Herbarium LE - Specimens

Araceae

Thyphonium sp.

Dac Lak Province, Yok Don National Park, terrestrial herb in open dipterocarp forest at elevation of about 200 m a.s.l.

August 2025, Nguyen Van Canh, AL4564.

Araceae

Thyphonium sp.

Dac Lak Province, Yok Don National Park, terrestrial herb in open dipterocarp forest at elevation of about 200 m a.s.l.

August 2025, Nguyen Van Canh, AL4564.

Begoniaceae

Begonia hanhhoae C.W.Lin, V.C.Nguyen, Vuong & Aver.

Khanh Hoa Province, Ninh Hoa District, secondary forest on granite at elevation 100 m a.s.l. Lithophytic herb on open rocks in wet place. Leaves with white nerves. Locally very common. May 2025, Nguyen Van Canh, AL4560 (topotype).

Araceae

Alocasia sp.

Khanh Hoa Province, Ninh Hoa District, secondary forest on granite at elevation 100 m a.s.l. Terrestrial herb among open rocks in wet place. Leaves with white nerves. Locally very common. May 2025, Nguyen Van Canh, AL4559.

Araceae

Alocasia sp.

Khanh Hoa Province, Ninh Hoa District, secondary forest on granite at elevation 100 m a.s.l. Terrestrial herb among open rocks in wet place. Leaves with white nerves. Locally very common. May 2025, Nguyen Van Canh, AL4559.

Convallariaceae

Aspidistra opaca Tillich?

Dac Lak Province, Lak District, evergreen broad-leaved forest on very steep slopes along stream. Locally very common. Flowers dark violet, stigma white.

June 2025, Nguyen Van Canh, AL4557.

Convallariaceae

Aspidistra opaca Tillich?

Dac Lak Province, Lak District, evergreen broad-leaved forest on very steep slopes along stream. Locally very common. Flowers dark violet, stigma white.

June 2025, Nguyen Van Canh, AL4557.

Zingiberaceae

Alpinia sp. (sp. nov.?)

Binh Dinh Province, Van Canh District, submontane forest, at elevation of about 900 m a.s.l. Terrestrial herb to 5 m tall. Locally common. Flowers white with little red.

March 2025, Nguyen Van Canh, AL4556.

Zingiberaceae

Alpinia sp. (sp. nov.?)

Binh Dinh Province, Van Canh District, submontane forest, at elevation of about 900 m a.s.l. Terrestrial herb to 5 m tall. Locally common. Flowers white with little red.

March 2025, Nguyen Van Canh, AL4556.

Acanthaceae

Phlogacanthus sp.

Plant obtained from local collectors in Son La Province in broad-leaved evergreen forest at about 700-800 m a.s.l. Flowers white with tube yellow inside.

Collected in Nguyen Van Canh Garden on 26 March 2026, Nguyen Van Canh, L. Averyanov, Truong Ba Vuong, T. Maisak, AL4553.

Euphorbiaceae

Breynia phuongiana N.K.T.Tram, Vuong, Aver. & V.C.Nguyen

Originally collected in Dak Lak Province, Ea H'leo District, evergreen forest at elevation of about 700 m a.s.l., 15 Feb. 2023, Nguyen Van Canh, AL 2456

Clonotype. Collected in Nguyen Van Canh Garden on 26 March 2026, Nguyen Van Canh, L. Averyanov, Truong Ba Vuong, T. Maisak, AL4552.

Convallariaceae

Aspidistra opaca Tillich

Dac Lak Province, Lac District, Evergreen broad-leaved forest on very steep slopes along stream. Locally very common. Flowers dark violet, stigma white.

Collected in Nguyen Van Canh Garden on 26 March 2026, Nguyen Van Canh, L. Averyanov, Truong Ba Vuong, T. Maisak, AL4550.

Convallariaceae

Peliosanthes “subsessiliflora” sp. new.? (P. micrantha aff.)

Cultivated plant originated from Binh Dinh Province, Van Canh District, lowland forest along stream, at elevation of about 300 m a.s.l., locally common.

Collected in Nguyen Van Canh Garden on 26 March 2026, Nguyen Van Canh, L. Averyanov, Truong Ba Vuong, T. Maisak, AL4548. Flowers dark violet, fruits glossy blue.

Selaginellaceae

Selaginella sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900-1000 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial erect to sub-prostrate herb to 0.5 m tall in open place along trail. Common.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4621.

