Herbarium LE - Specimens

FLORA OF VIETNAM

Fam. Balanophoraceae Rich.

Balanophora fungosa J.R.Forst.et Forst.

S. Vietnam, Prov. Kontum. Evergreen primary forest along stream on N slope of Ngoc Linh mountain system at 1700 m alt. Reddish purple parasite on roots of Saurauia sp. Not rare.

No VH 737 13.03.1995

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 5094-93)

Participants: L. Averyanov, N.T. Ban, A. Budantzev, L. Budantzev, L. Budantzev, N.T. Hiep, D.D. Huyen, P.K. Loc, N.X. Tam, G. Yakovlev

Orchidaceae

Coelogyne sanderae

S. Vietnam, Lam Dong Province, Dalat area, LX-VN s.n., Garden number 0-110 a. 1984

L. Averyanov

Photos © L. Averyanov

FLORA OF VIETNAM

Fam. Balanophoraceae Rich.

Balanophora fungosa J.R.Forst.et Forst.

S. Vietnam, Prov. Kontum. Evergreen primary forest along stream on N slope of Ngoc Linh mountain system at 1700 m alt. Reddish purple parasite on roots of Saurauia sp. Not rare.

No VH 737 13.03.1995

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 5094-93)

Participants: L. Averyanov, N.T. Ban, A. Budantzev, L. Budantzev, L. Budantzev, N.T. Hiep, D.D. Huyen, P.K. Loc, N.X. Tam, G. Yakovlev

Fam. Acanthaceae

N. Vietnam, prov. Ha Giang, distr. Yen Minh, municipality Du Gia, E slopes of Phu Tha Ca mountain system at 1900-2000 m elev. (N 23ш56' E 105ш13'). Primary evergreen broadleaved very wet mossy cloud forest on

quartzite very steep slopes. Terrestrial and lithophyte herb on mossy quartzite cliffs. Common.

No CBL 2157 03 May 1999

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 6300-98)

Collectors: P.K.Loc, P.H.Hoang, Averyanov L.

Fam. Acanthaceae

N. Vietnam, prov. Lao Cai, distr. Than Uyen, municipality Ho Mit (N 22є06' E 103є52'). Elevation to Mang Han San quartzite peak by SW slope from Ban Thao village (SSW macroslope of Hoang Lien Son ridge).

Primary wet evergreen broadleaved forest on very steep rocky slope at 1900 m elev. Lithophyte forb on wet shady bluffs. Common.

No NTH 2834 21-22 May 1999

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 6300-98)

Collectors: N.T.Hiep, P.H.Hoang, Averyanov L.

FLORA OF VIETNAM

Fam. Acanthaceae

Staurogyne chapaensis Benoist.⁣

Lao Cai Prov.

Phan Ke Loc et al.

P 7359, 20.04.1996

Fam. Acanthaceae

?

S. Vietnam, Prov. Kon Tum, Distr. Dak Gley, about 10-14 km to the N of Dak Gley town. Primary wet evergreen mountain forest with domination of Rhodoleia championi at 900-1000 m alt. Shrub up to 0.5 m high. Flowers white. Rare.

No VH 2154 30.11.1995

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 5094-93)

Participants: L. Averyanov, N.T. Ban, A. Budantzev, L. Budantzev, N.T. Hiep, D.D. Huyen, P.K. Loc, G. Yakovlev

Staurogyne major R.Ben.

S. Vietnam, Prov. Ninh Thuan, distr. Ninh Son, municipalite Phuoc Binh, 37 km to NE from Dalat city. Closed primary broadleaved wet mt forest on E macroslope of Gia Rinh mt. ridge at 1700-1800 m alt. (N 12° 06', E 108° 43'). Shrublet up to 1 m hg. Flowers light blue-violet. Common.

