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LE 01318771

LE 01318771

Davallia griffithiana Hook.⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: Tien Hiep Nguyen; Averyanov, Leonid Vladimirovich; et al. Field collecting number: LA-VN1001. Collection date: 03.04.2015.
Administrative regions: LA - Xiangkhouang.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Laos
Original label text:
FLORA OF LAOS
Polypodiaceae s.l.
Davallia sp.

Xiangkhouang Prov., Peak Distr., Khoang village, Pa Khao mt., around point 19°33'02.2'N 103°39'56.3'E. Primary and secondary broad-leaved evergreen dry forest on solid rocky highly eroded limestone at elev. 1200-1450 m a.s.l. Creeping epiphyte on tall mossy tree. Common.
LA-VN 1001 3 April 2015
Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318771 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318771
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LE 01318765

LE 01318765

Microlepia sp.⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: Tien Hiep Nguyen; Averyanov, Leonid Vladimirovich; et al. Field collecting number: LA-VN873. Collection date: 02.04.2015.
Administrative regions: LA - Xiangkhouang.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Laos
Original label text:
FLORA OF LAOS
Polypodiaceae s.l.
Microlepia sp.

Xiangkhouang Prov., Peak Distr., Oran village, around point 19°38’53”N 103°23’12”E. Primary and secondary broad-leaved evergreen dry forest on shale sandstone slopes and along ridge edge at elev. about 1750-1850 m a.s.l. Terrestrial fern in shady places. Common.
LA-VN 873 2 April 2015
Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318765 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318765
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LE 01318764

LE 01318764

Hypolepis sp.⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: Tien Hiep Nguyen; Averyanov, Leonid Vladimirovich; et al. Field collecting number: LA-VN892. Collection date: 02.04.2015.
Administrative regions: LA - Xiangkhouang.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Laos
Original label text:
FLORA OF LAOS
Polypodiaceae s.l.
Microlepia sp.?

Xiangkhouang Prov., Peak Distr., Oran village, around point 19°38’53”N 103°23’12”E. Primary and secondary broad-leaved evergreen dry forest on shale sandstone slopes and along ridge edge at elev. about 1750-1850 m a.s.l. Terrestrial fern in shady wet place. Common.
LA-VN 892 2 April 2015
Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318764 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318764
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LE 01318749

LE 01318749

Lepisorus carnosus (Hook.) C.F.Zhao, R.Wei & X.C.Zhang⁣ det. Cheng-Wei Chen at 21.02.2026

Additional identifications: Lemmaphyllum carnosum (Hook.) C.Presl.⁣ det. Averyanov, Leonid Vladimirovich at 04.02.2026
Collectors: Averyanov, Leonid Vladimirovich; Tien Hiep Nguyen; et al. Field collecting number: CPC7850. Collection date: 21.11.2014.
Administrative regions: VN - Tinh Cao Bang.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
FLORA OF VIETNAM
Polypodiaceae s.l.
Lematosorus sp.??

Cao Bang Prov., Bao Lac Distr., Hong An municipality, Mi Lung village. Primary broad-leaved and mixed humid evergreen forest (with Podocarpus, Pinus, Fokienia and Tsuga) on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation about 1500-1550 m a.s.l. around point 22°49’15.4’’N 105°49’53’’E. Creeping epiphyte on old mossy tree on rocky mossy mountain slope near mountain top. Occasional.
21 November 2014 CPC 7580
Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318749 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318749
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LE 01318736

LE 01318736

Lepisorus sp.⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: Averyanov, Leonid Vladimirovich; Tien Hiep Nguyen; et al. Field collecting number: CPC7610. Collection date: 24.11.2014.
Administrative regions: VN - Tinh Cao Bang.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
FLORA OF VIETNAM
Polypodiaceae s.l.
Lepisorus sp.

Cao Bang Prov., Bao Lac Distr., Dinh Phung municipality, Ban Meo village. Secondary broad-leaved evergreen forest on steep stream slope composed by shale stratified limestone at elevation 800-900 m a.s.l. around point 22°44’32.0’’N 105°46’29.1’’E. Lithophytic fern on large mossy stream boulders. Not common.
24 November 2014 CPC 7610
Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318736 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318736
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LE 01318710

LE 01318710

Diplazium megaphyllum (Baker) Christ⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: Nguyen Quang Hieu; Tien Hiep Nguyen; et al. Field collecting number: CPC7772. Collection date: 08.05.2015.
Administrative regions: VN - Tinh Ha Giang.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
FLORA OF VIETNAM
Polypodiaceae
Gen. sp.

