Herbarium LE - Specimens

Pandanaceae

Pandanus sp.

Gia Lai Province, Mang Yang District, Ayune Commune, Kon Ka Kinh National Park. Secondary and primary dry evergreen broad-leaved forest on shale and lava rocks at elevation of 900-1000 m a.s.l. around point 14º13N 108º20E. Palm-like few branching treelet to 4 m tall in shady wet place. Common.

3 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4713.

Gesneriaceae

Petrocosmea sp.

Lao Cai Prov., Sapa Distr., the territory of Hoang Lien National Park allied to Sapa Town, route to Ham Rong mountain, around point N 22º20’02’’ E 103º50’55”, elevation 1600-1680 m a.s.l. Humid mossy secondary forest and scrub on very steep slopes of mountain composed with solid marble-like highly eroded limestone on the territory of town botanical garden. Lithophytic rosulate herb in crevices of vertical shady mossy limestone cliff. Not common.

26 May 2022, L. Averyanov, Tran Huy Thai, Khang Sinh Nguyen, T. Maisak, VR 1821.

Orchidaceae

Oberonia sp.

Lam Dong Province, Bidoup Nui Ba National Park, humid primary evergreen broad-leaved and secondary mixed (with Pinus kesiya) forest at elevation of 1200-2100 m a.s.l.

14-26 January 2026, A. Petrov, AL4534 (living plant only).

Flowered in culture on 30 July 2026, L. Averyanov, AL4534.

Vitaceae

Tetrastigma sp.?

Lai Chau Province, Muong Te District, Mu Ca Commune. Old rich secondary evergreen broad-leaved forest on stratified shale at elevation of about 540 m a.s.l. around point 22º29’40’’N 102º24’15’’E. Small treelet or woody vine to 1 m long. Not common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5019.

Convallariaceae

Aspidistra sp. (“macrocarpa”)

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 1000-1200 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial rhizomatous herb in shady place. Not rare.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4640.

Convallariaceae

Aspidistra sp. (“macrocarpa”)

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 1000-1200 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial rhizomatous herb in shady place. Not rare.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4640.

Polypodiaceae s.l.

Vittaria sp.

Xiangkhouang Prov., Peak Distr., Oran village, around point 19°38’53”N 103°23’12”E. Primary and secondary broad-leaved evergreen dry forest on shale sandstone slopes and along ridge edge at elev. about 1750-1850 m a.s.l. Epiphyte on mossy tree in shady, humid place. Not common.

LA-VN 901 2 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

Polypodiaceae s.l.

Tectaria sp.

Cao Bang Prov., Bao Lac Distr., Hong An municipality, Mi Lung village. Primary broad-leaved and mixed humid evergreen forest on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation 1400-1450 m a.s.l. around point 22°49’23.4’’N 105°49’54.5’’E. Terrestrial fern in shady humid places among boulders. Fronds to 1.5 m long. Common.

19 November 2014 CPC 7501

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz

Polypodiaceae s.l.

Tectaria sp.?

Houphan Prov., Hem Distr., Khon Ngua village, around point 19°55'53.2''N 103°16'24.2''E.Secondary broad-leaved evergreen open dry forest with bamboo on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. 700-950 m a.s.l.

Lithophytic fern on shady cliff near mountain top. Common.

LA-VN 1490 15 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham, Somchanh Bounphanmy.

Polypodiaceae s.l.

Thelypteris sp.

Houphan Prov., Hem Distr., Nacoc village, Pu He mountain around point 20°10'36.7''N 103°11'39.2''E. Secondary broad-leaved evergreen open dry forest with bamboo on very steep rocky mountain slopes composed by shaly limestone at elev. 700-900 m a.s.l. Terrestrial fern in open wet place along rocky mountain stream. Common.

LA-VN 1370 12 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

Polypodiaceae

Pteris sp.

Ha Giang Prov., Vi Xuyen Distr., Tung Ba municipality, Khau Ca NR. Primary broad-leaved humid evergreen forest on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation 700-1150 m a.s.l. around point 22°50’39.4’’N 105°07’50.3’’E. Terrestrial herb 0.5 m in tall, common.

