Herbarium LE - Specimens

Plants of Bi Doup – Nui Ba National Park

Pinaceae

Pinus dalatensis

Vietnam, Lam Dong Prov., Lac Duong Distr., Da Chais Municipality, around point 12º08’17’N 108º40’47’’E, at elevation about 1600 m a.s.l. Primary mixed forest with Pinus dalatensis and P. krempfii on W slope of Gia Rich ridge. Tree up to 30 m tall. Common. One of dominant of first forest stratum.

HLF 5318 19 October 2005. Photo.

Coll. T.T.T.Trang, T.V.Thao, N.T.Vinh.

Grant from Henry Luce Foundation.

All photos © - L. Averyanov

Pinaceae

Pinus dalatensis?

S. Vietnam, Kontum, Ngoc Linh Mt., main summit,

1995, L. Averyanov.

All photos © - L. Averyanov

Pinaceae

Pinus dalatensis

S. Vietnam, Kontum Prov., Kon Plong Distr., Hieu Municipality, Mang La forest enterprise (14º39'N, 108º25'E). Primary wet mossy mountain coniferous forest on sandstone (with Pinus dalatensis, Fokienia hodginsii, Dacrydium elatum, Dacrycarpus imbricatus) along ridge tops at elev. 1200 m. Tree up to 25 m hg. Common on slopes and ridge tops. One of main codominants of coniferous forest.

No VH 5354 18.04.2000

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 6383-98)

Collectors: L.Averyanov, P.K.Loc, P.H.Hoang, D.X.Du, N.T.Vinh

All photos © - L. Averyanov

Pinaceae

Pinus dalatensis Ferr?

Saravan Prov., Sa Mouay Distr., to the SEE of Sa Mouay town, elevation from Ta Oy village (700 m elev.) to mountain ridge with highest point at 1350 m around 16°08’47.0”N 106°52’35.4”E. Primary evergreen humid mixed and coniferous forest on slopes composed with silicate rocks. Tree to 40 m tall and to 1.5 m DBH on tops and on ridge slopes from 1150 to the tops of ridge. Large old trees not common, seedlings, saplings and young trees to 0.15 m DBH abundant. Cones 6-8 cm long in old trees and to 18 cm long in younger samples.

LA-VN 195 04 March 2013

Coll.: L.Averyanov, N.T.Hiep, V.Lamxay, N.S.Khang, P.V.The, S.Lorphengsy

All photos © - L. Averyanov & Nguyen Sinh Khang

Podocarpaceae

Dacrydium elatum (Roxb.) Wall. ex Hook.

Saravan Prov., Sa Mouay Distr., to the SEE of Sa Mouay town, elevation from Ta Oy village (700 m elev.) to mountain ridge with highest point at 1350 m around 16°08’47.0”N 106°52’35.4”E. Primary evergreen humid mixed and coniferous forest on slopes composed with silicate rocks. Tree to 12 m tall on tops of ridge. Very common, important co- dominant of coniferous forest.

LA-VN 180a 04 March 2013

Coll.: L.Averyanov, N.T.Hiep, V.Lamxay, N.S.Khang, P.V.The, S.Lorphengsy

All photos © - L. Averyanov & Nguyen Sinh Khang

Triuridaceae

Sciaphila clemensiae Hemsl.

S. Vietnam, Prov. Khanh Hoa, distr. Khanh Son, 42 km to NE from Dalat city. Primary closed broadleaved evergreen forest on E macroslope of Hon Giao mt. ridge at 1500-1600 m alt. (N 12° 12', E 108° 44'). Chlorophyll less pinkish saprophyte on steep shadow slopes. Very rare. No VH 4269 24.04.1997 International Botanical Expedition of the U.S.A. National Geographic Society (grant No 5803-96) Collectors: L. Averyanov, N.Q. Binh, N.T. Hiep

All drawings © - L. Averyanov

Orchidaceae

Vanda fuscoviridis Lindl.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Epiphyte on tall trees along ridge edge above tall cliff. Occasional.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2855.