Ha Giang Prov., Vi Xuyen Distr., Tung Ba municipality, Khau Ca NR. Primary broad-leaved humid evergreen forest on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation 700-1150 m a.s.l. around point 22°50’39.4’’N 105°07’50.3’’E. Terrestrial herb 0.5-1-1,5 m in tall, common.
08 May 2015 CPC 7772
Coll.: N.Q. Hieu, N.T.Hiep, T.B. Ngan, C.Q. Ngan, D.K. Harder, C. P. Tuy, G.C.Pao
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318710 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318710
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LE 01318696

LE 01318696

Lepisorus sp.⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: Tien Hiep Nguyen; Averyanov, Leonid Vladimirovich; et al. Field collecting number: LA-VN1094. Collection date: 04.04.2015.
Administrative regions: LA - Xiangkhouang.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Laos
Original label text:
FLORA OF LAOS
Polypodiaceae s.l.
Lepisorus sp.

Xiangkhouang Prov., Kham Distr., Huad village, Phou tat Vinh mt., around point 19°32'31.6'N 103°39'40.4'E. Primary and secondary broad-leaved evergreen and semideciduous dry forest on very steep rocky slopes of limestone highly eroded mountains composed by solid marble like limestone at elev. 1200-1400 m a.s.l. Creeping epiphyte on tall old mossy tree along ridge edge. Occasional.
LA-VN 1094 4 April 2015
Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318696 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318696
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LE 01318693

LE 01318693

Lepisorus superficialis (Blume) C.F.Zhao, R.Wei & X.C.Zhang⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: Tien Hiep Nguyen; Averyanov, Leonid Vladimirovich; et al. Field collecting number: LA-VN888. Collection date: 02.04.2015.
Administrative regions: LA - Xiangkhouang.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Laos
Original label text:
FLORA OF LAOS
Polypodiaceae s.l.
Lepisorus sp.?

Xiangkhouang Prov., Peak Distr., Oran village, around point 19°38’53”N 103°23’12”E. Primary and secondary broad-leaved evergreen dry forest on shale sandstone slopes and along ridge edge at elev. about 1750-1850 m a.s.l. Creeping epiphytic vine 3-5 m long in shady place. Common.
LA-VN 888 2 April 2015
Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318693 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318693
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LE 01318690

LE 01318690

Davallia griffithiana Hook.⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: Tien Hiep Nguyen; Averyanov, Leonid Vladimirovich; et al. Field collecting number: LA-VN1230. Collection date: 11.04.2015.
Administrative regions: LA - Houaphan.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Laos
Original label text:
FLORA OF LAOS
Polypodiaceae s.l.
Davallia sp.

Houphan Prov., Xon Distr., Nong Hua Pa Noi village, around point 20°13'02.1''N 103°24'23.3''E. Primary and secondary broad-leaved evergreen forest on very steep rocky mountain slopes composed by shaly limestone at elev. 1100-1200 m a.s.l. Epiphyte on tall trees. Flowers buds white-yellowish. Locally very common.
LA-VN 1230 11 April 2015
Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318690 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318690
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LE 01318668

LE 01318668

Pteridrys sp.⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: Tien Hiep Nguyen; Averyanov, Leonid Vladimirovich; et al. Field collecting number: LA-VN1341. Collection date: 11.04.2015.
Administrative regions: LA - Houaphan.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Laos
Original label text:
FLORA OF LAOS
Polypodiaceae s.l.
Dryopteris sp.?

Houphan Prov., Hem Distr., Na Puok village, Pu Tham Nhon mountain around point 20°10'00.0''N 103°24'37.0''E. Primary and secondary broad-leaved evergreen dry forest on very steep rocky mountain slopes composed by solid marble-like highly eroded limestone at elev. 1050-1150 m a.s.l. Terrestrial fern in shady slope near mountain base. Very common.
LA-VN 1341 11 April 2015
Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318668 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318668
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LE 01318633

LE 01318633

Drynaria coronans (Wall. ex Mett.) J.Sm.⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: Cuong, Huu Nguyen Field collecting number: CLL0134. Collection date: 2019.
Administrative regions: VN - Vietnam.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318633 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318633
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LE 01318629

LE 01318629

Diplazium sp.⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH7441. Collection date: 07.03.2001.
Administrative regions: VN - Tinh Son La.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
DRYOPTERIDACEAE
Diplazium