08 May 2015 CPC 7773

Coll.: N.Q. Hieu, N.T.Hiep, T.B. Ngan, C.Q. Ngan, D.K. Harder, C. P. Tuy, G.C.Pao

VIETNAM

Polipodiaceae

Thelypteris sp.

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 1070, 20.10.1990

VIETNAM

Polipodiaceae

Thelypteris sp.

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 1070, 20.10.1990

Polypodiaceae s.l.

Tectaria sp.

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 816, 15.10.1990

Polypodiaceae s.l.

Tectaria sp.

Quang Ngai Prov.

Thai Tran Bai

412, 07.10.1990

VIETNAM

Polipodiaceae

Thelypteris sp.

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 551, 09.10.1990

Polypodiaceae s.l.

Thelipteris sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12º30'N, 108º30'E) to main peak of

Chu Yang Sinh mt system (12º24'N, 108º26'E) along Dak Tour river (N slope). Primary mossy cloud mixed forest (with Pinus, Fokienia,

Dacrydium and Dacrycarpus) on very steep granite slopes at elev. 2200-2400 m. Terrestrial fern on steep mossy cliff. Common.

No VH 6207 09.05.2000

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 6383-98)

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

Polypodiaceae

Vittaria sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12º30'N, 108º30'E) along Dak Tour river on N macroslope of Chu Yang Sinh mt system to main peak (12º24'N, 108º26'E). Primary broad-leaved mountain evergreen closed forest on granites along river slopes at elev. 1000-1200 m. Epiphyte with pendent fronds up to 1.2 m lg in shady river canyon. Not rare.

No VH 6025 05-06.05.2000

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 6383-98)

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

Polypodiaceae s.l.

Vittaria sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12º30'N, 108º30'E) to main peak of

Chu Yang Sinh mt system (12º24'N, 108º26'E) along Dak Tour river (N slope). Primary wet closed mountain broad-leaved and mixed forest on

granites along very steep shady slopes of river canyon at elev. 1700 m. Epiphyte on vertical mossy shady cliffs along river. Not common.

No VH 6349 11.05.2000

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 6383-98)

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

Polypodiaceae s.l.

Vittaria sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12º30'N, 108º30'E) to main peak of

Chu Yang Sinh mt system (12º24'N, 108º26'E) along Dak Tour river (N slope). Primary wet closed mixed forest (with Pinus, Fokienia,

Dacrydium and Dacrycarpus) along ridge at elev. 1600 m. Epiphyte in shady place. Not rare.

No VH 6185 09.05.2000

International Botanical Expedition of the U.S.A. National Geographic Society

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

Polypodiaceae s.l.

Tectaria sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12º30'N, 108º30'E) to main peak of

Chu Yang Sinh mt system (12º24'N, 108º26'E) along Dak Tour river (N slope). Primary closed mountain broad-leaved evergreen forest on very

steep granite slopes along river canyon at elev. 1350-1400 m. Terrestrial fern on very steep streamlet slope. Very common.

No VH 6386 12.05.2000

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 6383-98)

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

Hymenophyllaceae

Trichomanes sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12º30'N, 108º30'E) to main peak of

Chu Yang Sinh mt system (12º24'N, 108º26'E) along Dak Tour river (N slope). Primary wet closed mixed forest (with Pinus, Fokienia,

Dacrydium and Dacrycarpus) along ridge at elev. 2000-2100 m. Epiphytic fern at the base of large trees on mossy cliff. Not rare.

No VH 6191 09.05.2000

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 6383-98)

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

DRYOPTERIDACEAE

Tectaria

Ha Giang

Quan Ba District; Bat Dai Son

Municipality; Bat Dai Son Provincial

Protected Area (2000); collected on

ridge just behind Bat Dai Son School

building just S of Bat Dai Son border

station; from lower disturbed

agricultural area to limestone ridge in

disturbed ridgetop forest.

23°09'10"N 105°00'11"E 870 - 867 m

23°09'01"N 105°00'10"E

Occasional, near disturbed agricultural

12 February 2001

D.K. Harder, P.K. Loc & G.E. Schatz 6204

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

DRYOPTERIDACEAE

Tectaria

Ha Giang

Vi Xuyen District; Phong Quang

Municipality; Man Village. Limestone

peaks and ridges NE of Village.

