Herbarium LE - Specimens

VLORA OF VIETNAM

Lygodium microphyllum (Cav.) R.Br.⁣?

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 1048, 20.10.1990

VLORA OF VIETNAM

Lygodium microphyllum (Cav.) R.Br.⁣?

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 1048, 20.10.1990

VLORA OF VIETNAM

Lygodium microphyllum (Cav.) R.Br.⁣

Quang Ngai Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 456, 07.10.1990

VLORA OF VIETNAM

Lygodium microphyllum (Cav.) R.Br.⁣

Quang Ngai Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 456, 07.10.1990

VLORA OF VIETNAM

Cibotium barometz (L.) J. Sm.⁣

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 814, 15.10.1990

VLORA OF VIETNAM

Cibotium barometz (L.) J. Sm.⁣

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 814, 15.10.1990

VLORA OF VIETNAM

Cibotium barometz (L.) J. Sm.⁣

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 871, 16.10.1990

VLORA OF VIETNAM

Grypothrix triphylla (Sw.) S.E.Fawc. & A.R.Sm.⁣

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 755, 15.10.1990

VLORA OF VIETNAM

Grypothrix triphylla (Sw.) S.E.Fawc. & A.R.Sm.⁣

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 755, 15.10.1990

VLORA OF VIETNAM

Blechnum orientale L.⁣

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 679a, 12.10.1990

VLORA OF VIETNAM

Blechnum orientale L.⁣

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 595, 10.10.1990

Polypodiaceae s.l.

Onichium orientale ?

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality,elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary closed mountain broad-leaved evergreen forest on very steep granite slopes along river canyon at elev. 1400 m. Terrestrial fern with fronds up to 3 m lg on very steep river side slope. Common.

No VH 6383 12.05.2000

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 6383-98)

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

VLORA OF VIETNAM

Blechnum orientale L.⁣

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 595, 10.10.1990

VLORA OF VIETNAM

Blechnum orientale L.⁣

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 817, 15.10.1990

VLORA OF VIETNAM

Adiantum philippense L.⁣

Khanh Hoa Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 011, 27.09.1990

VLORA OF VIETNAM

Adiantum philippense L.⁣

Khanh Hoa Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 011, 27.09.1990

FLORA OF VIETNAM

Polypodiaceae s.l.

Cyclosurus sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of

Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary wet closed mountain broad-leaved and mixed forest on

granites along very steep shady slopes of river canyon at elev. 1500-1700 m. Terrestrial and lithophytic fern in open mossy slopes. Not rare.

No VH 6050 11.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

Polypodiaceae s.l.

Cyclosurus sp. ?

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary mossy cloud mixed forest (with Pinus, Fokienia, Dacrydium and Dacrycarpus) on very steep granite slopes at elev. 2440 m. Creeping epiphyte in short tall forest on the top of main ridge. Not common.

No VH 6199 09.05.2000

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 6383-98)

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

VIETNAM

Pteridophyta J.Y. Bergen & B.M. Davis

Ha Giang

Quan Ba District; Bat Dai Son

Municipality; Bat Dai Son Provincial

Protected Area (2000); ridge N of Bat

Dai Son border station.

23°09'11"N 104°59'06"E 1287 m

Occasional, on rocks. Rachis and rhizome

hair dark brown to black; fronds with

light yellow-green areas; sporangia

brown.

11 February 2001

D.K. Harder, N.T. Hiep, L.V. Averyanov &

P.K. Loc 6161

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

VIETNAM

MARATTIACEAE

Angiopteris

Lao Cai

Van Ban District; Liem Phu Municipality;

Khuoi Ngoa Village. Collections on

ridges just west of village.

21°59'31"N 104°19'05"E 323 - 1050 m

Occasional, in shade. Rachis red-purple;

shiny, wiry, with glaucous bloom at

"nodes," marginal sporangia light brown.

22 February 2001

D.K. Harder, P.K. Loc, L.V. Averyanov,

G.E. Schatz & S. Bodine 6734

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

VIETNAM

HYMENOPHYLLACEAE

Trichomanes

Lao Cai

Van Ban District; Liem Phu Municipality;

Khuoi Ngoa Village.