Son La
Moc Chau District; Van Ho Municipality,
Hua Tat Village. Collections on
limestone ridge from degraded forests to
cultivation on lower slopes to primary
broadleaved, closed wet forest along
limestone ridge and slopes near tops.
20°46'13"N 104°46'12"E 1100 - 1300 m
Lithophytic fern on vertical shady
cliffs. Not rare.
7 March 2001
D.K. Harder, P.K. Loc, L.V. Averyanov &
N.X. Tam 7441
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318629 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318629
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LE 01318596

LE 01318596

Drynaria quercifolia (L.) J. Sm.⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: Averyanov, Leonid Vladimirovich; Tien Hiep Nguyen; et al. Field collecting number: VH5963a. Collection date: 03.05.2000.
Administrative regions: VN - Tinh GJak Lak.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
FLORA OF VIETNAM
Polypodiaceae s.l.
Drynaria sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) along Dak Tour
river on N macroslope of Chu Yang Sinh mt system. Primary broad-leaved mountain evergreen forest on granites along river slopes at elev.
500-550 m. Lithophytic and epiphytic fern with fronds up to 1.5 m lg in open places along river. Very common.
No VH 5963 03.05.2000
International Botanical Expedition of the U.S.A. National Geographic
Society (grant No 6383-98)
___________________________________________________________
Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318596 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318596
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LE 01318595

LE 01318595

Drynaria quercifolia (L.) J. Sm.⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: Averyanov, Leonid Vladimirovich; Tien Hiep Nguyen; et al. Field collecting number: VH5963b. Collection date: 03.05.2000.
Administrative regions: VN - Tinh GJak Lak.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
FLORA OF VIETNAM
Polypodiaceae s.l.
Drynaria sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) along Dak Tour
river on N macroslope of Chu Yang Sinh mt system. Primary broad-leaved mountain evergreen forest on granites along river slopes at elev.
500-550 m. Lithophytic and epiphytic fern with fronds up to 1.5 m lg in open places along river. Very common.
No VH 5963 03.05.2000
International Botanical Expedition of the U.S.A. National Geographic
Society (grant No 6383-98)
___________________________________________________________
Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318595 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318595
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LE 01318594

LE 01318594

Drynaria quercifolia (L.) J. Sm.⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: Averyanov, Leonid Vladimirovich; Tien Hiep Nguyen; et al. Field collecting number: VH5963c. Collection date: 03.05.2000.
Administrative regions: VN - Tinh GJak Lak.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
FLORA OF VIETNAM
Polypodiaceae s.l.
Drynaria sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) along Dak Tour
river on N macroslope of Chu Yang Sinh mt system. Primary broad-leaved mountain evergreen forest on granites along river slopes at elev.
500-550 m. Lithophytic and epiphytic fern with fronds up to 1.5 m lg in open places along river. Very common.
No VH 5963 03.05.2000
International Botanical Expedition of the U.S.A. National Geographic
Society (grant No 6383-98)
___________________________________________________________
Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318594 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318594
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LE 01318577

LE 01318577

Drynaria coronans (Wall. ex Mett.) J.Sm.⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: Averyanov, Leonid Vladimirovich; et al. Field collecting number: VH6437. Collection date: 12.05.2000.
Administrative regions: VN - Tinh GJak Lak.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
FLORA OF VIETNAM
Polypodiaceae s.l.
Drynaria sp. ?

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of
Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary closed mountain broad-leaved evergreen forest on very steep granite slopes along river canyon at elev. 1100 m. Large epiphytic and Lithophytic rosette fern. Very common.
No VH 6437 12.05.2000
International Botanical Expedition of the U.S.A. National Geographic
Society (grant No 6383-98)
___________________________________________________________
Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318577 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318577
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LE 01318558

LE 01318558

Diplazium petri Tardieu.⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6364. Collection date: 15.02.2001.
Administrative regions: VN - Tinh Ha Giang.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
Pteridophyta J.Y. Bergen & B.M. Davis

Ha Giang
Vi Xuyen District; Phong Quang
Municipality; Man Village. Limestone
peaks and ridges NE of Village.
22°53'21"N 104°56'45"E 595 - 787 m
Occasional, in understory among rocks.
Lower rachis dark brown-black; sporangia
brown.
15 February 2001
D.K. Harder, P.K. Loc, L.V. Averyanov,
G.E. Schatz & S. Bodine 6364
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318558 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318558
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LE 01318551

LE 01318551

Lepisorus sp.⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6303. Collection date: 14.02.2001.
Administrative regions: VN - Tinh Ha Giang.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
POLYPODIACEAE
Lepisorus

