Herbarium LE - Specimens

Araceae

Amorphophallus krausei Engl.

Vietnam, sine loc. Flowered in culture on 9 June 2026, inflorescence with straight erect peduncle to 1.2 m tall, flowers with strong unpleasant smell L. Averyanov, AL5044.

Convallariaceae

Ophiopogon spathulifolius N.Tanaka & Aver.⁣

N. Vietnam, Ha Giang province, Bac Me district, Phieng Luong municipality, around Phieng Luong village, primary broad-leaved humid evergreen forest on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation 1100–1150 m a.s.l. around point 22°38’07.1’’N 105°19’22.1’’E, terrestrial and lithophytic rosulate herb on very steep rocky slope near and along ridge edge, common, 13 November 2014, L. Averyanov et al., CPC7419/AL5048.

Flowered in culture on 9 June 2026, L. Averyanov, CPC7419/TM1082/14397.

Araceae

Amorphophallus sp.

Phu Yen Province, Tuy Hoa City, Hoa Xuan Commune. Highly degraded secondary evergreen broad-leaved dry forest with bamboo on very steep rocky mountain slope on granite at elevation of about 400 m a.s.l., around point 12º53’N 109º24’E (Bia Mountain). Terrestrial and lithophytic tuberiferous herb on shady rocky slope. Spathe and inflorescence apex white, flowers yellowish-brown. Odorless. Occasional.

14 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, T. Maisak, АL4907.

Araceae

Amorphophallus sp.

Phu Yen Province, Tuy Hoa City, Hoa Xuan Commune. Highly degraded secondary evergreen broad-leaved dry forest with bamboo on very steep rocky mountain slope on granite at elevation of about 400 m a.s.l., around point 12º53’N 109º24’E (Bia Mountain). Terrestrial and lithophytic tuberiferous herb on shady rocky slope. Spathe and inflorescence apex white, flowers yellowish-brown. Odorless. Occasional.

14 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, T. Maisak, АL4907.

Araceae

Typhonium circinnatum Hett. & Mood

Dak Nong Province, Krong No District, Dray Sap Waterfall, secondary evergreen broad-leaved forest at elevation of about 400 m a.s.l., 15 6 2025, Nguyen Van Canh s.n. Photo prepared from cultivated plant on 14 June 2026, Nguyen Van Canh, AL5047.

All photos © - Nguyen Van Canh

Cycadaceae

Cycas micholitzii Dyer

Dak Lak Province, Lak District, secondary broad-leaved forest with bamboo at elevation of about 600 m a.s.l. 12 June 2026, Nguyen Van Canh, AL5045 (photo only).

All photos © - Nguyen Van Canh

Gesneriaceae

Henkelia sp.?

Lai Chau Province, Muong Te District, Mu Ca Commune. Old rich secondary evergreen broad-leaved forest on stratified shale at elevation of about 540 m a.s.l. around point 22º29’40’’N 102º24’15’’E. Terrestrial or lithophytic rosulate herb in shady place, Flowers blue. Not common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5033.

All photos © - L. Averyanov

Meliaceae

Aglaia sp.?

Lai Chau Province, Muong Te District, Mu Ca Commune. Old rich secondary evergreen broad-leaved forest on stratified shale at elevation of about 540 m a.s.l. around point 22º29’40’’N 102º24’15’’E. Tree to 10 m tall. Fruits orange. Not common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5032.

All photos © - L. Averyanov

Apocynaceae

Amalocalyx microlobus Pierre

Lai Chau Province, Muong Te District, Mu Ca Commune. Old rich secondary evergreen broad-leaved forest on stratified shale at elevation of about 540 m a.s.l. around point 22º29’40’’N 102º24’15’’E. Woody vine to 10 m long in rather open place. Flowers pink-purple. Not common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5031.

All photos © - L. Averyanov

Melastomataceae

Melastoma sp.

Lai Chau Province, Muong Te District, Mu Ca Commune. Old rich secondary evergreen broad-leaved forest on stratified shale at elevation of about 540 m a.s.l. around point 22º29’40’’N 102º24’15’’E. Shrub to 2 m tall in open place. Flowers purple. Locally common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5030.

All photos © - L. Averyanov

Capparidaceae

Capparis assamica Hook.f. & Thomson

Lai Chau Province, Muong Te District, Mu Ca Commune. Old rich secondary evergreen broad-leaved forest on stratified shale at elevation of about 540 m a.s.l. around point 22º29’40’’N 102º24’15’’E. Woody vine to 5 m long. Flowers white, pleasantly fragrant. Not common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5029.

Det.: Silvio Fici, 1 May 2026.

All photos © - L. Averyanov

Rubiaceae

Morinda officinalis F.C.How

Lai Chau Province, Muong Te District, Mu Ca Commune. Old rich secondary evergreen broad-leaved forest on stratified shale at elevation of about 540 m a.s.l. around point 22º29’40’’N 102º24’15’’E. Treelet 2-3 m tall. Flowers white with strong spacy fragrance. Not common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5028.

All photos © - L. Averyanov

Meliaceae

Aglaia sp.?