21°58'20"N 104°20'16"E 527 - 1120 m

21°58'08"N 104°21'00"E

Occasional, climbing epiphytic. Pods

deep green, transparent; sporangial

"cups" green.

21 February 2001

D.K. Harder, L.V. Averyanov & P.K. Loc

6596

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

VIETNAM

HYMENOPHYLLACEAE

Trichomanes

Lao Cai

Van Ban District; Liem Phu Municipality;

Khuoi Ngoa Village.

21°58'20"N 104°20'16"E 527 - 1120 m

21°58'08"N 104°21'00"E

Climbing, epiphytic, transplant,

occasional. Fronds dull green; rachis

stiff dark brown.

21 February 2001

D.K. Harder, L.V. Averyanov & P.K. Loc

6616

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

FLORA OF VIETNAM

Polypodiaceae s.l.

Adianthum sp. ?

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of

Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary wet closed mountain broad-leaved and mixed forest on

granites along very steep shady slopes of river canyon at elev. 1700-1750 m. Terrestrial fern on very steep slope of river canyon.

Common.

No VH 6423 11.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

VIETNAM

DENNSTAEDTIACEAE

Lindsaea

Lao Cai

Van Ban District; Nam Xe Municipality;

Nam Xi Tan Village. Collection in forest

on steep W-facing slope along stream;

just above village; on N side of road

between Nam Xe and Khau Co Pass at near

40 km to Than Uyen mark.

22°02'31"N 103°58'32"E 700 - 1113 m

Occasional, on slope. Rachis

brown-black, shiny; rhizome brown-black;

26 February 2001

D.K. Harder, P.K. Loc, N.T. Hiep, G.E.

Schatz, S. Bodine & N.Q. Hieu 6890

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

----------------------------------------

Page 2

sporangia brown.

VIETNAM

DRYOPTERIDACEAE

Cyrtogonellum

Son La

Moc Chau District; Long Luong

Municipality; Sang Cai Village.

Collections in primary, wet,

broad-leaved, evergreen and

semi-deciduous, mossy forest along

limestone ridge situated between

1400-1500 m elevation.

20°45'51"N 104°51'26"E 1300 - 1500 m

Terrestrial and lithophytic fern on

8 March 2001

D.K. Harder, P.K. Loc, L.V. Averyanov &

N.X. Tam 7485

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

----------------------------------------

Page 2

steep rocky slope. Not rare.

VIETNAM

DICKSONIACEAE

Cibotium

Ha Giang

Vi Xuyen District; Phong Quang

Municipality; Man Village. Collection

on ridge and small limestone peak just

east of house of village head; very

disturbed, secondary vegetation on lower

slopes and on limestone rocks up to

ridgetop.

22°53'11"N 104°56'43"E 437 - 645 m

22°53'23"N 104°56'48"E

Tree fern, to 5 m. Rachis brown;

14 February 2001

D.K. Harder, P.K. Loc, G.E. Schatz &

L.V. Averyanov 6301

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

----------------------------------------

Page 2

someplaces shiny; stem to 42 cm;

sporangia dark brown; fronds to 2 m.

VIETNAM

DRYOPTERIDACEAE

Cyrtogonellum

Son La

Moc Chau District; Van Ho Municipality,

Hua Tat Village. Collections on

limestone ridge from degraded forests to

cultivation on lower slopes to primary

evergreen, closed forest on limestone

ridge and cliffs, some coniferous forest

with Pinus kwangtungensis and Calocedrus

macrolepis.

20°46'35"N 104°46'31"E 1400 - 1500 m

6 March 2001

D.K. Harder, P.K. Loc, L.V. Averyanov &

N.X. Tam 7344

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

----------------------------------------

Page 2

Terrestrial and lithophytic fern on

steep slope. Not rare.

VIETNAM

DENNSTAEDTIACEAE

Lindsaea

Lao Cai

Van Ban District; Liem Phu Municipality;

Khuoi Ngoa Village. Collections on

ridges just west of village.

21°59'31"N 104°19'05"E 323 - 1050 m

Widespread, terrestrial. Rachis

purple-black; with dark brown hairs at

base; sporangia brown.