Ha Giang
Vi Xuyen District; Phong Quang
Municipality; Man Village. Collection
on ridge and small limestone peak just
east of house of village head; very
disturbed, secondary vegetation on lower
slopes and on limestone rocks up to
ridgetop.
22°53'11"N 104°56'43"E 437 - 645 m
22°53'23"N 104°56'48"E
Occasional, in shade, understory.
14 February 2001
D.K. Harder, P.K. Loc, G.E. Schatz &
L.V. Averyanov 6303
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
----------------------------------------
Page 2
Sporangia yellow-brown, raised.
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318551 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318551
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LE 01318547

LE 01318547

Diplazium dilatatum Blume⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6871. Collection date: 26.02.2001.
Administrative regions: VN - Tinh Lao Cai.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
DRYOPTERIDACEAE
Diplazium

Lao Cai
Van Ban District; Nam Xe Municipality;
Nam Xi Tan Village. Collection in forest
on steep W-facing slope along stream;
just above village; on N side of road
between Nam Xe and Khau Co Pass at near
40 km to Than Uyen mark.
22°02'31"N 103°58'32"E 700 - 1113 m
Occasional, in shade on slope. Fronds to
2 m; rachis green; sporangia brown.
26 February 2001
D.K. Harder, P.K. Loc, N.T. Hiep, G.E.
Schatz, S. Bodine & N.Q. Hieu 6871
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318547 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318547
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LE 01318545

LE 01318545

Lepisorus sp.⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6654. Collection date: 22.02.2001.
Administrative regions: VN - Tinh Lao Cai.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
POLYPODIACEAE
Lepisorus

Lao Cai

Van Ban District; Liem Phu Municipality;
Khuoi Ngoa Village. Collections on
ridges just west of village.
21°59'31"N 104°19'05"E 323 - 1050 m
Epiphytic. Fronds light green with
darker purple midvien; sporangia light
yellow-brown.
22 February 2001
D.K. Harder, P.K. Loc, L.V. Averyanov,
G.E. Schatz & S. Bodine 6654
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318545 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318545
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LE 01318533

LE 01318533

Austroblechnum melanocaulon (Brack.) Gasper & V.A.O.Dittrich⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: Averyanov, Leonid Vladimirovich; et al. Field collecting number: VH6039. Collection date: 11.05.2000.
Administrative regions: VN - Tinh GJak Lak.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
FLORA OF VIETNAM
Polypodiaceae s.l.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of
Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary wet closed mountain broad-leaved and mixed forest on
granites along very steep shady slopes of river canyon at elev. 1500-1700 m. Woody fern up to 1 m hg. Not common.
No VH 6039 11.05.2000
International Botanical Expedition of the U.S.A. National Geographic
Society (grant No 6383-98)
___________________________________________________________
Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318533 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318533
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LE 01318529

LE 01318529

Diplazium petri Tardieu.⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6622. Collection date: 21.02.2001.
Administrative regions: VN - Tinh Lao Cai.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
ASPLENIACEAE
Asplenium

Lao Cai
Van Ban District; Liem Phu Municipality;
Khuoi Ngoa Village.
21°58'20"N 104°20'16"E 527 - 1120 m
21°58'08"N 104°21'00"E
Occasional, in shady place. Lower rachis
with dark brown hairs; rachis shiny
green; sporangia brown.
21 February 2001
D.K. Harder, L.V. Averyanov & P.K. Loc
6622
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318529 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318529
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LE 01318508

LE 01318508

Lepisorus superficialis (Blume) C.F.Zhao, R.Wei & X.C.Zhang⁣ det. Cheng-Wei Chen at 21.02.2026

Additional identifications: Lepisorus superficialis (Blume) C.F.Zhao, R.Wei & X.C.Zhang⁣ det. Averyanov, Leonid Vladimirovich at 04.02.2026
Collectors: D.K. Harder; et al. Field collecting number: DKH6585. Collection date: 21.02.2001.
Administrative regions: VN - Tinh Lao Cai.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
POLYPODIACEAE
Microsorum

Lao Cai
Van Ban District; Liem Phu Municipality;
Khuoi Ngoa Village.
21°58'20"N 104°20'16"E 527 - 1120 m
21°58'08"N 104°21'00"E
Occasional, epiphytic. Rhizome green
with dark brown hairs; rachis stiff
green; sporangia light yellow-brown.
21 February 2001
D.K. Harder, L.V. Averyanov & P.K. Loc
6585
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318508 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318508
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LE 01318506