No VH 3514 01.04.1997

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 5803-96)

Collectors: L. Averyanov, N.Q. Binh, P.K. Loc

Staurogyne major R.Ben

S. Vietnam, Prov. Lam Dong, distr. Lac Duong, municipalite Da Chay, 32.5 km to NE from Dalat city. Closed primary broadleaved mt forest along mt ridge in 5-6 km to NNW from main Bi Dup peak at 1800-1900 m alt. (N 12° 07', E 108° 41'). Herb up to 0.3 m hg. Flowers light blue. Common. purple-red. Common.

No VH 3335 29.03.1997

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 5803-96)

Collectors: L. Averyanov, N.Q. Binh, P.K. Loc

Acanthaceae

Staurogyne major Benoist

Det.: Yun-Fei, Deng 2008

S. Vietnam, Prov. Lam Dong, 31 km to NE from Dalat city, distr. Lac Duong, municipalite Da Chay, vicinities Klong Lanh village (N 12° 08', E 108° 39'). Closed primary broadleaved mt forest on NW macroslope of

Bi Dup mt system at 1850-1950 m alt. Terrestrial herb on wet shadow places. Flowers white. Not rare.

No VH 2644 15.03.1997

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 5803-96)

Collectors: L. Averyanov, N.Q. Binh, N.V. Duy, P.K. Loc

Fam. Acanthaceae Juss.

?

S. Vietnam, Prov. Quangnam-Danang. Evegreen close primary forest on SE slope of Ngoc Linh mountain system at 1700 m alt. Herb shrub up to 1.5 m high. Flowers yellowish white. Not rare and sometimes dominante on SE and S slopes of Ngoc Linh mountain system at 1400-1700 m alt. (rare on N, NW and W slopes).

No VH 906 24.03.1995

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 5094-93)

Participants: L. Averyanov, N.T. Ban, A. Budantzev, L. Budantzev, N.T. Hiep, D.D. Huyen, P.K. Loc, G. Yakovlev

Fam. Acanthaceae

Gen.?

Vietnam, Thua Thien – Hue Prov., Nam Dong Distr., Huong Son Municipality, around point 16°09'58''N, 107°36'07''E. Fractionally logged primary closed broad-leaved evergreen lowland forest along La Ma River on very steep hill slopes composed with clayey shale and sandstone at elevation 300-400 m a.s.l. Lithophytic and terrestrial herb on rocky steep hill slope. Flowers light blue. Locally common.

No HAL 7146 31 March 2005. Photo.

Collectors: L.Averyanov, P.K.Loc, P.V.The, A.Averyanova, N.T.Vinh.

Acanthaceae

Staurogyne sp.?

Quang Nam Prov., Dai Loc Distr., Dai Hong municipality, environs of Dai Hong town around point 15°49’19”N 107°56’58”E. Rich secondary broadleaved evergreen forest on very steep rocky slopes and cliffs composed with eroded sandstone at elev. 400-500 m a.s.l. Terrestrial rosulate herb among large shady mossy boulders on very steep humid rocky slope. Flower white. Common.

CPC 3423 19 May 2011

Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, P.V.The, N.T.Vinh.

FLORA OF VIETNAM

Acanthaceae

Staurogyne sp.⁣?

Quang Nam Prov.

Averyanov L., et al.

CPC 3205b, 14.05.2011

Acanthaceae

Nghe An Prov., Ky Son Distr., Nam Can municipality, along the road to international Gate (Vietnam - Laos) Nam Can. Open secondary forest, woodlands and open secondary scrub at elevation 750-1000 m a.s.l. around point 19°27’19.4’’N 104°06’25.8’’E. Terrestrial herb in wet open place. Flowers vuolet. Common.

29 October 2013 CPC 6395

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, L.M.Tuan

Acanthaceae

Staurogyne sp.

Vietnam, Quang Binh Prov., Quang Ninh Distr., Truong Son municipality, around point 17є20’20’’N 106є27’39’’E.

