Herbarium LE - Specimens

Asteraceae

Blumea sp.

Vientiane Prov., Kasi Distr., Thong Mout village, around point 19°25’19.7”N 102°08’32.7”E. Primary broad-leaved evergreen forest on very steep slopes and on cliffs of remnant mountain composed with solid crystalline highly eroded limestone at elevation about 1550 m. Terrestrial herb to 1.5 m tall on open alluvial slope among secondary shrubs. Flowers yellow. Common.

LA-VN 669 20 March 2013

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, P.V.The, S.Lorphengsy

Asteraceae

Vernonia sp.?

Vientiane Prov., Kasi Distr., Thong Mout village, Pudang Ridge, around point 19°23’33.7”N 102°08’05.7”E. Remnants of primary broad-leaved evergreen forest on mountain tops composed with shale at elevation 1950-2000 m. Terrestrial erect herb to 1.5 in open grassy place. Flowers light violet-purple. Not common.

LA-VN 707 22 March 2013

Coll.: L.Averyanov, N.S.Khang, S.Lorphengsy

Asteraceae

Vernonia sp.

Hoa Binh Prov., Lac Son Distr., Tu Do Municipality, Mon village, around point 20°24’52”N 105°19’41”E. Secondary evergreen dry broad-leaved forest on steep rocky slopes of remnant mountain composed with highly eroded marble-like solid crystalline limestone at elevation about 300 m a.s.l. Herbaceous vine 5-6 m long in open rocky place. Flowers yellow-orange. Not rare.

CPC 1377 23 March 2011

Coll.: N.Q.Hieu, L.Averyanov, N.T.Hiep, P.K.Loc, P.V.The, N.T.Vinh, T.B.Ngan, N.Tap, D.D.Dao, V.T.Ha.

Asteraceae

Vernonia sp.

Hoa Binh Prov., Lac Son Distr., Tu Do Municipality, Cho village, around point 20°25’21”N 105°18’01”E. Secondary scrub along stream in alluvial valley between limestone hills at elevation about 230 m a.s.l. Herbaceous vine to 5 m long. Flowers yellow. Common.

CPC 1419 24 March 2011

Coll.: N.Q.Hieu, L.Averyanov, N.T.Hiep, P.K.Loc, P.V.The, N.T.Vinh, T.B.Ngan, N.Tap, D.D.Dao, V.T.Ha.

Asteraceae

Vernonia sp.?

Nghe An Prov., Ky Son Distr., Na Ngoi Municipality, eastern slopes of Phu Xai Lai Leng Mountain. Primary and secondary broad-leaved forest on steep mountain slopes composed of sandstone and gray shale at elevation 1300-1500 m a.s.l. around point 19°13’53.4’’N 104°12’09.7’’E.

Shrub 3 m tall in rather open place. Common.

26 October 2013 CPC 6334

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, L.M.Tuan, N.A.Trang, L.H.Dan

Asteraceae

Vernonia sp.?

Nghe An Prov., Ky Son Distr., Na Ngoi Municipality, eastern slopes of Phu Xai Lai Leng Mountain. Primary and secondary broad-leaved forest on steep mountain slopes composed of sandstone and gray shale at elevation 1300-1500 m a.s.l. around point 19°13’53.4’’N 104°12’09.7’’E.

Shrub 3 m tall in rather open place. Common.

26 October 2013 CPC 6334

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, L.M.Tuan, N.A.Trang, L.H.Dan

Asteraceae

Vernonia sp.

Hoa Binh Prov., Lac Son Distr., Ngoc Son Municipality, Khu village, around point 20°26’31”N 105°19’32”E. Primary broadleaved forest on steep rocky slopes of remnant mountain composed with solid crystalline highly eroded limestone at elevation about 550-650 m a.s.l.

Herbaceous vine to 6 m long on rather open rocky slope. Flowers purple-violet. Locally common.

CPC 1607 27 March 2011

Coll.: N.Q.Hieu, L.Averyanov, N.T.Hiep, P.K.Loc, P.V.The, N.T.Vinh, T.B.Ngan, N.Tap, D.D.Dao, V.T.Ha.

Asteraceae

Vernonia sp.

