Groups of specimens:Образцы сектора Центральной и Восточной АзииRecord creation: 2026-02-26, Ivanenko, PhotoScan D2Citation: Specimen LE 01329247 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01329247
Groups of specimens:Образцы сектора Центральной и Восточной АзииRecord creation: 2026-02-26, Ivanenko, PhotoScan D2Citation: Specimen LE 01329246 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01329246
Groups of specimens:Образцы сектора Центральной и Восточной АзииRecord creation: 2026-02-26, Ivanenko, PhotoScan D2Citation: Specimen LE 01329245 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01329245
Groups of specimens:Образцы сектора Центральной и Восточной АзииRecord creation: 2026-02-26, Ivanenko, PhotoScan D2Citation: Specimen LE 01329244 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01329244
Groups of specimens:Образцы сектора Центральной и Восточной АзииRecord creation: 2026-02-26, Ivanenko, PhotoScan D2Citation: Specimen LE 01329243 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01329243
Groups of specimens:Образцы сектора Центральной и Восточной АзииRecord creation: 2026-02-26, Ivanenko, PhotoScan D2Citation: Specimen LE 01329242 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01329242
Groups of specimens:Образцы сектора Центральной и Восточной АзииRecord creation: 2026-02-26, Ivanenko, PhotoScan D2Citation: Specimen LE 01329241 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01329241
Groups of specimens:Образцы сектора Центральной и Восточной АзииRecord creation: 2026-02-26, Ivanenko, PhotoScan D2Citation: Specimen LE 01329240 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01329240
Groups of specimens:Образцы сектора Центральной и Восточной АзииRecord creation: 2026-02-26, Ivanenko, PhotoScan D2Citation: Specimen LE 01323302 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01323302
Groups of specimens:Образцы сектора Центральной и Восточной АзииRecord creation: 2026-02-26, Ivanenko, PhotoScan D2Citation: Specimen LE 01323301 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01323301
Groups of specimens:Образцы сектора Центральной и Восточной АзииRecord creation: 2026-02-26, Ivanenko, PhotoScan D2Citation: Specimen LE 01323300 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01323300
Groups of specimens:Образцы сектора Центральной и Восточной АзииRecord creation: 2026-02-26, Ivanenko, PhotoScan D2Citation: Specimen LE 01323299 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01323299
Groups of specimens:Образцы сектора Центральной и Восточной АзииRecord creation: 2026-02-26, Ivanenko, PhotoScan D2Citation: Specimen LE 01286506 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01286506
Groups of specimens:Образцы сектора Центральной и Восточной АзииRecord creation: 2026-02-26, Ivanenko, PhotoScan D2Citation: Specimen LE 01286505 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01286505
Groups of specimens:Образцы сектора Центральной и Восточной АзииRecord creation: 2026-02-26, Ivanenko, PhotoScan D2Citation: Specimen LE 01286504 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01286504
Groups of specimens:Образцы сектора Центральной и Восточной АзииRecord creation: 2026-02-26, Ivanenko, PhotoScan D2Citation: Specimen LE 01286503 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01286503
Groups of specimens:Образцы сектора Центральной и Восточной АзииRecord creation: 2026-02-26, Ivanenko, PhotoScan D2Citation: Specimen LE 01286502 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01286502
Groups of specimens:Образцы сектора Центральной и Восточной АзииRecord creation: 2026-02-26, Ivanenko, PhotoScan D2Citation: Specimen LE 01286501 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01286501
Groups of specimens:Образцы сектора Центральной и Восточной АзииRecord creation: 2026-02-26, Ivanenko, PhotoScan D2Citation: Specimen LE 01286500 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01286500
Groups of specimens:Образцы сектора Центральной и Восточной АзииRecord creation: 2026-02-26, Ivanenko, PhotoScan D2Citation: Specimen LE 01286499 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01286499
Groups of specimens:Образцы сектора Центральной и Восточной АзииRecord creation: 2026-02-26, Ivanenko, PhotoScan D2Citation: Specimen LE 01286498 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01286498
Groups of specimens:Образцы сектора Центральной и Восточной АзииRecord creation: 2026-02-26, Ivanenko, PhotoScan D2Citation: Specimen LE 01286497 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01286497
Groups of specimens:Образцы сектора Центральной и Восточной АзииRecord creation: 2026-02-26, Ivanenko, PhotoScan D2Citation: Specimen LE 01286496 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01286496
Groups of specimens:Образцы сектора Центральной и Восточной АзииRecord creation: 2026-02-26, Ivanenko, PhotoScan D2Citation: Specimen LE 01286495 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01286495
Groups of specimens:Образцы сектора Центральной и Восточной АзииRecord creation: 2026-02-26, Ivanenko, PhotoScan D2Citation: Specimen LE 01286494 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01286494
Groups of specimens:Образцы сектора КавказаRecord creation: 2026-02-26, Andrey Sytin, PhotoScan D2Citation: Specimen LE 01357776 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01357776
Groups of specimens:Образцы сектора КавказаRecord creation: 2026-02-26, Andrey Sytin, PhotoScan D2Citation: Specimen LE 01357775 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01357775
N.Vietnam, Ha Tinh Prov., Huong Son Distr., Son Hong Municipality, around point 18º34'06''N, 105º11'40''E. Primary broad-leaved evergreen riparian forest with Platanus kerrii and Duabanga grandiflora along rocky Song Con river valley composed with shale and gray sandstone at elevation 250-300 m a.s.l. Terrestrial fern on open wet river rocks. Very common.
