis cordifolia Hook.f.

N VIetnam. Prov. Hoang Lien Son, Sapa, 13 03 1972, Takhtajan A. No 040

Det.: Averyanov L. 1989

FLORA OF VIETNAM
Orchidaceae
Liparis cordifolia Hook.f.

N. Viet Nam. Ha Giang Province, Quang Ba District, at Sin Suoi Ho
village on east side of Sin Suoi Ho river. Degraded forest along north
ridge slope. Limestone substrate and exposed rock outcrops at elev.
about 1000-1100 m. 23°06′57′'N, 105°01′48′'E. Lithophyte. Rare.

DKH 5061 3 April 2000

Collectors: D.K.Harder, N.T.Hiep, L.V.Averyanov, N.Q.Hieu, K.Daria.

FLORA OF VIETNAM
Orchidaceae
Liparis cordifolia Hook.f.

N. Viet Nam. Ha Giang Province, Quang Ba District, at Sin Suoi Ho
village and river. Degraded agricultural land among limestone rocks to
ridgetop of exposed limestone under light canopy with many epiphytes
and lithophytes on north and east facing slopes, at 23°06′57″N,
105°01′48″E, at 514-900 m. Leaves green, shiny. Occasional.

DKH 4824 1 April 2000

Collectors: D.K.Harder, N.T.Hiep, L.V.Averyanov, N.Q.Hieu, K.Daria.

Orchideaceae

Liparis cordifolia Hook.f.⁣