Herbarium LE - Specimens

Paraboea sinensis (Oliv.) B.L. Burtt

Rutaceae

Ha Giang province, Quan Ba district, Tung Vai commune, Thang village. Remnant primary evergreen broad-leaved forest on limestone mountains, around point 23.05186° N 104.86735° E, at elevation of 1000-1050 m a.s.l. Treelet about 5 m tall, flower white, not common, 12 March 2019, Nguyen Sinh Khang et al., NSK 1193.

Urticaceae

Elatostema sp.

Ha Giang province, Quan Ba district, Tung Vai commune, Thang village. Remnant primary evergreen broad-leaved forest on limestone mountains, along stream bank, around point 23.05186° N 104.86735° E, at elevation of 1000-1050 m a.s.l. Terrestrial herb about 20-35 cm tall, flower white, locally common.

12 March 2019, Nguyen Sinh Khang et al., NSK 1189.

Loganiaceae

Mitreola pedicellata Benth.

Ha Giang province, Quan Ba district, Tung Vai commune, Thang village. Remnant primary evergreen broad-leaved forest on limestone mountains, along stream bank, around point 23.05186° N 104.86735° E, at elevation of 1000-1050 m a.s.l. Terrestrial herb about 20-35 cm tall, flower white, locally common.

12 March 2019, Nguyen Sinh Khang et al., NSK 1186.

Rubiaceae

Ophiorrhiza sp.

Ha Giang province, Quan Ba district, Tung Vai commune, Thang village. Remnant primary evergreen broad-leaved forest on steep slopes of limestone mountains, around point 23.05186° N 104.86735° E, at elevation of 1000-1050 m a.s.l. Terrestrial herb about 40-60 cm tall, corolla tube dull white, corolla lobe yellowish green, calyx, corolla densely villous, locally common.

Rubiaceae

Leptomischus funingensis H. S. Lo

Ha Giang province, Quan Ba district, Tung Vai commune, Thang village. Remnant primary evergreen broad-leaved forest on steep slopes of limestone mountains, around point 23.05186° N 104.86735° E, at elevation of 1000-1050 m a.s.l. Terrestrial herb about 40-60 cm tall, corolla tube dull white, corolla lobe yellowish green, calyx, corolla densely villous, locally common.

12 March 2019, Nguyen Sinh Khang et al., NSK 1183.

Lobeliaceae

Lobelia alsinoides Lam.

Ha Giang province, Quan Ba district, Tung Vai commune, Thang village. Remnant primary evergreen broad-leaved forest on slopes of limestone mountains, around point 23.05186° N 104.86735° E, at elevation of 1000-1100 m a.s.l. Terrestrial herb about 20-30 cm tall, flower light blue, inside lower lip with dull white and yellowish green patches, locally common.

12 March 2019, Nguyen Sinh Khang et al., NSK 1180.

Asparagaceae s.l.

Disporum pedunculatum H.Li & J.L.Huang

Ha Giang province, Quan Ba district, Tung Vai commune, Thang village. Remnant primary evergreen broad-leaved forest on slopes of limestone mountains, around point 23.05186° N 104.86735° E, at elevation of 1000-1100 m a.s.l. Terrestrial herb about 70-100 cm tall, inflorescence and leave base dark purple to purple, flower at first green then turning greenish yellow to yellow during the anthesis, sepals green at base and greenish yellow at apex, filament white, anther yellow, locally common.

12 March 2019, Nguyen Sinh Khang et al., NSK 1179.

Acanthaceae

Rungia yunnanesis H.S. Lo

Ha Giang province, Quan Ba district, Tung Vai commune, Thang village. Remnant primary evergreen broad-leaved forest on slopes of limestone mountains, around point 23.05186° N 104.86735° E, at elevation of 1000-1100 m a.s.l. Terrestrial herb about 40-70 cm tall, flower white with purple patches inside lower lip, locally occasion.

12 March 2019, Nguyen Sinh Khang et al., NSK 1178.

Balsaminaceae

Impatiens sp.