Record creation: 2026-02-25, Аверьянов Леонид ВладимировичCitation: Specimen LE 01203664 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01203664
N.Vietnam, Bac Kan Prov., Cho Don Distr., Ban Thi Municipality, village Phia Khao, Lung Li valley, around point 22º16'10''N, 105º31'01''E, Cot Moc mountain at elevation 800-900 m a.s.l. Primary broad-leaved evergreen closed forest on very steep slopes and cliffs near tops of rocky ridge composed with solid marble-like stratified limestone. Lithophytic fern on vertical cliffs. Very common.
Record creation: 2026-02-25, Аверьянов Леонид ВладимировичCitation: Specimen LE 01203663 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01203663
N.Vietnam, Cao Bang Prov., Trung Khanh Distr., Ngoc Khe Municipality, Pac Nga village, around point 22º54'59''N, 106º31'44''E, at elevation 800-850 m a.s.l. Remnants of primary evergreen broad-leaved forest along tops of rocky ridges and mesae composed with light gray solid marble-like highly eroded crystalline limestone. Lithophyte on wet cliff. Common.
Record creation: 2026-02-25, Аверьянов Леонид ВладимировичCitation: Specimen LE 01203662 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01203662
Record creation: 2026-02-25, Аверьянов Леонид ВладимировичCitation: Specimen LE 01203661 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01203661
N.Vietnam, Cao Bang Prov., Trung Khanh Distr., Ngoc Khe Municipality, Pac Nga village, around point 22º54'59''N, 106º31'44''E, at elevation 800-850 m a.s.l. Remnants of primary broad-leaved evergreen and coniferous forest along tops of rocky ridges and mesae composed with light gray solid marble-like highly eroded crystalline limestone. Lithophytic fern on mossy exposed rocks and on vertical shady cliffs. Very common.
Record creation: 2026-02-25, Аверьянов Леонид ВладимировичCitation: Specimen LE 01203660 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01203660
N.Vietnam, Cao Bang Prov., Trung Khanh Distr., Ngoc Khe Municipality, Pac Nga village, around point 22º54'59''N, 106º31'44''E, at elevation 500-550 m a.s.l. Secondary shrubs on very steep mossy rocky slope of N exposition of rocky ridge composed with light gray solid marble-like highly eroded crystalline limestone along Quy Son River valley. Lithophytic fern on vertical mossy cliffs. Rare.
Record creation: 2026-02-25, Аверьянов Леонид ВладимировичCitation: Specimen LE 01203659 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01203659
N.Vietnam, Bac Kan Prov., Cho Don Distr., Ban Thi Municipality, village Phia Khao, around point 22º17'31''N, 105º31'51''E, Cau Muc mountain at elevation 800-900 m a.s.l. Primary broad-leaved evergreen closed forest on very steep slopes and cliffs near tops of rocky ridge composed with solid marble-like stratified limestone. Lithophytic fern in crevices of vertical open cliffs. Common.
Record creation: 2026-02-25, Аверьянов Леонид ВладимировичCitation: Specimen LE 01203658 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01203658
N.Vietnam, Bac Kan Prov., Cho Don Distr., Ban Thi Municipality, village Phia Khao, around point 22º15'35''N, 105º31'17''E at elevation 800-900 m a.s.l. Remnants of primary broad-leaved evergreen closed wet forest on very steep slopes and cliffs of rocky ridges and mesae composed with solid marble-like stratified limestone. Lithophyte on vertical cliff. Common.
Record creation: 2026-02-25, Аверьянов Леонид ВладимировичCitation: Specimen LE 01203657 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01203657
Ha Giang Prov., Bac Me Distr., Phieng Luong municipality, around Phieng Day village. Degraded primary broad-leaved humid evergreen forest along rocky canyon of small stream at the base of mountains composed with solid crystalline highly eroded limestone at elevation 900-1000 m a.s.l. around point 22°38’43.8’’N 105°19’19.8’’E. Lythophytic fern on mossy rocks. Not common.
Record creation: 2026-02-25, Аверьянов Леонид ВладимировичCitation: Specimen LE 01203656 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01203656
Record creation: 2026-02-25, Аверьянов Леонид ВладимировичCitation: Specimen LE 01203655 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01203655
N.Vietnam, Bac Kan Prov., Na Ri Distr., Liem Thuy Municipality, village Na Bo, around point 21º56'44''N, 106º05'09''E, at elevation 300-700 m a.s.l. Primary broad-leaved evergreen closed wet forest on very steep slopes and cliffs of rocky ridges and mesae composed with white solid marble-like crystalline limestone. Lithophytic fern on shady vertical cliffs. Very common.