Rubiaceae

Gaertnera sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900-1000 m a.s.l. around the point 12º10’N 107º46’E. Treelet or shrub to 2 m tall in open place along trail. Flowers white. Not common.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4620.

Rubiaceae

Brachytome walichii Hook.f

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900-1000 m a.s.l. around the point 12º10’N 107º46’E. Treelet or shrub to 2.5 m tall in open place along trail. Flowers pale pink to pink orange. Not rare.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4619.

Apocynaceae

Wrightia dubia (Sims) Spreng.?

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900-1000 m a.s.l. around the point 12º10’N 107º46’E. Treelet or shrub to 3 m tall in open place along trail. Flowers red to red orange. Not rare.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4618.

Pandanaceae

Pandanus sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Wetland thicket in marshland depression with domination of Pandanus and Livistona along stream. Palm-like tree to 4 m tall and leaves to 6 m long. Locally common, co-dominant of plant community.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4610.

Convallariaceae

Tupistra thangii N.Tanaka & D.D.Nguyen?

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial herb with erect stem bearing many stout stilt roots on shady stream slope. Locally common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4609.

Oleaceae

Jasminum sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Scandent shrub of semi-woody vine to 3 m long in open place along road. Flowers white. Common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4606.

Potamogetonaceae

Aponogeton robinsonii A.Camus?

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Aquatic herb with immersed linear leaves in shallow slow water of rocky forest stream. Flowers white. Locally common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4605.

Potamogetonaceae

Aponogeton robinsonii A.Camus?

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Aquatic herb with immersed linear leaves in shallow slow water of rocky forest stream. Flowers white. Locally common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4605.

Acanthaceae

Acanthus sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Creeping ascending herb on shady wet stream slope. Flowers white. Locally very common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4604.

Rubiaceae

Psychotria langbianensis Wernham

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial herb to 1 m tall in rather open place along trail. Flowers white. Not rare.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4603.

Selaginellaceae

Selaginella sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Wetland thicket in marshland depression with domination of Pandanus and Livistona along stream. Terrestrial scandent herb to 1.5 m long in wet open place. Common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4602.

Rubiaceae

Brachytome walichii Hook.f?

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Treelet or shrub to 2 m tall in rather open place along trail. Flowers yellow to yellow orange. Occasional.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4601.

Araceae

Typhonium sp. “simplicifolium”

Dac Lak Province without location. Plant cultivated in Nguyen Van Canh Garden. Spathe reddish brown about 3 cm long, inflorescence very short.

July 2025, Nguyen Van Canh, AL4561.

Orchidaceae

Dendrobium sp.

Gia Lai Province, K'Bang District. Highly degraded secondary evergreen broad-leaved dry forest at elevation 600-800 m a.s.l., around point 14º26N 108º22E. Clustering epiphyte on tall tree. Not rare.

7 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4810.

Passifloraceae

Adina sp.?

Gia Lai Province, K'Bang District. Secondary and remnants of primary evergreen broad-leaved dry forest with Dacrydium on lava rocks (red soils) at elevation about 1000 m a.s.l., around point 14º15N 108º35E. Herbaceous vine 3-4 m long in open place. Fruits bright red. Not common.

6 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4785.

Solanaceae

Solanum sp.

Gia Lai Province, K'Bang District. Highly degraded secondary evergreen broad-leaved dry forest at elevation 600-800 m a.s.l., around point 14º26N 108º22E. Shrub to 2 m tall in open place along road. Flowers white, anthers yellow. Common weed in the area

7 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4764.

Zingiberaceae

Alpinia sp.

Gia Lai Province, K'Bang District. Highly degraded secondary evergreen broad-leaved dry forest at elevation 600-800 m a.s.l., around point 14º26N 108º22E. Terrestrial herb to 3 m tall in open place along road. Flowers white, median tepal red. Common.

7 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4767.

Selaginellaceae

Selaginella sp.

Gia Lai Province, K'Bang District. Secondary and remnants of primary evergreen broad-leaved dry forest with Dacrydium on lava rocks (red soils) at elevation about 1000 m a.s.l., around point 14º15N 108º35E. Prostrate herb on shady slope along trail. Common.

6 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4781.

Podocarpaceae

Dacrydium elatum (Roxb.) Wall.

Gia Lai Province, K'Bang District. Highly degraded secondary evergreen broad-leaved dry forest at elevation 600-800 m a.s.l., around point 14º26N 108º22E. Sapling about 2 m tall of tree to 35 m tall. Former co-dominant of the first forest stratum.

7 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4778.