Lai Chau Province, Muong Te District, Mu Ca Commune. Old rich secondary evergreen broad-leaved forest on stratified shale at elevation of about 540 m a.s.l. around point 22º29’40’’N 102º24’15’’E. Tree 8-10 m tall. Fruits white with pink tint. Common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5027.

All photos © - L. Averyanov

Rubiaceae

Ixora sp.

Lai Chau Province, Muong Te District, Mu Ca Commune. Old rich secondary evergreen broad-leaved forest on stratified shale at elevation of about 540 m a.s.l. around point 22º29’40’’N 102º24’15’’E. Shrub to 4 m tall. Flowers bright red. Not common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5026.

All photos © - L. Averyanov

Rubiaceae

Pavetta sp.?

Lai Chau Province, Muong Te District, Mu Ca Commune. Old rich secondary evergreen broad-leaved forest on stratified shale at elevation of about 540 m a.s.l. around point 22º29’40’’N 102º24’15’’E. Shrub to 3 m tall. Flowers white. Not common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5025.

All photos © - L. Averyanov

Scrophulariaceae

Lindernia sp.

Lai Chau Province, Muong Te District, Mu Ca Commune. Old rich secondary evergreen broad-leaved forest on stratified shale at elevation of about 540 m a.s.l. around point 22º29’40’’N 102º24’15’’E. Creeping herb in wet open place. Flowers light pink, yellow inside. Not common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5024.

All photos © - L. Averyanov

Rubiaceae

Gen. sp.

Lai Chau Province, Muong Te District, Mu Ca Commune. Old rich secondary evergreen broad-leaved forest on stratified shale at elevation of about 540 m a.s.l. around point 22º29’40’’N 102º24’15’’E. Tree 4-6 m tall. Flowers white. Not common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5023.

All photos © - L. Averyanov

Orchidaceae

Anoectochilus roxburghii (Wall.) Lindl.

Lai Chau Province, Muong Te District, Mu Ca Commune. Old rich secondary evergreen broad-leaved forest on stratified shale at elevation of about 540 m a.s.l. around point 22º29’40’’N 102º24’15’’E. Subprostrate terrestrial and lithophytic herb on shady rocky slope. Leaves dark green or brown with network of yellow or pink nerves. Not common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5022.

All photos © - L. Averyanov

Rubiaceae

Spiradiclis sp.⁣

Lai Chau Province, Muong Te District, Mu Ca Commune. Old rich secondary evergreen broad-leaved forest on stratified shale at elevation of about 540 m a.s.l. around point 22º29’40’’N 102º24’15’’E. Subprostrate terrestrial and lithophytic herb on shady rocky slope. Leaves dark green with white median stripe, flowers white apically pink purple. Occasional.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5021.

All photos © - L. Averyanov

Begoniaceae

Begonia sp.

Lai Chau Province, Muong Te District, Mu Ca Commune. Old rich secondary evergreen broad-leaved forest on stratified shale at elevation of about 540 m a.s.l. around point 22º29’40’’N 102º24’15’’E. Erect terrestrial and lithophytic herb to 1 m tall on shady rocky slope. Leaves uniform green. Not common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5020.

All photos © - L. Averyanov

Vitaceae

Tetrastigma sp.?

Lai Chau Province, Muong Te District, Mu Ca Commune. Old rich secondary evergreen broad-leaved forest on stratified shale at elevation of about 540 m a.s.l. around point 22º29’40’’N 102º24’15’’E. Small treelet or woody vine to 1 m long. Not common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5019.

All photos © - L. Averyanov

Balsaminaceae

Impatiens sp.

Lai Chau Province, Muong Te District, Mu Ca Commune. Old rich secondary evergreen broad-leaved forest on stratified shale at elevation of about 540 m a.s.l. around point 22º29’40’’N 102º24’15’’E. Terrestrial and lithophytic herb in wet rocks along stream. Flowers yellow with pink and red. Not common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5018.

All photos © - L. Averyanov

Liliaceae s.l. (Phormiaceae)

Chlorophytum malayense Ridl.

Lai Chau Province, Muong Te District, Mu Ca Commune. Old rich secondary evergreen broad-leaved forest on stratified shale at elevation of about 540 m a.s.l. around point 22º29’40’’N 102º24’15’’E. Terrestrial and lithophytic herb in wet shady slope. Flowers white. Not rare.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5017.

All photos © - L. Averyanov

Amaranthaceae

Amaranthus viridis L.

Lai Chau Province, Muong Te District, Mu Ca Commune. Highly degraded secondary evergreen broad-leaved forest with bamboo along rocky river on stratified shale and limestone at elevation of about 500 m a.s.l. around point 22º24’42’’N 102º24’35’’E. Erect herb to 0.8 m tall on sandy soil among open wet river rocks. Very common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5016 (photo only).

All photos © - L. Averyanov

Asteraceae

Crassocephalum rubens (Jacq.) S.Moore

Lai Chau Province, Muong Te District, Mu Ca Commune. Highly degraded secondary evergreen broad-leaved forest with bamboo along rocky river on stratified shale and limestone at elevation of about 500 m a.s.l. around point 22º24’42’’N 102º24’35’’E. Erect herb to 0.6 m tall on sandy soil among open wet river rocks. Flowers blue. Common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5015 (photo only).