Ha Giang prov., Quan Ba distr., Bat Dai Son Nature Reserve, Can Ty commune, Dau Cau 1 village, around point N 23.08650º E105.02312º, elevation of 750-900 m a.s.l., karstic, highly eroded mountains composed of solid marble-like limestone, primary evergreen broad-leaved humid forest on steep rocky slopes near mountain tops. Lithophytic herb, flower white-violetish blue and purple, middle vein silver, common, 14 March 2019, Nguyen Sinh Khang et al., NSK 1210.

Balsaminaceae

Impatiens rubrostriata Hook.f.

Ha Giang province, Quan Ba district, Tung Vai commune, Thang village. Remnant primary evergreen broad-leaved forest on limestone mountains, around point 23.05186° N 104.86735° E, at elevation of 1000-1050 m a.s.l. Terrestrial herb about 50-70 cm tall, flower red or scarlet, common, 12 March 2019, Nguyen Sinh Khang et al., NSK 1198.

Rosaceae

Eriobotrya elliptica

Begoniaceae

Begonia cavalerei?

Ha Giang province, Quan Ba district, Tung Vai commune, Thang village. Remnant primary evergreen broad-leaved forest on limestone mountains, around point 23.05186° N 104.86735° E, at elevation of 1000-1050 m a.s.l. Liana on cliff and tree, flower white, very common, 12 March 2019, Nguyen Sinh Khang et al., NSK 1202.

Begoniaceae

Begonia sp.

Ha Giang province, Quan Ba district, Tung Vai commune, Thang village. Remnant primary evergreen broad-leaved forest on limestone mountains, around point 23.05186° N 104.86735° E, at elevation of 1000-1050 m a.s.l. Terrestrial herb about 50-70 cm tall, flower red or scarlet, common, 12 March 2019, Nguyen Sinh Khang et al., NSK 1199.

Begoniaceae

Begonia sp.

Ha Giang province, Quan Ba district, Tung Vai commune, Thang village. Remnant primary evergreen broad-leaved forest on limestone mountains, along stream bank, around point 23.05186° N 104.86735° E, at elevation of 1000-1050 m a.s.l. Terrestrial herb about 20-35 cm tall, flower white, locally common.

12 March 2019, Nguyen Sinh Khang et al., NSK 1190.

Orchidaceae

Calanthe argenteo-striata C.Z.Tang & S.J.Cheng

Ha Giang prov., Quan Ba distr., Bat Dai Son Nature Reserve, Can Ty commune, Dau Cau 1 village, around point N 23.08650º E105.02312º, elevation of 750-900 m a.s.l., karstic, highly eroded mountains composed of solid marble-like limestone, primary evergreen broad-leaved humid forest on steep rocky slopes near mountain tops. Terrestrial and lithophytic herb, 14 March 2019, Nguyen Sinh Khang et al., NSK 1207d.

Orchidaceae

Paphiopedilum hirsutissimum (Lindl. ex Hook.) Stein var. esquirolei (Schltr.) P.J.Cribb

Ha Giang prov., Quan Ba distr., Bat Dai Son Nature Reserve, Can Ty commune, Dau Cau 1 village, around point N 23.08650º E105.02312º, elevation of 750-900 m a.s.l., karstic, highly eroded mountains composed of solid marble-like limestone, primary evergreen broad-leaved humid forest on steep rocky slopes near mountain tops. Lithophytic herb, 12 March 2019, Nguyen Sinh Khang et al., NSK 1207c.

Orchidaceae

Paphiopedilum malipoense S.C.Chen & Z.H.Tsi

Ha Giang prov., Quan Ba distr., Bat Dai Son Nature Reserve, Can Ty commune, Dau Cau 1 village, around point N 23.08650º E105.02312º, elevation of 750-900 m a.s.l., karstic, highly eroded mountains composed of solid marble-like limestone, primary evergreen broad-leaved humid forest on steep rocky slopes near mountain tops. Terrestrial and lithophytic herb, 12 March 2019, Nguyen Sinh Khang et al., NSK 1207b.

Orchidaceae

Vanda concolor Blume

Ha Giang prov., Quan Ba distr., Bat Dai Son Nature Reserve, Can Ty commune, Dau Cau 1 village, around point N 23.08650º E105.02312º, elevation of 750-900 m a.s.l., karstic, highly eroded mountains composed of solid marble-like limestone, primary evergreen broad-leaved humid forest on steep rocky slopes near mountain tops. Epiphyte, 12 March 2019, Nguyen Sinh Khang et al., NSK 1207a.