Primary and secondary broad-leaved forest on steep shale hill slopes along Khe Viet stream at elevation about 70 m a.s.l. Lithophytic rosulate herb on shady wet shale rocky steep slope along stream. Flowers white. Occasional.

HAL 11553 14 April 2008

Coll.: L.Averyanov, P.K.Loc, N.T.Vinh, N.S.Khang

Fam. Acanthaceae

Gen.?

Vietnam, Thua Thien – Hue Prov., Huong Thuy Distr., Duong Hoa Municipality, Huong Thuy forest enterprise territory, around point 16°13'20''N, 107°34'23''E, SW slopes of Mang Chan ridge at elevation 500-600 m a.s.l.

Fractionally logged primary closed evergreen broad-leaved forest along steep slopes composed with shale.

Terrestrial herb on shady wet slopes along stream. Flowers white. Very common.

No HAL 8080 10 May 2005. Photo.

Collectors: L.Averyanov, P.K.Loc, T.V.Thao, N.T.Vinh.

Balanophoraceae

Balanophora fungosa J.R.Forst. & G.Forst.

Lam Dong Prov.

16.04.1984

Det. Averyanov L. 1991

Fam. Balanophoraceae

Balanophora abbreviata Blume

S. Vietnam, Prov. Kon Tum, Distr. Dak Gley, about 15-18 km to NW of Dak Gley town, near Mang Khen (Dak Che) village. Primary evergreen mountain very wet forest in deep stream canyon at 1000 m alt. Reddosh-brown root parasite (on roots of Poikilospermum sp.). Not common.

No VH 1942 23.11.1995

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 5094-93)

Participants: L. Averyanov, N.T. Ban, A. Budantzev, L. Budantzev, N.T. Hiep, D.D. Huyen, P.K. Loc, G. Yakovlev

Fam. Balanophoraceae Rich.

Balanophora fungosa J.R.Forster et Forster

S. Vietnam, Prov. Kontum. NW slopes of Ngoc Linh mountain system at 1600 m alt. Shadow secondary forest along mountain stream. Parasite (on roots of Saurauia sp. ?). Inflorescence purple.

No VH 326 02.03.1995

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 5094-93)

Participants: L. Averyanov, N.T. Ban, N. Q. Binh, A. Budantzev, L. Budantzev, N.T. Hiep, D.D. Huyen, P.K. Loc, N.X. Tam, G. Yakovlev

Fam. Balanophoraceae

Balanophora fungosa ?

S. Vietnam, Prov. Kon Tum, Distr. Dak Gley, about 10-12 km to the SE of Dak Gley town (4-6 km to the E of Dak Tung village). Primary evergreen mountain mixed (with Pinus khesya) forest at 1100-1200 m alt. Large reddish root parasite up to 0.2 m high. Very rare.

No VH 2383 06.12.1995

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 5094-93)

Participants: L. Averyanov, N.T. Ban, A. Budantzev, L. Budantzev, N.T. Hiep, D.D. Huyen, P.K. Loc, G. Yakovlev

Balanophora fungosa J.R. et G.Forst.

S. Vietnam, Prov. Lam Dong, distr. Lac Duong, municipalite Da Chay, 32.5 km to NE from Dalat city. Closed primary broadleaved mt forest along mt ridge in 5-6 km to NNW from main Bi Dup peak at 1800-1900 m alt. (N 12° 07', E 108° 41'). Reddish root parasite. Not common.

No VH 3331 29.03.1997

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 5803-96)

Collectors: L. Averyanov, N.Q. Binh, P.K. Loc

Fam. Balanophoraceae

Balanophora latisepala (Tiegh.) Lecomte⁣.

Khammouan Prov.

Konstantinov E.

K 215, 30.10.2013

Det. Nuraliev M. 17.09.2026

Fam. Balanophoraceae

Balanophora sp. (male samples)

N. Vietnam, Ha Giang prov., Meo Vac distr., vicinities of Meo Vac town (N 23є10' E 105є24'), 48.2 km to 36є of Ha Giang city. Primary evergreen broadleaved wet closed forest on sandstones at elev. 1400-1600 m a.s.l. Brightly red root parasite on very steep shady slope. Not common.