Thanh Hoa Prov., Thuong Xuan Distr., Bat Mot Municipality, Vin village, Xuan Lien Natural Reserve. Primary broad-leaved evergreen wet forest on shale at elevation 1000-1200 m a.s.l. around point 19°58’18.2’’N 104°59’24.0’’E. Semiwoody vine 3-4 m long in open places along stream slope. Rare.

2 November 2013 CPC 6572

L.Averyanov, N.T.Hiep, N.S.Khang

Asteraceae

Blumea sp.?

Son La Prov., Van Ho Distr., Tan Xuan Municipality, Cot Moc village, territory of Xuan Nha natural reserve, eastern slopes of Pha Luong Mountain. Remnants of primary and secondary broad-leaved evergreen humid forest on very steep mountain slopes composed by shale and sandstone at elevation 1000-1400 m a.s.l. around point 20°40’33.3’’N 104°39’00.3’’E. Terrestrial erect herb 1.5-2 m tall on grassy steep slope. Flowers purple-violet. Locally common.

15 November 2013 CPC 7184

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, N.D.Thang, L.D.Qui

Asteraceae

Vernonia sp.

Quang Binh Prov., Minh Hoa Distr., Dan Hoa Municipality, Bai Dinh village around point 17°44’04”N 105°49’14”E. Remnants of primary evergreen broad-leaved forest on very steep slopes and on tops of remnant hills composed with highly eroded marble-like solid limestone at elevation about 400-500 m a.s.l. Semi-woody vine to 15 m long in shrubs on rocky hill top. Flowers light violet. Locally very common.

HAL 12476 8 February 2009

Coll.: L.Averyanov, P.K.Loc, N.T.Vinh, L.T.Son.

Orchidaceae

Bulbophyllum astelidum Aver.

Lam Dong Province, Bidoup Nui Ba National Park, humid primary evergreen broad-leaved and secondary mixed (with Pinus kesiya) forest at elevation of 1200-2100 m a.s.l.

14-26 January 2026, A. Petrov, AL4511 (living plant only).

Flowered in culture on 26.02.2026, L. Averyanov, AL4511.

Convallariaceae

Aspidistra sp.

Son La Province, Chiong Pac District, evergreen limestone forest, wild collected plant from street market.

March 2023, Nguyen Van Canh, L. Averyanov, T. Maisak, AL2418.

Flowered in culture on 04 March 2026, L. Averyanov, T. Maisak, AL2418.

Asteraceae

Adenostemma sp.

Quang Nam Prov., Dong Giang Distr., Ma Cooih Municipality around point 15°58’11”N 107°40’15”E. Remnants of primary closed evergreen broad-leaved forests on steep slopes composed with shale at elevation 450-500 m a.s.l. Terrestrial herb in open wet place along stream. Occasional.

HAL 12098 9 January 2009

Coll.: L.Averyanov, P.K.Loc, P.V.The, N.T.Vinh

Asteraceae

Solidago sp.

Son La Prov., Thuan Chau Distr., Ban Lam Municipality, Ban Lam village, around pick mountain Hua Bay, 21º12’45’’N 103º46’44’’E.

Secondary grasslands and open secondary shrubs on very steep mountain slopes composed with clay shale at elev. 1500-1700 m a.s.l.

Terrestrial herb up to 1 m tall. Flowers yallow. Locally common.

HAL 9679 10 Nov. 2006

Coll.: Nguyen Tien Hiep, L.Averyanov, Pham Van The.

Pinaceae

Pinus cernua P.K.Loc ex Aver., K.S.Nguyen & T.H.Nguyen

Son La Prov., Van Ho Distr., Tan Xuan comm., A Lay village, around point 20°40’46.2N 104°39’49.6E. Primary evergreen broad-leaved forest on very steep rocky slopes and on mountain tops composed with red-brown sandstone at elevation about 1500 m a.s.l. Tree 15-20 m tall and 70-80 cm DBH. Rare. Only three trees were seen.

1 October 2016 CPC 8169 Photos

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.V.Maisak, N.T.Son.

All photos © - L. Averyanov & Nguyen Sinh Khang

Pinaceae
Pinus cernua P.K.Loc ex Aver., K.S.Nguyen & T.H.Nguyen

Son La Prov., Moc Chau Distr., Chieng Son comm., Pha Luong village, Pha Luong Mountain system.
Primary broad-leaved evergreen forest with bamboo on eroded red-brown sandstone near mountain top at elevation 1425 m a.s.l. around point 20°41′30.9N 104°38′08.9E.
Tree. Rare. Only 3 trees were found.
25 September 2016 CPC 8022
Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.V.Maisak, N.T.Son.