Flowered in culture on 11 March 2026, L. Averyanov, AL2855.

Orchidaceae

Phaius wallichii Lindl.

Lam Dong Province, the border of Lac Duong and Dam Rong districts, Bidoup – Nui Ba National Park, mountain evergreen broad-leaved and mixed forest (with Pinus kesiya) at elevation 1600–1800 m a.s.l. Terrestrial herb in shady place along rocky mountain stream. Not common. 21 February 2023, L. Averyanov, Nguyen Van Canh, Nong Van Duy, AL 2173. Living plant only.

Flowered in culture on 17 March 2026, L. Averyanov, AL2173.

Asteraceae

Gerbera sp.?

Dien Bien Prov., Tua Chua Distr., Trung Thu Municipality, Trung Thu Village, around point 21°55’38”N 103°18’05”E. Highly degraded primary evergreen broadleaved forest on very steep slopes and along remnant ridge composed with solid limestone at elevation 1350-1450 m a.s.l.

Lithophytic herb on vertical shady cliff along ridge tops. Rare.

CPC 2166 10 April 2011

Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, N.T.Vinh.

Asteraceae

Thanh Hoa Prov., Thuong Xuan Distr., Van Xuan Municipality, Quan village, Xuan Lien Natural Reserve. Secondary broad-leaved evergreen forest on sandstone hills at elevation 300-500 m a.s.l. around point 19°51’27.0’’N 105°12’46.0’’E. Terrestrial herb in open rocky wet place along stream. Flowers white. Common.

7 November 2013 CPC 6818

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, N.D.Thang, L.D.Qui

Asteraceae

Elephanthopus scaber

Nghe An Prov., Ky Son Distr., Na Ngoi Municipality, vicinities of Phu Kha village, eastern slopes of Phu Xai Lai Leng mountain system.

Secondary broad-leaved open forest and secondary scrub on very steep clay laterite hill slopes on shale along road at elevation 700-900 m a.s.l. around point 19°13’21.9’’N 104°15’04.0’’E. Terrestrial herb to 1 m tall in open grasslands along road. Flowers white. Very common.

27 October 2013 CPC 6374

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, L.M.Tuan

Asteraceae

Elephantopus scaber L.?

Cao Bang Prov., Nguyen Binh Distr., Ca Thanh Municipality, Cao Lu village, around point 22°42'47.7"N, 105°51'50.8"E. Primary coniferous forest with Pseudotsuga sinensis along highly eroded rocky limestone ridge at elevation about 1300 m a.s.l. Terrestrial rosulate herb on short tall grassland on alluvial slope along road. Flowers purple-violet. Common.

4 October 2013 CPC 5410

L.Averyanov, N.T.Hiep, L.M.Tuan, N.S.Khang, T.Maisak, L.Osinovets

FLORA OF VIETNAM

Asteraceae

Elephantopus mollis Kunth.

Binh Phuoc Prov.

Bochkin, V. D.

22/133, 17.05.2014

FLORA OF VIETNAM

Asteraceae

Bidens pilosa L.

Tinh Quang Tri Prov.

N. Tap, N. V. Trai

HLF 6023, 20.03.2006

FLORA OF VIETNAM

Asteraceae

Synedrella nodiflora (L.) Gaertn.⁣

Tinh Quang Tri Prov.

N. Tap, N. V. Trai

HLF 6460, 02.04.2006

Asteraceae

Ageratum sp.

Quang Nam Prov., Dong Giang Distr., Ma Cooih Municipality around point 15°49’07”N 107°39’51”E. Secondary evergreen woodlands and scrubs on slopes composed with shale at elevation 400-500 m a.s.l. Open wet grassland along road. Flowers light dull blue-violet. Very common. Commonest weedy plants in the area.

HAL 12070 8 January 2009

Coll.: L.Averyanov, P.K.Loc, P.V.The, N.T.Vinh

FLORA OF VIETNAM

Asteraceae

Ageratum conyzoides L.⁣

Tinh Quang Tri Prov.