No NTH 3410 07 Oct. 1999

Collectors: N.T. Hiep, N.Q. Binh, L. Averyanov, P. Cribb.

Balanophora sp.

S. Vietnam, Prov. Ninh Thuan, distr. Ninh Son, municipalite Phuoc Binh in 34 km to NE from Dalat city. Closed primary broadleaved forest along mt ridge between Bi Dup and Gia Rinh mt. mt at 1800-1900 m alt.

(N 12° 07', E 108° 42'). Root chlorophyll less yellowish parasite on Tetrastigma sp. ? Not common.

No VH 3579 03.04.1997

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 5803-96)

Collectors: L. Averyanov, N.Q. Binh, P.K. Loc

Balabophoraceae

Balanophora sp.

Cao Bang Prov., Nguyen Binh Distr., Ca Thanh Municipality, Cao Lu village, around point 22°42'47.7"N, 105°51'50.8"E. Primary coniferous forest with Pseudotsuga sinensis along highly eroded rocky limestone ridge at elevation about 1300 m a.s.l. Terrestrial root chlorophyll less red parasite on shady rocky slope. Not common.

4 October 2013 CPC 5391

L.Averyanov, N.T.Hiep, L.M.Tuan, N.S.Khang, T.Maisak, L.Osinovets

Balanophoraceae

Balanophora sp.

Khammouan Prov.

Konstantinov E.

K 254, 03.11.2013

Det. Nuraliev M. 17.09.2026

Balanophoraceae

Balanophora sp.

Son La Prov., Thuan Chau Distr., Co Ma Municipality, Hua Ty village, Copia nature reserve, around point 21є20’07’’N 103є35’07’’E. Remnants of broad-leaved (former coniferous with Fokienia hodginsii) primary and secondary forest on steep slopes and on tops of ridge composed with clay shale at elev. 1500-1600 m a.s.l.

Terrestrial achlorophyllous root brightly red parasite on shady wet stream slope. Rare.

HAL 9725 12 Nov. 2006

Coll.: Nguyen Tien Hiep, L.Averyanov, Pham Van The.

Plants of Ta Dung Nature Reserve

Balanophoraceae

Balanophora sp.

Vietnam, Dak Nong Prov., Dak Glong Distr., Dak Plao Municipality, steep mountain slopes along main ridge of Ta Dung mountain system at elevation 1700-1985 m a.s.l. around point of highest peak at 11є51’30’’N 108є02’08’’E.Primary broad-leaved evergreen closed wet mossy cloudy forest on crystalline highly metamorphosed sandstone. Ground root achlorophyllous pure white parasite on shady slope. Flowers white. Occasional.

HLF 5653 9 November 2005.

Coll. L.Averyanov, P.K.Loc, T.V.Thao, N.T.Vinh.

Grant from Henry Luce Foundation.

Balanophoraceae

Balanophora sp.

Lam Dong Prov.

Averyanov L., et al.

LX-VN 1118, 31.12.1982

Balanophoraceae

Balanophora sp.

Vietnam.

Averyanov L., et al.

LX-VN 1072, 29.12.1982

Balanophoraceae

Balanophora sp.

Vietnam.

Averyanov L., et al.

LX-VN 1117, 31.12.1982

Plants of Phong Nha - Ke Bang National Park

Fam. Balanophoraceae

Balanophora sp.

Vietnam, Quang Binh Prov., Bo Trach Distr., Tan Trach Municipality, vicinities of A Rem village (17є23'32''N, 106є12'46''E). Heavily logged primary closed evergreen broad-leaves forest on steep slopes of rocky ridge composed with solid marble-like highly eroded stratified crystalline limestone at elevation about 650-750 m a.s.l.