LE 01318506

Microlepia speluncae (L.) T.Moore.⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6529. Collection date: 20.02.2001.
Administrative regions: VN - Tinh Lao Cai.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
DENNSTAEDTIACEAE
Microlepia

Lao Cai
Van Ban District; Liem Pho Municipality;
Khuoi Ngoa Village. Collections in
disturbed secondary vegetation and
cultivated tree plantation, along road
between tree [Fokienia (Pou Nhou)]
extraction camp along road to near Khuoi
Ngoa school.
21°58'14"N 104°19'51"E 606 m
Occasional in understory. Fronds to 1.5
20 February 2001
D.K. Harder, P.K. Loc, L.V. Averyanov,
G.E. Schatz & S. Bodine 6529
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
----------------------------------------
Page 2
m; young sporangial indusia light green;
rachis green with red-brown markings.
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318506 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318506
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LE 01318490

LE 01318490

Diplazium stenochlamys C.Chr.⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6805. Collection date: 24.02.2001.
Administrative regions: VN - Tinh Lao Cai.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
DRYOPTERIDACEAE
Diplazium

Lao Cai
Van Ban District; Nam Xe Municipality.
Collection in small forest patch near
road between Nam Xe Village and Khau Co
Pass.
22°02'26"N 103°59'21"E 612 m
Occasional, among rocks, in wet area.
Rachis brown-purple; sporangia light
brown; stem to 10 cm; pinnae light
green; roots at tips.
24 February 2001
D.K. Harder & et al. 6805
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318490 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318490
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LE 01318484

LE 01318484

Lepisorus fortuni (T.Moore) C.M.Kuo⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH7443. Collection date: 07.03.2001.
Administrative regions: VN - Tinh Son La.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
POLYPODIACEAE
Lepisorus

Son La
Moc Chau District; Van Ho Municipality,
Hua Tat Village. Collections on
limestone ridge from degraded forests to
cultivation on lower slopes to primary
broadleaved, closed wet forest along
limestone ridge and slopes near tops.
20°46'13"N 104°46'12"E 1100 - 1300 m
Lithophytic creeping fern on wet rocks.
Common.
7 March 2001
D.K. Harder, P.K. Loc, L.V. Averyanov &
N.X. Tam 7443
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318484 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318484
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LE 01318476

LE 01318476

Microlepia platyphylla (Don) J.Sm.⁣ det. Cheng-Wei Chen at 21.02.2026

Additional identifications: Microlepia marginata (Houtt.) C.Chr.⁣ ⟮prelim. ID⟯ det. Averyanov, Leonid Vladimirovich at 04.02.2026
Collectors: D.K. Harder; et al. Field collecting number: DKH7513. Collection date: 08.03.2001.
Administrative regions: VN - Tinh Son La.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
DENNSTAEDTIACEAE
Microlepia

Son La
Moc Chau District; Long Luong
Municipality; Sang Cai Village.
Collections in primary, wet,
broad-leaved, evergreen and
semi-deciduous, mossy forest along
limestone ridge situated between
1400-1500 m elevation.
20°45'51"N 104°51'26"E 1300 - 1500 m
Terrestrial fern. Common.
8 March 2001
D.K. Harder, P.K. Loc, L.V. Averyanov &
N.X. Tam 7513
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318476 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318476
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LE 01318464

LE 01318464

Lepisorus fortuni (T.Moore) C.M.Kuo⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH7118. Collection date: 02.03.2001.
Administrative regions: VN - Tinh Son La.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
POLYPODIACEAE
Lepisorus

Son La
Yen Chau District; Muong Lum
Municipality; Lum Village. Collections
on ridge S of village. Collected with
P.K. Loc, N.T. Hiep, L.V. Averyanov,
G.E. Schatz, S. Bodine & N.Q. Hieu.
21°00'47"N 104°29'30"E 900 - 1134 m
Occasional, lithophytic. Fronds shiny
green; sporangia yellow-brown.
2 March 2001
D.K. Harder & et al. 7118
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318464 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318464
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LE 01318456

LE 01318456

Drynaria fortunei (Kunze ex Mett.) J.Sm.⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6884a. Collection date: 26.02.2001.
Administrative regions: VN - Tinh Lao Cai.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
POLYPODIACEAE
Drynaria