N. Tap, N. V. Trai

HLF 6021, 20.03.2006

FLORA OF VIETNAM

Asteraceae

Ageratum conyzoides L.⁣

Tinh Quang Tri Prov.

N. Tap, N. V. Trai

HLF 6031, 20.03.2006

FLORA OF VIETNAM

Asteraceae

Ageratum conyzoides L.⁣

Tinh Quang Tri Prov.

N. Tap, N. V. Trai

HLF 6371, 31.03.2006

FLORA OF VIETNAM

Asteraceae

Dichrocephala integrifolia (L. f.) Kuntze⁣

Tinh Quang Tri Prov.

N. Tap, N. V. Trai

HLF 6374, 31.03.2006

Plants of Ha Giang Province

Compositae

Ha Giang Prov., Dong Van distr., Pai Lung mun., 23?15’15’’ N, 105?22’16’’ E, 1232 m. Scattered in abandoned shifting areas of limestone mountains. Annual herb. Flowers white.

HAL 8671 26 May 06 Photos

Collectors: Phan Ke Loc, Nguyen Sinh Khang & Vu Xuan Manh.

Grant from Program of Basic Researchs in Life Sciences, Vietnam

FLORA OF VIETNAM

Asteraceae

Spilanthes paniculata Wall. ex DC.⁣

Tinh Quang Tri Prov.

N. Tap, N. V. Trai

HLF 6027, 20.03.2006

Asteraceae

Aster sp. s.l.

Lai Chau Prov., Phong Tho Distr., Sin Suoi Ho Municipality, Sa Ma Pho village, around point 22°32’20’’N 103°33’31’’E. Secondary open evergreen broad-leaved forest and secondary woodlands on very steep mountain slopes and cliffs composed predominantly with weathered granite at elev. about 1700-1900 m a.s.l. Terrestrial herb on very steep open rocky grassy slope. Flowers light violet to nearly white. Common.

HAL 10610 12 Dec. 2006 Photo.

Coll.: Nguyen Tien Hiep, L.Averyanov, Phan Ke Loc, Pham Van The, N.T.Vinh.

Asteraceae

Solidago virgaurea L.

Vietnam.

Averyanov L., Kudryavtzeva E.

Averyanov 1327, 1987

Asteraceae

Aster sp. s.l.

Son La Prov., Mai Son Distr., Muong Tranh Municipality, Bong village, around point 21º14’51’’N 103º49’17’’E. Broad-leaved secondary open dry forest on steep slopes composed with clay shale at elev. 900-1200 m a.s.l. Terrestrial herb up to 1 m tall in short tall open secondary grasslands on mountain top. Flowers white. Occasional.

HAL 9577 07 Nov. 2006

Coll.: Nguyen Tien Hiep, L.Averyanov, Pham Van The.

Asteraceae

Thanh Hoa Prov., Thuong Xuan Distr., Van Xuan Municipality, Quan village, Xuan Lien Natural Reserve. Secondary broad-leaved evergreen forest on sandstone hills at elevation 300-500 m a.s.l. around point 19°51’27.0’’N 105°12’46.0’’E. Terrestrial herb in open rocky wet place along stream. Flowers white. Common.

7 November 2013 CPC 6818

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, N.D.Thang, L.D.Qui

Asteraceae

Aster sp. s.l.

Thanh Hoa Prov., Thuong Xuan Distr., Bat Mot Municipality, Vin village, Xuan Lien Natural Reserve. Remnants of primary broad-leaved evergreen humid forest on rocky, highly eroded, crystalline limestone at elevation 750-800 m a.s.l. around point 19°59’11.4’’N 104°58’31.8’’E.

Terrestrial herb to 1 m tall among open secondary shrubs along stream slpe in alluvial valley between limestone hills. Marginal flowers light violet, central flowers yellow. Not rare.

1 November 2013 CPC 6727

L.Averyanov, N.T.Hiep, N.S.Khang

Orchidaceae

Bulbophyllum semiteretifolium Gagnep.