22 February 2001

D.K. Harder, P.K. Loc, L.V. Averyanov,

G.E. Schatz & S. Bodine 6698

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

VIETNAM

DENNSTAEDTIACEAE

Lindsaea

Lao Cai

Van Ban District; Liem Pho Municipality;

Khuoi Ngoa Village. Collections in

disturbed secondary vegetation and

cultivated tree plantation, along road

between tree [Fokienia (Pou Nhou)]

extraction camp along road to near Khuoi

Ngoa school.

21°58'14"N 104°19'51"E 606 m

Common, on roadside. Lower rachis

20 February 2001

D.K. Harder, P.K. Loc, L.V. Averyanov,

G.E. Schatz & S. Bodine 6517

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

----------------------------------------

Page 2

red-brown; edge sporangia light green.

VIETNAM

HYMENOPHYLLACEAE

Trichomanes

Son La

Moc Chau District; Long Luong

Municipality; Sang Cai Village.

Collections in primary, wet,

broad-leaved, evergreen and

semi-deciduous, mossy forest along

limestone ridge situated between

1400-1500 m elevation.

20°45'51"N 104°51'26"E 1300 - 1500 m

Lithophytic fern on shady wet rocks. Not

8 March 2001

D.K. Harder, P.K. Loc, L.V. Averyanov &

N.X. Tam 7475

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

----------------------------------------

Page 2

common.

VIETNAM

PTERIDACEAE

Adiantum

Ha Giang

Vi Xuyen District; Phong Quang

Municipality; Man Village, Phong Quang

Nature Reserve. Collection on S-facing

slope from secondary grasslands to

limestone cliffs to area of primary

forest on ridge, on limestone.

22°52'04"N 104°57'32"E 300 - 1050 m

Frequent, on vertical cliffs. Rachis

brown; sporangia dark, dull brown.

16 February 2001

D.K. Harder & L.V. Averyanov 6427

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

VIETNAM

Pteridophyta J.Y. Bergen & B.M. Davis

Ha Giang

Vi Xuyen District; Phong Quang

Municipality; Man Village. Limestone

peaks and ridges NE of Village.

22°53'21"N 104°56'45"E 595 - 787 m

Occasional, in disturbed forest patch;

near wetter area. Rachis dark

purple-black; marginal sporangia dull

brown.

15 February 2001

D.K. Harder, P.K. Loc, L.V. Averyanov,

G.E. Schatz & S. Bodine 6387

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

VIETNAM

PTERIDACEAE

Pteris

Ha Giang

Vi Xuyen District; Phong Quang

Municipality; Man Village, Phong Quang

Nature Reserve. Collection on S-facing

slope from secondary grasslands to

limestone cliffs to area of primary

forest on ridge, on limestone.

22°52'04"N 104°57'32"E 300 - 1050 m

Occasional, in open, in middle forest.

Rachis brown; sporangia dark brown;

fronds to 2.5 m.

16 February 2001

D.K. Harder & L.V. Averyanov 6425

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

FLORA OF VIETNAM

Polypodiaceae

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) along Dak Tour

river on N macroslope of Chu Yang Sinh mt system to main peak (12ш24'N, 108ш26'E). Primary broad-leaved mountain evergreen closed

forest on granites along river slopes at elev. 1000-1200 m. Epiphyte fern on mossy trees along shady waterfall sides. Very common.

No VH 6022 05-06.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

VIETNAM

PTERIDACEAE

Pteris semipinnata L.

det. D.K. Harder, 2001

Lao Cai

Van Ban District; Liem Phu Municipality;

Khuoi Ngoa Village. Collections on

ridges just west of village.

21°59'31"N 104°19'05"E 323 - 1050 m

Frequent, in shade, widespread. Rachis

shiny dark purple-black; marginal

sporangia light brown; fronds to 1.0 m.

22 February 2001

D.K. Harder, P.K. Loc, L.V. Averyanov,

G.E. Schatz & S. Bodine 6668

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

FLORA OF VIETNAM

Polypodiaceae s.l.

Lindsaea sp. ?