Lao Cai
Van Ban District; Nam Xe Municipality;
Nam Xi Tan Village. Collection in forest
on steep W-facing slope along stream;
just above village; on N side of road
between Nam Xe and Khau Co Pass at near
40 km to Than Uyen mark.
22°02'31"N 103°58'32"E 700 - 1113 m
Epiphytic, 10 m from ground. "Clasping"
frond light brown; rachis on fertile
26 February 2001
D.K. Harder, P.K. Loc, N.T. Hiep, G.E.
Schatz, S. Bodine & N.Q. Hieu 6884
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
----------------------------------------
Page 2
frond shiny black; sporangia brown.
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318456 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318456
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LE 01318455

LE 01318455

Drynaria fortunei (Kunze ex Mett.) J.Sm.⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6884b. Collection date: 26.02.2001.
Administrative regions: VN - Tinh Lao Cai.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
POLYPODIACEAE
Drynaria

Lao Cai
Van Ban District; Nam Xe Municipality;
Nam Xi Tan Village. Collection in forest
on steep W-facing slope along stream;
just above village; on N side of road
between Nam Xe and Khau Co Pass at near
40 km to Than Uyen mark.
22°02'31"N 103°58'32"E 700 - 1113 m
Epiphytic, 10 m from ground. "Clasping"
frond light brown; rachis on fertile
26 February 2001
D.K. Harder, P.K. Loc, N.T. Hiep, G.E.
Schatz, S. Bodine & N.Q. Hieu 6884
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
----------------------------------------
Page 2
frond shiny black; sporangia brown.
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318455 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318455
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LE 01318442

LE 01318442

Lepisorus sp.⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH7498. Collection date: 08.03.2001.
Administrative regions: VN - Tinh Son La.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
POLYPODIACEAE
Lepisorus

Son La
Moc Chau District; Long Luong
Municipality; Sang Cai Village.
Collections in primary, wet,
broad-leaved, evergreen and
semi-deciduous, mossy forest along
limestone ridge situated between
1400-1500 m elevation.
20°45'51"N 104°51'26"E 1300 - 1500 m
Epiphyte fern. Not common.
8 March 2001
D.K. Harder, P.K. Loc, L.V. Averyanov &
N.X. Tam 7498
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318442 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318442
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LE 01318440

LE 01318440

Lepisorus sp.⁣ det. Cheng-Wei Chen at 21.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6819. Collection date: 24.02.2001.
Administrative regions: VN - Tinh Lao Cai.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
POLYPODIACEAE
Lepisorus

Lao Cai
Van Ban District; Nam Xe Municipality.
Collection in small forest patch near
road between Nam Xe Village and Khau Co
Pass.
22°02'26"N 103°59'21"E 612 m
Occasional, lithophytic. Fronds light
green; lower older rachis brown;
sporangia light brown.
24 February 2001
D.K. Harder & et al. 6819
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-24, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318440 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318440
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LE 01311653

LE 01311653

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-23, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01311653 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01311653
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LE 01311652

LE 01311652

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-23, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01311652 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01311652
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LE 01311651

LE 01311651

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-23, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01311651 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01311651
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LE 01311650

LE 01311650

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-23, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01311650 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01311650
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LE 01311649

LE 01311649

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-23, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01311649 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01311649
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LE 01311648

LE 01311648

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-23, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01311648 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01311648
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LE 01311647

LE 01311647

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-23, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01311647 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01311647
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LE 01311646

LE 01311646

Eremostachys iliensis Regel⁣

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-23, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01311646 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01311646
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LE 01311645

LE 01311645

Eremostachys iliensis Regel⁣

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-23, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01311645 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01311645
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LE 01311644

LE 01311644

Eremostachys iliensis Regel⁣

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-23, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01311644 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01311644
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LE 01311643

LE 01311643

Eremostachys iliensis Regel⁣

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-23, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01311643 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01311643
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LE 01311642

LE 01311642

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-23, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01311642 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01311642
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LE 01311641

LE 01311641

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-23, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01311641 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01311641
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LE 01311640

LE 01311640

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-23, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01311640 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01311640
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LE 01311639

LE 01311639

Eremostachys gymnocalyx Schrenk⁣

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-23, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01311639 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01311639
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LE 01311638

LE 01311638

Eremostachys gymnocalyx Schrenk⁣

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-23, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01311638 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01311638
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LE 01311637

LE 01311637

Eremostachys gymnocalyx Schrenk⁣

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-23, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01311637 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01311637
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LE 01311636

LE 01311636

Eremostachys gymnocalyx Schrenk⁣

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-23, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01311636 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01311636
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