Lam Dong Province, Bidoup Nui Ba National Park, humid primary evergreen broad-leaved and secondary mixed (with Pinus kesiya) forest at elevation of 1200-2100 m a.s.l.

14-26 January 2026, A. Petrov, AL4520 (living plant only).

Flowered in culture on 11 March 2026, L. Averyanov, AL4520.

Asteraceae

Vernonia sp.

Quang Tri Prov., Huong Hoa Distr., Huong Viet municipality around point 16°51’06”N 106°34’38”E. Partially destroyed primary broadleaved evergreen forest on very steep rocky slopes of remnant hills composed with solid highly eroded crystalline limestone at elev. about 600-700 m a.s.l. Undershrub to 2 m tall in grassy places along road. Flowers yellow. Very common.

CPC 2825 7 May 2011

Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, P.V.The, N.T.Vinh.

Asteraceae

Vernonia sp.

Quang Tri Prov., Huong Hoa Distr., Huong Lap municipality, Cup village, around point 16°55’11”N 106°34’36”E. Poor remnants of primary broadleaved evergreen forest on very steep slopes of river valley composed with stratified shale at elev. about 350 m a.s.l. Herbaceous vine 4-6 m long in open place. Occasional.

CPC 2910 8 May 2011

Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, P.V.The, N.T.Vinh.

Asteraceae

Vernonia sp.

Dien Bien Prov., Tua Chua Distr., Sin Chai Municipality, around point 22°03’38”N 103°19’56”E. Primary humid evergreen broad-leaved forest on very steep rocky slopes and on tops of remnant mountain composed with highly eroded marble-like solid crystalline limestone at elevation 1350-1500 m a.s.l. Herbaceous vine to 10 m long in rocky place along road. Not common.

CPC 978 14 December 2010

Coll.: L.Averyanov, P.K.Loc, P.V.The, N.T.Vinh.

Asteraceae

Vernonia sp.

Dien Bien Prov., Tua Chua Distr., Sin Chai Municipality, around point 22°03’38”N 103°19’56”E. Primary humid evergreen broad-leaved forest on very steep rocky slopes and on tops of remnant mountain composed with highly eroded marble-like solid crystalline limestone at elevation 1350-1500 m a.s.l. Herbaceous vine to 4 m long on shady mossy cliff. Flowers white to yellowish. Not common.

CPC 907 13 December 2010

Coll.: L.Averyanov, P.K.Loc, P.V.The, N.T.Vinh.

Asteraceae

Gynura sp.? (or Vernonia sp.?)

Kham Mouan Prov., about 10 km to the E from Thaket City by road 12, around point 17°26’53.3”N 104°52’09.4”E at elev. about 150 m.

Very dry broad-leaved secondary semideciduous open forest and scrub on rocky crystalline limestone at elevation about 200 m.

Lithophytic herb to 0.5 m tall with fleshy leaves on shady vertical cliff. Flowers light violet. Not rare.

LA-VN 334 11 March 2013

Coll.: L.Averyanov, N.T.Hiep, V.Lamxay, N.S.Khang, P.V.The, S.Lorphengsy

Asteraceae

Cirsium sp. s.l.

Son La Prov., Thuan Chau Distr., Ban Lam Municipality, Ban Lam village, around pick mountain Hua Bay, 21º12’45’’N 103º46’44’’E.

Secondary grasslands and open secondary shrubs on very steep mountain slopes composed with clay shale at elev. 1500-1700 m a.s.l.

Terrestrial herb up to 2 m tall. Flowers white to light yellowish. Locally common.

HAL 9678 10 Nov. 2006

Coll.: Nguyen Tien Hiep, L.Averyanov, Pham Van The.

Fam. Asteraceae

Cirsium sp. (or Carduus sp.?)