22°53'21"N 104°56'45"E 595 - 787 m

Occasional, among rocks, in understory.

Rachis purple-black, shiny; sporangia

dark brown.

15 February 2001

D.K. Harder, P.K. Loc, L.V. Averyanov,

G.E. Schatz & S. Bodine 6381

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

DRYOPTERIDACEAE

Tectaria

Lao Cai

Van Ban District; Liem Phu Municipality;

Khuoi Ngoa Village.

21°58'20"N 104°20'16"E 527 - 1120 m

21°58'08"N 104°21'00"E

Frequent, at only restricted elevation

ca. 750 m. Rachis dark brown to black,

shiny; sporangia dark brown.

21 February 2001

D.K. Harder, L.V. Averyanov & P.K. Loc

6614

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

HYMENOPHYLLACEAE

Trichomanes

Lao Cai

Van Van District; Liem Phu Municipality;

Khuoi Ngoa Village. Collections near

logging camp [Fokienia (Pou Nhou)] on

road through village; collections from

logging camp and along stream.

21°58'14"N 104°19'51"E 606 m

Occasional, lithophytic creeping. Pinnae

clear green; rachis tinged black

medially; sporangia green.

21 February 2001

D.K. Harder, L.V. Averyanov, P.K. Loc &

G.E. Schatz 6557

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

DRYOPTERIDACEAE

Tectaria

Ha Giang

Vi Xuyen District; Phong Quang

Municipality; Man Village. Collection

on ridge and small limestone peak just

east of house of village head; very

disturbed, secondary vegetation on lower

slopes and on limestone rocks up to

ridgetop.

22°53'11"N 104°56'43"E 437 - 645 m

22°53'23"N 104°56'48"E

Occasional on lower slopes. Rachis

14 February 2001

D.K. Harder, P.K. Loc, G.E. Schatz &

L.V. Averyanov 6280

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

Hymenophyllaceae

Trichomanes sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12º30'N, 108º30'E) to main peak of

Chu Yang Sinh mt system (12º24'N, 108º26'E) along Dak Tour river (N slope). Primary mossy cloud mixed forest (with Pinus, Fokienia,

Dacrydium and Dacrycarpus) on very steep granite slopes at elev. 2200-2400 m. Epiphytic and lithophytic fern on shady cliffs. Common.

No VH 6213 09.05.2000

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 6383-98)

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

Pteridophyta J.Y. Bergen & B.M. Davis

Lao Cai

Van Ban District; Liem Phu Municipality;

Khuoi Ngoa Village.

21°58'20"N 104°20'16"E 527 - 1120 m

21°58'08"N 104°21'00"E

Occasional, in shade. Sporangia light

brown.

21 February 2001

D.K. Harder, L.V. Averyanov & P.K. Loc

6602

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

HYMENOPHYLLACEAE

Trichomanes

Lao Cai

Van Van District; Liem Phu Municipality;

Khuoi Ngoa Village. Collections near

logging camp [Fokienia (Pou Nhou)] on

road through village; collections from

logging camp and along stream.

21°58'14"N 104°19'51"E 606 m

Occasional, lithophytic creeping vine;

rachis black-purple; pinnae clear green;

sporangia terminal, green.

21 February 2001

D.K. Harder, L.V. Averyanov, P.K. Loc &

G.E. Schatz 6556

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

VITTARIACEAE

Vittaria

Lao Cai

Van Ban District; Liem Pho Municipality;

Khuoi Ngoa Village. Collections in

disturbed secondary vegetation and

cultivated tree plantation, along road

between tree [Fokienia (Pou Nhou)]

extraction camp along road to near Khuoi

Ngoa school.

21°58'14"N 104°19'51"E 606 m

Occasional, pendant, epiphytic, on

20 February 2001

D.K. Harder, P.K. Loc, L.V. Averyanov,

G.E. Schatz & S. Bodine 6532

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

FLORA OF VIETNAM

Polypodiaceae

Thelipteris sp. ?