All photos © - L. Averyanov

Polygonaceae

Polygonum aviculare L. ?

Lai Chau Province, Muong Te District, Mu Ca Commune. Highly degraded secondary evergreen broad-leaved forest with bamboo along rocky river on stratified shale and limestone at elevation of about 500 m a.s.l. around point 22º24’42’’N 102º24’35’’E. Prostrate herb on sandy soil among open wet river rocks. Common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5014.

All photos © - L. Averyanov

Polygonaceae

Polygonum aviculare L.

Lai Chau Province, Muong Te District, Mu Ca Commune. Highly degraded secondary evergreen broad-leaved forest with bamboo along rocky river on stratified shale and limestone at elevation of about 500 m a.s.l. around point 22º24’42’’N 102º24’35’’E. Prostrate herb on sandy soil among open wet river rocks. Common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5013.

All photos © - L. Averyanov

Caryophyllaceae

Herniaria sp.

Lai Chau Province, Muong Te District, Mu Ca Commune. Highly degraded secondary evergreen broad-leaved forest with bamboo along rocky river on stratified shale and limestone at elevation of about 500 m a.s.l. around point 22º24’42’’N 102º24’35’’E. Prostrate herb on sandy soil among open wet river rocks. Common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5012.

All photos © - L. Averyanov

Amaranthaceae

Alternanthera sessilis (L.) DC

Lai Chau Province, Muong Te District, Mu Ca Commune. Highly degraded secondary evergreen broad-leaved forest with bamboo along rocky river on stratified shale and limestone at elevation of about 500 m a.s.l. around point 22º24’42’’N 102º24’35’’E. Prostrate herb on sandy soil among open wet river rocks. Common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5011.

All photos © - L. Averyanov

Chenopodiaceae

Suaeda sp.?

Lai Chau Province, Muong Te District, Mu Ca Commune. Highly degraded secondary evergreen broad-leaved forest with bamboo along rocky river on stratified shale and limestone at elevation of about 500 m a.s.l. around point 22º24’42’’N 102º24’35’’E. Erect herb to 0.7 m tall on sandy soil among open wet river rocks. Common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5010.

All photos © - L. Averyanov

Polygonaceae

Rumex sp.

Lai Chau Province, Muong Te District, Mu Ca Commune. Highly degraded secondary evergreen broad-leaved forest with bamboo along rocky river on stratified shale and limestone at elevation of about 500 m a.s.l. around point 22º24’42’’N 102º24’35’’E. Erect herb to 0.5 m tall on sandy soil among open wet river rocks. Common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5009.

All photos © - L. Averyanov

Lythraceae ?

Rotala sp. ?

Lai Chau Province, Muong Te District, Mu Ca Commune. Highly degraded secondary evergreen broad-leaved forest with bamboo along rocky river on stratified shale and limestone at elevation of about 500 m a.s.l. around point 22º24’42’’N 102º24’35’’E. Prostrate herb on sandy soil among open wet river rocks. Stem and flower buds purple. Common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5008.

All photos © - L. Averyanov

Melastomataceae

Sonerila sp.

Lai Chau Province, Muong Te District, Mu Ca Commune. Highly degraded secondary evergreen broad-leaved forest with bamboo along rocky river on stratified shale and limestone at elevation of about 500 m a.s.l. around point 22º24’42’’N 102º24’35’’E. Terrestrial herb to 6 cm tall on shady mossy steep roadcut. Petals, filaments and style purple, anthers pale yellow. Not common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5007.

All photos © - L. Averyanov

Elaeocarpaceae

Elaeocarpus sp.

Lai Chau Province, Muong Te District, Mu Ca Commune. Highly degraded secondary evergreen broad-leaved forest with bamboo along rocky river on stratified shale and limestone at elevation of about 500 m a.s.l. around point 22º24’42’’N 102º24’35’’E. Tree to 6-8 m tall in open place along rocky river. Sepals white, light greenish to pale pink, petals white. Common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5006.

All photos © - L. Averyanov

Selaginellaceae

Selaginella sp.

Lai Chau Province, Muong Te District, Mu Ca Commune. Highly degraded secondary evergreen broad-leaved forest with bamboo along rocky river on stratified shale and limestone at elevation of about 500 m a.s.l. around point 22º24’42’’N 102º24’35’’E. Erect terrestrial herb to 40 cm tall on shady soil cliff along small rocky stream. Locally common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5005.

All photos © - L. Averyanov

Polypodiaceae s.l.

Tectaria zeilanica (Houtt.) Sledge

Lai Chau Province, Muong Te District, Mu Ca Commune. Highly degraded secondary evergreen broad-leaved forest with bamboo along rocky river on stratified shale and limestone at elevation of about 500 m a.s.l. around point 22º24’42’’N 102º24’35’’E. Terrestrial fern on shady soil cliff along river. Common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5004.

All photos © - L. Averyanov