Vietnam, Ha Giang Prov., Yen Minh Distr, Lao Va Chai Municipality, vicinities of Chi Sang village around point 23º06’20’’N, 105º04’25’’E at elevation about 1450 m a.s.l. Primary mossy coniferous forest with Tsuga sinensis on tops of high rocky ridge composed with highly eroded marble-like limestone. Terrestrial and lithophytic herb up to 2.5 m tall in open places. Flowers whitish. Locally common.

No HAL 8431 9 Dec 2005.

Collectors: L.Averyanov, P.K.Loc, T.V.Thao, P.V.The, N.S.Khang.

Fam. Asteraceae

Cirsium sp. (or Carduus sp.?)

Vietnam, Ha Giang Prov., Yen Minh Distr, Lao Va Chai Municipality, vicinities of Chi Sang village around point 23º06’20’’N, 105º04’25’’E at elevation about 1450 m a.s.l. Primary mossy coniferous forest with Tsuga sinensis on tops of high rocky ridge composed with highly eroded marble-like limestone. Terrestrial and lithophytic herb up to 2.5 m tall in open places. Flowers whitish. Locally common.

No HAL 8431 9 Dec 2005.

Collectors: L.Averyanov, P.K.Loc, T.V.Thao, P.V.The, N.S.Khang.

FLORA OF VIETNAM

Fam. Asteraceae

Hemistepta lyrata Bunge⁣.

Nghe An Prov.

Do Thi Xuyen et al.

HLF. TR7018, 03.2007

Orchidaceae

Bulbophyllum retusiusculum Rchb.f.

Dak Nong Province, Dak Song District, Nam Nung Nature Reserve around point 12.1200N 107.4233E. Fractionally logged primary evergreen broad-leaved and mixed forest with Dacrydium, Dacrycarpus and Pinus latteri on granite at elevation about 900 m a.s.l. Creeping epiphyte on tall tree. Not rare.

27 November 2022, L. Averyanov, Nguyen Van Canh, T. Maisak, Truong Ba Vuong, Nguyen Quoc Bao, AL1835.

Flowered in culture on 11 March 2026, L. Averyanov, AL1835.

Asteraceae

Vientiane Prov., Kasi Distr., Namken village, Phachao Mt., around point 19°18’45.5”N 102°22’31.4”E. Primary broad-leaved evergreen forest on very steep rocky slopes of remnant mountain composed with highly eroded solid crystalline limestone at elevations 1500-1750 m.

Terrestrial and lithophytic erect herb on exposed mossy rocks near top of mountain. Common.

LA-VN 805 24 March 2013

Coll.: L.Averyanov, N.S.Khang, S.Lorphengsy

Asteraceae

Cirsium sp.?

Nghe An Prov., Ky Son Distr., Na Ngoi Municipality, Phu Kha village, eastern slopes of Phu Xai Lai Leng mountain system.

Primary broad-leaved forest on steep eastern granite slopes of Phu Xai Lai Leng Mountain at elevation 1700- 2300 m a.s.l. around point 19°12’54’’N 104°12’01’’E. Terrestrial herb to 1.5 m tall in open grassland along road. Common.

24 October 2013 CPC 6109

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, L.M.Tuan, N.A.Trang, L.H.Dan

Asteraceae

Cirsium sp.?

Nghe An Prov., Ky Son Distr., Na Ngoi Municipality, Phu Kha village, eastern slopes of Phu Xai Lai Leng mountain system.

Primary broad-leaved forest on steep eastern granite slopes of Phu Xai Lai Leng Mountain at elevation 1700- 2300 m a.s.l. around point 19°12’54’’N 104°12’01’’E. Terrestrial herb to 1.5 m tall in open grassland along road. Common.

24 October 2013 CPC 6109

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, L.M.Tuan, N.A.Trang, L.H.Dan

FLORA OF VIETNAM

Asteraceae

Eupatorium odoratum L.⁣

Tinh Quang Tri Prov.

N. Tap, N.V. Trai.

HLF 6026, 20.03.2006

Asteraceae

Cirsium sp.?

Dien Bien Prov., Tua Chua Distr., Ta Phin Municipality, around point 22°00’21”N 103°21’39”E. Open secondary scrub on very steep rocky humid mossy slopes of remnant hills composed with highly eroded marble-like solid crystalline limestone at elevation about 1150 m a.s.l.