S. Vietnam, Kontum Prov., Kon Plong Distr., Hieu Municipality, Mang La forest enterprise (14ш39'N, 108ш25'E). Primary broad-leaved closed

evergreen wet mountain forest on sandstone steep slopes of forest river at elev. 1100-1200 m. Terrestrial fern on dry steep slope.

Locally common.

No VH 5677 24.04.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, P.H.Hoang, D.X.Du, N.T.Vinh

Marattiaceae

Archangiopteris sp. ?

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary closed wet broad-leaved forest on steep granite slopes at elev. 1550-1600 m. Terrestrial fern with fronds up to 1 m lg in wet cliffs of very deep shady river canyon. Rare.

No VH 6300 07.05.2000

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 6383-98)

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

Convallariaceae

Tupistra thangii N.Tanaka & D.D.Nguyen

Lam Dong Province, Bao Lam District, B 40 Pass, around point N11.4366 E107.4266. Fractionally logged primary evergreen broad-leaved and mixed forest with Dacrydium, Dacrycarpus and Pinus latteri on granite at elevation 800–900 m a.s.l. Terrestrial herb in dense forest. Locally common. 28 November 2022, L. Averyanov, Nguyen Van Canh, T. Maisak, Truong Ba Vuong, Nguyen Quoc Bao, AL 1862.

Living plant obtained at 25 Nov. 2022, Nguyen Van Canh, L. Averyanov, T. Maisak, AL 1700.

Clonotype

Flowered in culture on 12.12.2025, L. Averyanov, T. Maisak, AL 1700

Orchidaceae

Nervilia gracilis Aver.

Son La Province, Thuan Chau District, limestone forest, at elevation about 900 m a.s.l.

May 2024, Nguyen Van Canh, AL3299.

Orchidaceae

Zeuxine nervosa?

Gia Lai Province, Krong Pa District, evergreen broad-leaved forest at elevation of about 600 m a.s.l.

2024, Nguyen Van Canh, AL3794.

Orchidaceae

Hetaeria affinis (Griff.) Seidenf. & Ormerod

Dak Lak Province, Buon Ma Thuot City area, forest park, not common.

2021, Nguyen Van Canh, AL3304.

Orchidaceae

Dendrobium heterocarpum Lindl.

Annonaceae

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Treelet to 6-7 m tall on shady slope. Flowers dark green. Not common.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4232 (NHC043).

Annonaceae

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Treelet or shrub 2-3 m tall. Flowers yellow, fruits green. Not common.

9 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4218 (NHC029).

Orchidaceae

Coelogyne sp.

Dak Lak Province, Krong Bong District, Yang Hanh Mountain, evergreen broad-leaved forest at elevation of about 1200 m a.s.l.

February 2025, Nguyen Van Canh, AL3812.

Orchidaceae

Hetaeria sp.

Dak Lak Province, Krong Bong District, Yang Hanh Mountain, evergreen broad-leaved forest at elevation of about 1200 m a.s.l.

February 2025, Nguyen Van Canh, AL3814.

Orchidaceae

Cleisostoma williamsonii (Rchb.f.) Garay

Kon Tum Province, Dak Gley District, W slopes of Ngoc Linh Mountain system. Remnants of primary and secondary evergreen broad-leaved forest on granite and shale at elevation about 1200 m a.s.l. Clustering epiphyte in canopies of tall trees. Flowers pinkish. Locally common.

02 May 2025, L. Averyanov, N.V. Canh, T. Maisak, AL4104.

Orchidaceae

Cleisostoma williamsonii (Rchb.f.) Garay

Kon Tum Province, Dak Gley District, W slopes of Ngoc Linh Mountain system. Remnants of primary and secondary evergreen broad-leaved forest on granite and shale at elevation about 1200 m a.s.l. Clustering epiphyte in canopies of tall trees. Flowers pinkish. Locally common.

02 May 2025, L. Averyanov, N.V. Canh, T. Maisak, AL4104.

Orchidaceae

Pomatocalpa grandis Seidenf.

Lam Dong Province, Duc Trong District, evergreen forest, at elevation 900-1000 m a.s.l.

2022, Nguyen Van Canh, AL3301.