Convallariaceae

Aspidistra sp. (A. atropurpurea?)

Son La Prov., Moc Chau Distr., Chieng Son comm., Hin Pen village, around point 20°42’36.7N 104°35’11.0E. Primary and secondary evergreen broad-leaved forest on very steep slopes and cliffs of highly eroded red-brown sandstone at elevation about 1200 m a.s.l. Terrestrial herb in shady rocky place. Very common and abundant from elevation 800 m to the top of mountain. Old, dry flowers dull gray. Leaves uniform green.

29 September 2016 CPC 8136

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.V.Maisak, N.T.Son.

Convallariaceae

Convallariaceae

Aspidistra arnautovii Tillich

Living plant was collected by N. Arnautov (88-115) in Vietnam, Prov. Hai Phong, Cat Ba. The plant was erroneously identified as A. acetabuliformis Y. Wan & C.C. Huang by N. Arnautov in (2002. Clone of the plant from which holotype was prepared. 30 November 2018, E. Lebedev & L. Averyanov, AL 466.

Convallariaceae

Aspidistra

N.Vietnam, Cao Bang Prov., Trung Khanh Distr., Ngoc Khe Municipality, Pac Nga village, around point 22º54'59''N, 106º31'44''E, at elevation 800-850 m a.s.l. Remnants of primary evergreen broad-leaved forest along tops of rocky ridges and mesae composed with light gray solid marble-like highly eroded crystalline limestone. Terrestrial and lithophytic herb. Flowers deep violet. Very common, but very rare in flowers. No HAL 5579 10 June 2004 (Photo) Collectors: L.Averyanov, P.K.Loc, P.V.The, N.T.Vinh.

Juglandaceae

Platycarya strobilacea

N.Vietnam, Cao Bang Prov., Trung Khanh Distr., Ngoc Khe Municipality, Pac Nga village, around point 22º54'59''N, 106º31'44''E, at elevation 800-850 m a.s.l. Remnants of primary broad-leaved evergreen and coniferous forest along rocky tops of rocky ridges and mesae composed with light gray solid marble-like highly eroded crystalline limestone. Tree 10-15 m tall. Inflorescences yellow-green. Very common. Co-dominant of 1-2 forest stratum.

No HAL 5618 11 June 2004

Collectors: L.Averyanov, P.K.Loc, P.V.The, N.T.Vinh.

All photos © L. Averyanov

Araceae

Typhonium sp.

Quang Binh Prov., Tuyen Hoa Distr., Lam Hoa municipality, Hung village, around point 17°55’41”N 105°49’43”E. Secondary shrubs along small river on flat alluvial valley between remnant limestone hills composed with highly eroded marble-like crystalline limestone at elev. About 100 m a.s.l. Terrestrial tuberiferous herb on short tall secondary grassland. Spathe dark velvety purple-brown. Locally very common.

CPC 2631 1 May 2011 Photos.

Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, P.V.The, N.T.Vinh.

Araceae

Typhonium sp.?

Vietnam, Nghe An Prov., Tuong Duong Distr., Tam Quang Municipality, Tung Huong village, along Khe Mat stream around point 19º03’09’’N 104º36’53’’E at elevation about 300-350 m a.s.l. Lowland closed secondary evergreen forest on steep slopes along rocky stream valley composed with stratified solid shale and sandstone. Lithophytic herb on crevices of river bluffs. Spathe purple-brown, flowers white, strong unpleasant smelled. Locally common.

HLF 6887 21 March 2007. Photo.

Coll. Averyanov L., P.K.Loc, N.V.Trai, N.T.Vinh, N.D.Phuong, Averyanova A.

Grant from Henry Luce Foundation.

Araceae

Symplocarpus foetidus

Russia, Vladivostok a.1989, L. Averyanov s.l.

Araceae

Steudnera sp.