Chlorophylless ground white parasite. Common.

No HAL 6105 24 January 2005. Photo.

Collectors: L.Averyanov, P.K.Loc, P.V.The, A.Averyanova, N.T.Vinh, N.Q.Vinh, N.T.Binh.

Fam. Balanophoraceae Rich.

Balanophora laxiflora Hemsl.

S. Vietnam, Prov. Kontum. Evergreen primary forest at 1800-1900 m alt. on NW slope of Ngoc Linh mountain system on elevation to Ngoc Gua peak. Yellowish pink parasite (on roots of Fagaceae Dumortier ssp.?).

No VH 1237 09.04.1995

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 5094-93)

Participants: L. Averyanov, N.T. Ban, N. Q. Binh, A. Budantzev, L. Budantzev, N.T. Hiep, D.D. Huyen, P.K. Loc, N.X. Tam, G. Yakovlev

Fam. Balanophoraceae

Rhopalocnemis phalloides ?

S. Vietnam, Prov. Kon Tum, Distr. Dak Gley, about 12 km to N of Dak Gley town (24 km by road), near Mang Khen village. Secondary evergreen mountain forest rich in primary elements at 1000-1100 m alt. Root parasite. Rare.

No VH 1867 21.11.1995

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 5094-93)

Participants: L. Averyanov, N.T. Ban, A. Budantzev, L. Budantzev, N.T. Hiep, D.D. Huyen, P.K. Loc, G. Yakovlev

Fam. Balanophoraceae

Rhopalocnemis phalloides ?

S. Vietnam, Prov. Kon Tum, Distr. Dak Gley, about 12 km to N of Dak Gley town (24 km by road), near Mang Khen village. Primary wet mountain evergreen forest on slope of N exposition at 1100-1200 m alt. Root parasite. Rare.

No VH 1662 14.11.1995

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 5094-93)

Participants: L. Averyanov, N.T. Ban, A. Budantzev, L. Budantzev, N.T. Hiep, D.D. Huyen, P.K. Loc, G. Yakovlev

Fam. Balanophoraceae Rich.

Balanophora fungosa Forst.

S. Vietnam, Prov. Kontum. Evergreen close primary forest on SE slope of Ngoc Linh mountain system at 2100 m alt., on the boundary of Kontum and Quangnam-Danang provinces. Parasite (on Fagaceae sp. roots ?). Male samples.

No VH 876 22.03.1995

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 5094-93)

Participants: L. Averyanov, N.T. Ban, N. Q. Binh, A. Budantzev, L. Budantzev, N.T. Hiep, D.D. Huyen, P.K. Loc, N.X. Tam, G. Yakovlev

Fam. Balanophoraceae

Rhopalocnemis phalloides ?

S. Vietnam, Prov. Kon Tum, Distr. Dak Gley, about 15-18 km to NW of Dak Gley town, between Mang Khen (Dak Che) and Dak Polo villages, near Laos boundary. Primary evergreen mountain forest at 1250-1450 m alt.

Yellowish root parasite (on Castanopsis sp.?). Rare.

No VH 1929 22.11.1995

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 5094-93)

Participants: L. Averyanov, N.T. Ban, A. Budantzev, L. Budantzev, N.T. Hiep, D.D. Huyen, P.K. Loc, G. Yakovlev

Acanthaceae

Staurogyne sp. nov.?

Herbarium specimen prepared on 22 December 2025,

by Fedor V. Kondrachuk, from cultivated plants originated

from Central Vietnam without indication of habitat and geographical location.

22 December 2025, Fedor V. Kondrachuk, sine n.

Upland. Steninge. Vajo. Leg. A. M. Fydberg.

A Smithii C. solida (male C. Halleri dicta) Deffert ut

C. Fabacea a C. pumila, ut speciem forsan dicas.

C. laxa Fries Isotypus

(= C. solida / pumila, introgressive hybridir.)

Teste Liden