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12º30'N, 108º30'E) along Dak Tour

river on N macroslope of Chu Yang Sinh mt system. Secondary dry evergreen broad-leaved forest with remnants of primary elements along

river slopes at elev. 450-550 m. Terrestrial fern with fronds up to 1.2 m lg in open wet place. Very common.

No VH 5892 02.05.2000

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 6383-98)

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

VITTARIACEAE

Vittaria

Lao Cai

Van Ban District; Nam Xe Municipality;

Nam Xi Tan Village. Collection in forest

on steep W-facing slope along stream;

just above village; on N side of road

between Nam Xe and Khau Co Pass at near

40 km to Than Uyen mark.

22°02'31"N 103°58'32"E 700 - 1113 m

Occasional, epiphyte on tree trunk.

Fronds light green; marginal sporangia

26 February 2001

D.K. Harder, P.K. Loc, N.T. Hiep, G.E.

Schatz, S. Bodine & N.Q. Hieu 6918

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

Polypodiaceae s.l.

Thelipteris sp. ?

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12º30'N, 108º30'E) to main peak of

Chu Yang Sinh mt system (12º24'N, 108º26'E) along Dak Tour river (N slope). Primary closed wet broad-leaved and mixed forest (with Pinus,

Fokienia, Dacrydium and Dacrycarpus) on steep granite slopes at elev. 1200-1300 m. Terrestrial fern on open wet very steep slope of

waterfall. Not rare.

No VH 6236 08.05.2000

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 6383-98)

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

DRYOPTERIDACEAE

Tectaria

Son La

Moc Chau District; Van Ho Municipality,

Hua Tat Village. Collections on

limestone ridge from degraded forests to

cultivation on lower slopes to primary

broadleaved, closed wet forest along

limestone ridge and slopes near tops.

20°46'13"N 104°46'12"E 1100 - 1300 m

Lithophytic fern on vertical shady

cliffs. Not rare.

7 March 2001

D.K. Harder, P.K. Loc, L.V. Averyanov &

N.X. Tam 7438

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

VITTARIACEAE

Vittaria

Son La

Moc Chau District; Long Luong

Municipality; Sang Cai Village.

Collections in primary, wet,

broad-leaved, evergreen and

semi-deciduous, mossy forest along

limestone ridge situated between

1400-1500 m elevation.

20°45'51"N 104°51'26"E 1300 - 1500 m

Epiphyte fern. Locally very common.

8 March 2001

D.K. Harder, P.K. Loc, L.V. Averyanov &

N.X. Tam 7496

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

Asteraceae

Gynura auriculata

Phu Tho Prov., Tan Son Distr., Xuan Son Municipality, Du village, Nhon Mountain, around mountain top, point 21°06’51”N 104°57’34”E.

Remnants of primary evergreen primary forest on very steep slopes and cliffs composed with solid highly eroded marble-like limestone at elevation about 600 m a.s.l. Herbaceous vine 2-3 m long in open exposed mossy rocks. Flowers orange. Leaves with violet tint. Not common.

HAL 12817 19 February 2009 Photo.

Coll.: L.Averyanov, P.K.Loc, N.T.Vinh, L.T.Son.

Asteraceae

Hypoxis aurea Lour.⁣

Son La Prov.

Avrorin N., Nguyen Anh Tiep.

Avrorin 12151a , 28.04.1963

Det. Averyanov L. 18.08.2026

Orchidaceae

Dendrobium truncatum Lindl.

Binh Dinh Province, Van Canh District, Canh Lien Municipality. Fractionally logged dense dry primary evergreen broad-leaved forest on steep mountain slopes composed by granite with quartzite at elevation 600-700 m a.s.l. Epiphyte on tall trees.

05 November 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3557.

Fl. In culture on 18 August 2026, L. Averyanov, AL3557.

Л. Ридель: Растения Бразилии 1821-1824 гг.

novbic. 1922

L. Riedel: Iter Brasisiense 1821-1824

0,29.

Pyrenacea Duranta ellisia ?

frytex - spinosus

flores coerulei

10

22

Л. Ридель: Растения Бразилии 1821-1824 гг.

L. Riedel: Iter Brasiliense 1821-1824.