N.Vietnam, Ha Tinh Prov., Huong Son Distr., Son Kim Municipality, Rao Bun stream around point 18º23'39''N, 105º15'12''E. Primary broad-leaved evergreen closed wet forest on very steep shale slopes of river canyon at elevation about 300 m a.s.l. Lithophytic herb on shady wet stream rocks, Spathe dull light yellow-brownish outside, deep brown-violet inside, flowers white, very fragrant. Common.

No HAL 5007 2 May 2004 (Photo)

Collectors: P.K.Loc, L.Averyanov, P.V.The, N.T.Vinh.

Araceae

Steudnera discolor W. Bull

Luang Prabang prov., Ngoy distr., Don Khun village, near waterfall, around point 20°31'36.9''N 102°35'15.8''E. Remnants of primary evergreen forest on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. 450-550 m a.s.l. along mountain stream. Terrestrial herb on shady rocky stream slope. Spathe purple-brown. Very common along stream.

LA-VN 2017 2 April 2017

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.Maisak, Vichith Lamxay, Koudkeo Phommachanh, Phizar Sayalath, Keooudone Souvannakhoummane.

Rubiaceae

Acranthera hoangii Hareesh & T.A.Le

Vietnam, Thua Thien – Hue Prov., Nam Dong Distr., Huong Son Municipality, around point 16°10'22''N, 107°36'24''E. Fractionally logged primary closed broad-leaved evergreen lowland forest along La Ma River on very steep hill slopes composed with clayey shale and sandstone at elevation about 300 m a.s.l. Undershrub or shrub 1-2 m tall on shady rocky stream slope. Flowers light lemon yellow. Occasional.

No HAL 7011 28 March 2005. Photo.

Collectors: L.Averyanov, N.T.Hiep, P.K.Loc, P.V.The, A.Averyanova, N.T.Vinh.

Rubiaceae

Acranthera hoangii Hareesh & T.A. Le

Thua Thien – Hue Prov., Nam Dong Distr., Thuong Lo Municipality, along Cha Mang stream, around point 16°07’57’’N 107°44’16’’E. Primary evergreen broad-leaved lowland forest on very steep hill slopes and along mountain ridge composed with stratified shale and sandstone at elev. about 450-550 m a.s.l. Undershrub 1-2 m tall on steep wet shady slope. Flowers light yellow. Not common.

HAL 10931 5 Apr. 2007. Photo.

Coll.: L.Averyanov, Phan Ke Loc, A.Averyanova, N.D.Phuong, L.V.Hung.

All photos © L. Averyanov

Rubiaceae

Acranthera sinensis

Vietnam, Nghe An Prov., Tuong Duong Distr., Tam Hop Municipality, Pa Lom village, around point 19º05’07’’N 104º20’55’’E at elevation 1300-1350 m a.s.l. Submontane primary evergreen closed forest on very steep mountain slopes composed with stratified solid shale. Terrestrial herb up to 2 m tall on very steep rocky shady mountain slope along forest stream. Flowers white, dark bright purple inside tube. Locally very common.

HLF 7061 25 March 2007. Photo.

Coll. Averyanov L., P.K.Loc, N.V.Trai, N.T.Vinh, N.D.Phuong, Averyanova A.

Grant from Henry Luce Foundation.

All photos © L. Averyanov

Araceae

Scindapsus sp.

Thua Thien – Hue Prov., Nam Dong Distr., Thuong Lo Municipality, along Cha Mang stream, around point 16°07’46’’N 107°44’24’’E. Remnants of primary evergreen broad-leaved lowland forest on very steep hill slopes composed with stratified shale and sandstone at elev. about 250-300 m a.s.l. Creeping epiphytic vine on shady rocky stream slopes. Flowers and spathe light greenish. Occasional.

HAL 10933 5 Apr. 2007. Photo.

Coll.: L.Averyanov, Phan Ke Loc, A.Averyanova, N.D.Phuong, L.V.Hung.

Araceae

Scindapsus poilanei Gagnep.

Quang Binh Prov., Minh Hoa Distr., Thuong Hoa municipality, environs of Mo O O O village, around point 17°40’03.6”N 105°55’08.5”E. Dense broad-leaved closed forest on wet flat seasonally flooded river valley on alluvial limestone clay at elevation about 336 m a.s.l. Small lithophytic and epiphytic vine to 1 m long on rocky slope. Spathe dark green. Locally very common.