Record creation: 2026-02-25, Аверьянов Леонид ВладимировичCitation: Specimen LE 01203654 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01203654
Record creation: 2026-02-25, Аверьянов Леонид ВладимировичCitation: Specimen LE 01203653 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01203653
Record creation: 2026-02-25, Аверьянов Леонид ВладимировичCitation: Specimen LE 01203652 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01203652
N.Vietnam, Bac Kan Prov., Cho Don Distr., Ban Thi Municipality, village Phia Khao, around point 22º15'52''N, 105º32'01''E at elevation 800-900 m a.s.l. (Po Luong mountain). Primary broad-leaved evergreen closed forest on very steep slopes and cliffs near and on tops of rocky ridge composed with solid marble-like stratified limestone. Lithophytic and epiphytic fern in shady place. Very common.
Record creation: 2026-02-25, Аверьянов Леонид ВладимировичCitation: Specimen LE 01203651 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01203651
Bolikhamsai prov., Thaphabat distr., Nam Bon village, Phou Khao Khouay national park, Tad Xai Waterfall, around point 18°27'25.9''N 103°08'17.5''E. Degraded primary and secondary evergreen dry forest along river on eroded sandstone at elev. About 340 m a.s.l. Lithophytic fern on shady mossy boulders and river cliffs. Common.
Record creation: 2026-02-25, Аверьянов Леонид ВладимировичCitation: Specimen LE 01203650 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01203650
Record creation: 2026-02-25, Аверьянов Леонид ВладимировичCitation: Specimen LE 01203649 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01203649
Record creation: 2026-02-25, Аверьянов Леонид ВладимировичCitation: Specimen LE 01203648 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01203648
N.Vietnam, Bac Kan Prov., Na Ri Distr., Liem Thuy Municipality, village Lung Vai, around point 21º57'18''N, 106º06'12''E, at elevation 500-700 m a.s.l. Primary broad-leaved evergreen closed wet forest on very steep slopes and cliffs of rocky ridges and mesae composed with white solid marble-like crystalline limestone. Epiphyte. Common.
Record creation: 2026-02-25, Аверьянов Леонид ВладимировичCitation: Specimen LE 01203647 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01203647
Additional identifications:Asplenium cardiophyllum (Hance) Baker det. Averyanov, Leonid Vladimirovich at 07.02.2009; Hymenasplenium cardiophyllum (Hance) Nakaike det. Averyanov, Leonid Vladimirovich at 25.02.2026Collectors:Averyanov, Leonid Vladimirovich; et al.Field collecting number: HAL12357. Collection date: 07.02.2009. Administrative regions:VN - Tinh Quang Binh. Groups of specimens:L. V. Averyanov specimens; L. V. Averyanov photos; Flora of VietnamOriginal label text:
FLORA OF VIETNAM
Polypodiaceae s.l.
Asplenium cardiophyllum
Quang Binh Prov., Minh Hoa Distr., Dan Hoa Municipality, Bai Dinh village around point 17°45’28”N 105°46’14”E. Remnants of primary evergreen broad-leaved forest on very steep slopes and on tops of low remnant hills composed with highly eroded marble-like solid limestone at elevation about 200-300 m a.s.l. Lithophytic fern on wet travertine deposits on tall vertical shady cliff composed with outcrops of orange-brown crystalline limestone. Rare.
Record creation: 2026-02-25, Аверьянов Леонид ВладимировичCitation: Specimen LE 01203646 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01203646
N.Vietnam, Cao Bang Prov., Trung Khanh Distr., Ngoc Khe Municipality, Pac Nga village, around point 22º54'59''N, 106º31'44''E, at elevation 500-600 m a.s.l. Secondary open shrubs on wet rocky slope of N exposition. Terrestrial and lithophytic fern on mossy places. Not common.
Record creation: 2026-02-25, Аверьянов Леонид ВладимировичCitation: Specimen LE 01203645 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01203645
N.Vietnam, Bac Kan Prov., Ba Be Distr., Nam Mau Municipality, Ba Be National Park around point 22o25'36''N 105o37'50''E. Heavily logged primary evergreen dry broad-leaved forest on very steep slopes and cliffs composed with crystalline highly eroded marble-like limestone along Ba Be lake at elevation 200-250 m a.s.l. Lithophytic herb on open cliffs. Very common.
HLF 584 22 Dec. 2002. Photo.
Coll. Phan Ke Loc, L.Averyanov, Hoang Van Chat, Do Hoang Chung, Nguyen Manh Hung, Vu Hong Van, Nguyen Tien Vinh, Mai Van Xinh. Grant from Henry Luce Foundation.
Record creation: 2026-02-25, Аверьянов Леонид ВладимировичCitation: Specimen LE 01203644 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01203644
Ad fluvios Olenek et Lenam inferiorem, et inter Lena et Olenek
Record creation: 2026-02-25, Marina Yarichina, PhotoScan D2Citation: Specimen LE 01357774 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01357774