Selaginellaceae

Selaginella sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Wetland thicket in marshland depression with domination of Pandanus and Livistona along stream. Terrestrial scandent herb to 2 m long in wet open place. Common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4600.

Selaginellaceae

Selaginella sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Wetland thicket in marshland depression with domination of Pandanus and Livistona along stream. Terrestrial scandent herb to 2 m long in wet open place. Common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4600.

Convallariaceae

Aspidistra sp. (“pumila”)

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 1000-1200 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial rhizomatous herb in shady place. Not rare.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4639.

Convallariaceae

Aspidistra paucitepala Vislobokov, Nuraliev & D.D.Sokoloff

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 1000-1200 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial herb with erect stem bearing numerous stilt roots. Fruits dirty olive. Locally abundant.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4638.

Convallariaceae

Aspidistra paucitepala Vislobokov, Nuraliev & D.D.Sokoloff

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 1000-1200 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial herb with erect stem bearing numerous stilt roots. Fruits dirty olive. Locally abundant.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4638.

Zingiberaceae

Etlingera sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 1000-1200 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial herb to 2 m tall in shady place. Flowers bright red. Occasional.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4627.

Zingiberaceae

Etlingera sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 1000-1200 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial herb to 2 m tall in shady place. Flowers bright red. Occasional.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4627.

Podocarpaceae

Podocarpus annamiensis N.E.Gray

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 1000-1200 m a.s.l. around the point 12º10’N 107º46’E. Treelet about 2 m tall on steep shady slope. Tree to 20 m tall. Occasional.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4626.

Pinaceae

Keteleeria evelyniana Mast.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 1000-1200 m a.s.l. around the point 12º10’N 107º46’E. Treelet about 3 m tall on steep shady slope. Tree to 40 m tall and to 3 m DBH. Occasional emergent at elevation of about 1200 m a.s.l.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4624.

Convallariaceae

Peliosanthes macrostegia Hance⁣ var. subcapitata Aver.

Gia Lai Province, K'Bang District. Highly degraded secondary evergreen broad-leaved dry forest at elevation 600-800 m a.s.l., around point 14º26N 108º22E. Terrestrial herb on shady slope. Flowers dirty violet, fruits glossy blue. Locally common.

7 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4768.

Convallariaceae

Peliosanthes macrostegia Hance⁣ var. subcapitata Aver.

Gia Lai Province, K'Bang District. Highly degraded secondary evergreen broad-leaved dry forest at elevation 600-800 m a.s.l., around point 14º26N 108º22E. Terrestrial herb on shady slope. Flowers dirty violet, fruits glossy blue. Locally common.

7 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4768.

Convallariaceae

Peliosanthes macrostegia Hance⁣ var. subcapitata Aver.

Gia Lai Province, K'Bang District. Highly degraded secondary evergreen broad-leaved dry forest at elevation 600-800 m a.s.l., around point 14º26N 108º22E. Terrestrial herb on shady slope. Flowers dirty violet, fruits glossy blue. Locally common.

7 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4768.

Convallariaceae

Tupistra fungilliformis F.T.Wang & S.Yun Liang

Lao Cai Province, Bat Xat district, Y Ty commune

28 November 2025. Truong Ba Vuong, BV 1955.

Epiphytic plant, inflorescence oblique to pendulous, rachis square (4 angles can be see at number IMG_1110), flower sessile, odorless.

Convallariaceae

Tupistra fungilliformis F.T.Wang & S.Yun Liang

Lao Cai Province, Bat Xat district, Y Ty commune

28 November 2025. Truong Ba Vuong, BV 1955.

Epiphytic plant, inflorescence oblique to pendulous, rachis square (4 angles can be see at number IMG_1110), flower sessile, odorless.

Convallariaceae

Aspidistra bicolor Tillich

Cao Bang Province, Phuc Hoa District (Phục Hòa), Hanh Phuc Commune (Hạnh Phúc),

Lung Luong Village (Lũng Luông). Plants in the population with 2 types of inflorescences (?), one is shorter quite close to the stem, the other form with much longer, erect inflorescence. Flower odorless. Leaves sometimes with sparsely yellowish dots.

17 December 2025, Truong Ba Vuong, BV 1956.

Gesneriaceae

Didymocarpus hanhnguyenae V.C.Nguyen, Vuong, Aver. & C.V.Lin⁣.

Kon Tum Province, Tu MA Rong District.

20 September 2025, Nguyen Van Canh, Nguyen Van Khuong, Truong Ba Vuong BV 1990