CPC 3910 29 July 2011

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Vinh.

Araceae

Remusatia vivipara (Roxb.) Schott

Vientiane prov., Kasi distr., between Pa Chao and Nam Ken villages, Pa Chao Mt., around point 19°19'50.6''N 102°21'20.5''E. Secondary broad-leaved evergreen dry forest on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. 1000-1100 m a.s.l. Lithophyte on shady vertical cliff. Spathe bright yellow. Common.

LA-VN 1693 26 March 2017 Photos.

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.Maisak, Khamla Inkhavilay, Hongthong Southyvohanh, Pinmany Sisavath, Chanthala Cheuaysamphanh.

Araceae

Remusatia vivipara (Roxb.) Schott

Dien Bien Prov., Tua Chua Distr., Tua Thang Municipality, Ta Xi Lang village, around point 21°57’17”N 103°25’31”E. Highly degraded primary and secondary evergreen broadleaved forest on very steep slopes and on tops of remnant mountains composed with solid limestone at elevation 850-950 m a.s.l. Lithophyte on mossy shady rocks on very steep rocky slope. Spathe yellow. Not rare.

CPC 2307 13 April 2011 Photos.

Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, N.T.Vinh.

Araceae

Raphidophora sp.

N.Vietnam, Bac Kan Prov., Cho Don Distr., Ban Thi Municipality, village Phia Khao, Lung Li valley, around point 22º16'10''N, 105º31'01''E, Cot Moc mountain at elevation 800-900 m a.s.l. Primary broad-leaved evergreen closed forest on very steep slopes and cliffs near tops of rocky ridge composed with solid marble-like stratified limestone. Creeping lithophytic and epiphytic vine. Spathe dull light brown-yellow, flowers white. Very common. Co-dominant of herbaceous lithophytic communities on lower and middle part of mountain slopes.

No HAL 4808 23 May 2004 (Photo).

Collectors: L.Averyanov, N.T.Hiep, P.V.The, N.T.Vinh.

Araceae

Rhaphidophora decursiva (Roxb.) Schott

Quang Binh Prov., Minh Hoa Distr., Thuong Hoa municipality, environs of Mo O O O village, around point 17°40’03.6”N 105°55’08.5”E. Dense broad-leaved closed forest along highly eroded rocky limestone ridge at elevation about 500 m a.s.l. Lithophytic vine to 5 m long on exposed rocks along ridge tops. Spathe dull yellow. Very common.

CPC 3915 29 July 2011

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Vinh.

Araceae

Raphidophora decursiva

N.Vietnam, Thanh Hoa, Co Lung HAL s.n. a. 2003, L. Averyanov s.n.

All photos © L. Averyanov

Araceae

Raphidophora decursiva

Cao Bang Province, Bao Lac, Son Lo & Son Lap, CPC s.n., 19 03 2013, L. Averyanov, s.n.

All photos © L. Averyanov

Araceae

Pseudodraconthium sp.?

Northern Vietnam, Ninh Binh Prov., Nho Quan Distr., Cuc Phuong Municipality, Cuc Phuong National Park, Dang locality, about 1 km WNW of Dang Forest Control point, 20ø16'28''N, 105ø40'57''. Secondary closed evergreen seasonal broad-leaved lowland forest on alluvial valley between limestone hills at elev. about 300 m a.s.l. Terrestrial and lithophytic herb with inflorescence up to 25 cm lg. Spathe light green, inflorescence white with green tint. Occasional.

No HAL 1640 (Photo) 19 May 2002

Collectors: L.Averyanov, P.K.Loc, M.V.Sinh, N.X.Tam, N.T.Vinh.

FLORA OF VIETNAM

Araceae

Pseudodracontium lacourii

S.Vietnam, Kien Giang, Tho Chu, a.1989, L. Averyanov s.n.

Araceae

Pseudodracontium anomalum

Vietnam, s. loc., cult. Komarov Bot. Gard.,1986, L. Averyanov s.n.

Araceae

Pothos sp.