Terrestrial herb to 1 m tall among limestone rocks in open place. Flowers light pink-violet. Uncommon weed.

CPC 1020 16 December 2010

Coll.: L.Averyanov, P.K.Loc, P.V.The, N.T.Vinh.

Asteraceae

Vernonia sp.

Dien Bien Prov., Tua Chua Distr., Sin Chai Municipality, around point 22°03’38”N 103°19’56”E. Primary humid evergreen broad-leaved forest on very steep rocky slopes and on tops of remnant mountain composed with highly eroded marble-like solid crystalline limestone at elevation 1350-1500 m a.s.l. Terrestrial scandent herb on steep clay rocky shady slope. Flowers light violet. Occasional.

CPC 898 13 December 2010

Coll.: L.Averyanov, P.K.Loc, P.V.The, N.T.Vinh.

Asteraceae

Vernonia arborea

Dien Bien Prov., Muong Cha Distr., Hua Ngai Municipality, Thien Pa Village, around point 21°52’27”N 103°09’45”E. Highly degraded primary evergreen broadleaved humid forest on very steep slopes of remnant mountains composed with solid limestone at elevation 1100-1300 m a.s.l.

Small tree to 6 m tall on open grassy slope. Flowers light violet. Very common.

CPC 2068 08 April 2011

Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, N.T.Vinh.

Asteraceae

Blumea sp.

Quang Binh Prov., Tuyen Hoa Distr., Lam Hoa municipality, Hung village, around point 17°55’41”N 105°49’43”E. Highly degraded primary and secondary evergreen broadleaved forest on very steep rocky slopes of remnant limestone hills composed with highly eroded marble-like crystalline limestone at elev. 200-300 m a.s.l. Lithophytic herb to 1.5 m tall in humid wet depression between limestone hills. Flowers yellow. Occasional.

CPC 2602 1 May 2011

Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, P.V.The, N.T.Vinh.

Asteraceae

Blumea lacera

Quang Binh Prov., Tuyen Hoa Distr., Lam Hoa municipality, near Ca Tang bridge crossing Gianh River, around point 17°57’31”N 105°48’43”E.

Secondary broadleaved evergreen forest along river valley composed with stratified soft shale limestone at elev. about 60 m a.s.l.

Terrestrial herb to 1 m tall in open wet places. Flowers light pinkish-violet. Very common.

CPC 2647 2 May 2011

Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, P.V.The, N.T.Vinh.

Asteraceae

Vernonia cinerea

Quang Binh Prov., Tuyen Hoa Distr., Lam Hoa municipality, near Ca Tang bridge crossing Gianh River, around point 17°57’31”N 105°48’43”E.

Secondary broadleaved evergreen forest along river valley composed with stratified soft shale limestone at elev. about 60 m a.s.l.

Terrestrial herb on open wet river slopes. Flowers light violet. Very common.

CPC 2661 2 May 2011

Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, P.V.The, N.T.Vinh.

Asteraceae

Vernonia sp.?

Vientiane Prov., Kasi Distr., Thong Mout village, around point 19°23’27.3”N 102°08’21.3”E. Primary broad-leaved evergreen forest on very steep slopes of shaly mountains at elevation about 1800 m. Terrestrial herb to 0.6 m tall in open places along road. Flowers light dull purple-violet. Very common.

LA-VN 652 20 March 2013

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, P.V.The, S.Lorphengsy

Asteraceae

Vernonia sp.

Dien Bien Prov., Tua Chua Distr., Trung Thu Municipality, Trung Thu Village, around point 21°55’38”N 103°18’05”E. Highly degraded primary evergreen broadleaved forest on very steep slopes and along remnant ridge composed with solid limestone at elevation 1350-1450 m a.s.l.

Undershrub to 3 m tall in rather open place along ridge tops. Common.

CPC 2179 10 April 2011

Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, N.T.Vinh.