All photos © - L. Averyanov & Nguyen Sinh Khang

Pinaceae
Pinus cernua P.K.Loc ex Aver., K.S.Nguyen & T.H.Nguyen

Son La Prov., Moc Chau Distr., Chieng Son comm., Pha Luong village, Pha Luong Mountain system.
Primary mixed evergreen forest with bamboo on eroded red-brown sandstone near mountain top at elevation 1350-1400 m a.s.l. around point 20°41′41.1N 104°37′48.9E (20°41′36.9N 104°37′47.6E; 20°41′37.9N 104°37′51.1E). Tree 10-15 m tall. Locally common. About 20 trees were recorded.
23 September 2016 CPC 8013.
Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.V.Maisak, N.T.Son.

All photos © - L. Averyanov & Nguyen Sinh Khang

Pinaceae

Pinus cernua P.K.Loc ex Aver., K.S.Nguyen & T.H.Nguyen

Son La Prov., Moc Chau Distr., Chieng Son comm., Pha Luong village, Pha Luong Mountains. Primary evergreen broad-leaved forest on mountain tops and steep summit slopes composed with highly eroded red-brown sandstone at elevation 1400-1500 m a.s.l. around point 20°41’33.2N 104°37’37.0E. Tree 15-20 m tall and 40-45 cm in diam., near mountain top. Rare. Two trees were seen.

22 September 2016 CPC 7959

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.V.Maisak, N.T.Son.

All photos © - L. Averyanov & Nguyen Sinh Khang

Pinaceae

Pinus cernua

Son La, Moc Chau, Chieng Xuan P 11177, Phan Ke Loc.

All photos © - Phan Ke Loc

Pinaceae

Picea tianschanica

Kazakhstan, Dzungharia mt., s.n. a. 1978, L. Averyanov.

All photos © - L. Averyanov

Pinaceae

Picea orientalis

Georgia, Bakuriani, 2200-2500 m. Aug. 2010, L. Averyanov.

All photos © - L. Averyanov

Pinaceae

Keteleeria lungcuensis

Ha Gieng, Dong Van, Lung Cu, Phan Ke Loc, 11416.

All photos © - Phan Ke Loc

Pinaceae

Keteleeria evelyniana Mast.

S. Vietnam, Prov. Lam Dong, distr. Lac Duong, municipalite Da Chay, 35 km to NE from Dalat city. Open secondary broadleaved evergreen mt forest along river valley on W macroslope of Bi Dup mt system at 1450 m alt. (N 12º 09', E 108º 41'). Tree about 25-30 (up to 35) m hg. Not common, occcur on W and NW macroslope of Bi Dup mt system mostly at 1450-1600 m alt. No VH 4553 04.05.1997

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 5803-96)

Collectors: L. Averyanov, N.Q. Binh, N.T. Hiep, P.K. Loc, P. Lowry

All photos © - L. Averyanov

Pinaceae

Keteleeria evelyniana Mast.

Xiangkhouang Prov., Kham Distr., Huad village, Phou tat Vinh mt., around point 19°32'31.6'N 103°39'40.4'E. Primary and secondary broad-leaved evergreen and semideciduous dry forest on very steep rocky slopes of limestone highly eroded mountains composed by solid marble like limestone at elev. 1200-1400 m a.s.l. Tree to 30 m tall and 1 m DBH. Occasional co-dominant of the forest along ridge edge. Rarely forms coniferous forests on mountain tops (together with Pinus kesiya).

LA-VN 1083 4 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

All photos © - L. Averyanov

Pinaceae

Keteleeria evelyniana Mast.⁣

Houphan Prov., Xon Distr., Nong Hua Pa Noi village, around point 20°13'02.1''N 103°24'23.3''E. Primary and secondary broad-leaved evergreen forest on very steep rocky mountain slopes composed by shaly limestone at elev. 1100-1200 m a.s.l. Tree 20-25 m tall in open secondary forest on rocky slope. Locally common. Occasional secondary forest co-dominant.

LA-VN 1264a 11 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

All photos © - L. Averyanov