Quang Binh Prov., Minh Hoa Distr., Dan Hoa Municipality, Bai Dinh village around point 17°45’28”N 105°46’14”E. Remnants of primary evergreen broad-leaved forest on very steep slopes and on tops of low remnant hills composed with highly eroded marble-like solid limestone at elevation about 200-300 m a.s.l. Creeping lithophytic vine to 15 m long on exposed tall vertical limestone cliff. Flowers and fruits white, spathe green to olive-green. Locally very common.

HAL 12368 7 February 2009 Photo.

Coll.: L.Averyanov, P.K.Loc, N.T.Vinh, L.T.Son.

Araceae

Pothos sp.

Vietnam, Thua Thien – Hue Prov., Nam Dong Distr., Huong Son Municipality, around point 16°10'22''N, 107°36'24''E. Fractionally logged primary closed broad-leaved evergreen lowland forest along La Ma River on very steep hill slopes composed with clayey shale and sandstone at elevation about 300 m a.s.l. Herbaceous and semi-woody creeping epiphytic vine up to 15 m long on tall trees along rocky stream valley. Inflorescence yellow. spathe olive-brown. Common.

No HAL 7079 30 March 2005. Photo.

Collectors: L.Averyanov, P.K.Loc, P.V.The, A.Averyanova, N.T.Vinh.

Araceae

Pothos sp.

N.Vietnam, Ha Tinh Prov., Huong Son Distr., Son Kim Municipality, Rao Bun stream around point 18º23'39''N, 105º15'12''E. Primary broad-leaved evergreen closed wet forest on very steep shale slopes of river canyon at elevation about 300-400 m a.s.l. Creeping epiphytic vine 15-20 m long. Flowers white to yellow, spathe

olive-violet. Very common.

No HAL 5109 4 May 2004 (Photo)

Collectors: P.K.Loc, L.Averyanov, P.V.The, N.T.Vinh.

Araceae

Pothos repens

Northern Vietnam, Ha Tinh Prov., Huong Son Distr., Son Kim Municipality, in the vicinity of Rao An village, around point 18ø22'00''N, 105ø13'13''E. Primary evergreen lowland seasonal broad-leaved forest on granite along rocky river valley at 240-260 m a.s.l. Creeping epiphyte vine on high tree. Fruits red. Very common.

No HAL 1288 (Photo) 29 April 2002

Collectors: P.K.Loc, L.Averyanov, N.X.Tam, N.T.Vinh.

All photos © L. Averyanov

Araceae

Pothos grandis

Northern Vietnam, Ninh Binh Prov., Nho Quan Distr., Cuc Phuong Municipality, Cuc Phuong National Park, Dang locality, about 1 km WNW of Dang Forest Control point, 20ø16'28''N, 105ø40'57''. Secondary closed evergreen seasonal broad-leaved lowland forest on alluvial valley between limestone hills at elev. about 300 m a.s.l. Lithophytic and epiphytic creeping vine. Fruits brightly red. Common.

No HAL 1641 (Photo) 19 May 2002

Collectors: L.Averyanov, P.K.Loc, M.V.Sinh, N.X.Tam, N.T.Vinh.

All photos © L. Averyanov

Araceae

Pothos grandis

N.Vietnam, Ha Tinh Prov., Huong Son Distr., Son Hong Municipality, around point 18º34'06''N, 105º11'40''E. Primary broad-leaved evergreen forest on steep mt slopes composed with granite along Khe Van stream at elev 400-450 m a.s.l. Creeping epiphytic vine. Not common.

No HAL 5217 8 May 2004

Collectors: P.K.Loc, L.Averyanov, P.V.The, N.T.Vinh.

Araceae

Pothos grandis

Vietnam, Ha Giang Prov., Hoang Su Phi Distr., Ho Thau Municipality, around Chien Thang village, about 4 km to NW of Ho Thau village (22º37’24’’N, 104º38’00’’E) at elev. 1200-1500 m a.s.l. Remnants of heavily logged primary and secondary closed evergreen broad-leaved forest on very steep slopes of mountains composed with shale and granite with quartzite. Lithophytic and epiphytic vine 5-8 m long in open place. Flowers yellowish-white, spathe green. Common.

No HAL 6633 10 March 2005. Photo.

Collectors: L.Averyanov, P.K.Loc, N.T.Vinh, A.Averyanova.