Herbarium LE - Specimens

VIETNAM

ASPLENIACEAE

Asplenium

Ha Giang

Vi Xuyen District; Minh Tan Municipality

Ban Ngay Village. Collection on E

facing ridge above river; on exposed

limestone rocks; collections in

disturbed agricultural land leading into

primary forest with some deforestation

of Burretiodendron (Tiliaceae) trees.

23°00'09"N 104°55'21"E 225 - 621 m

23°00'37"N 104°54'46"E

Frequent epiphyte and occasional

17 February 2001

D.K. Harder, P.K. Loc, N.T. Hiep, G.E.

Schatz & S. Bodine 6464

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

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Page 2

lithophyte. Fronds light green;

sporangia dull, light brown; midrib of

frond purple-brown, producing "nests"

when large.

VIETNAM

ASPLENIACEAE

Asplenium

Son La

Moc Chau District; Van Ho Municipality,

Hua Tat Village. Collections on

limestone ridge from degraded forests to

cultivation on lower slopes to primary

broadleaved, closed wet forest along

limestone ridge and slopes near tops.

20°46'13"N 104°46'12"E 1100 - 1300 m

Lithophytic fern. Common.

7 March 2001

D.K. Harder, P.K. Loc, L.V. Averyanov &

N.X. Tam 7392

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

Asplenium antrophyoides Christ

Northern Vietnam, Thanh Hoa, Pu Luong Nature Reserve.

2019

Coll. : Cuong Huu Nguyen CLL 529

Det. Averyanov L. 26.12.2025

VIETNAM

GRAMMITIDACEAE

Grammitis

Son La

Moc Chau District; Van Ho Municipality,

Hua Tat Village. Collections on

limestone ridge from degraded forests to

cultivation on lower slopes to primary

broadleaved, closed wet forest along

limestone ridge and slopes near tops.

20°46'13"N 104°46'12"E 1100 - 1300 m

Epiphyte in shady places. Common.

7 March 2001

D.K. Harder, P.K. Loc, L.V. Averyanov &

N.X. Tam 7411

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

VIETNAM

GRAMMITIDACEAE

Grammitis

Son La

Moc Chau District; Van Ho Municipality,

Hua Tat Village. Collections on

limestone ridge from degraded forests to

cultivation on lower slopes to primary

evergreen, closed forest on limestone

ridge and cliffs, some coniferous forest

with Pinus kwangtungensis and Calocedrus

macrolepis.

20°46'35"N 104°46'31"E 1400 - 1500 m

6 March 2001

D.K. Harder, P.K. Loc, L.V. Averyanov &

N.X. Tam 7335

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

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Page 2

Epiphyte in shady place. Not rare.

POLYPODIACEAE

Pyrrosia

Ha Giang

Quan Ba District; Bat Dai Son

Municipality; Bat Dai Son Provincial

Protected Area (2000). On limestone

ridge of degraded forest just northwest

of Bai Dai Son border station;

collections in forest above cultivations

(Forest possibly burned ca. 25 years ago

or so, only small trees and few fire

tolerant herbs).

13°09'13"N 104°59'36"E 1064 m

10 February 2001

D.K. Harder, Hiep, P.K. Loc, L.V.

Averyanov, G.E. Schatz & S.A. Bodin 6092

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

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Page 2

Epiphytic. Fronds light green; sporangia

rusty-brown.

VIETNAM

GRAMMITIDACEAE

Grammitis

Ha Giang

Vi Xuyen District; Phong Quang

Municipality; Man Village. Limestone

peaks and ridges NE of Village.

22°53'21"N 104°56'45"E 595 - 787 m

Rare, lithophytic, in shade. Sporangia

brown.

15 February 2001

D.K. Harder, P.K. Loc, L.V. Averyanov,

G.E. Schatz & S. Bodine 6359

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

GRAMMITIDACEAE

Grammitis

Son La

Moc Chau District; Long Luong

Municipality; Sang Cai Village.

Collections in primary, wet,

broad-leaved, evergreen and

semi-deciduous, mossy forest along

limestone ridge situated between

1400-1500 m elevation.

20°45'51"N 104°51'26"E 1300 - 1500 m

Epiphyte in shady places. Common.

8 March 2001

D.K. Harder, P.K. Loc, L.V. Averyanov &

N.X. Tam 7488

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

VIETNAM

Grammitideae C. Presl

Son La

Yen Chau District; Muong Lum

Municipality; On Oc Village; collection

along N-S ridge along western side.

Collection along trail from agricultural

field to secondary vegetation to primary

forest vegetation with some trees

removed along ridge top. Limestone

substrate (crystalline). GPS coordinates

at top of ridge.

20°59'06"N 104°28'46"E 1468 m

4 March 2001

D.K. Harder, P.K. Loc & N.T. Huong 7261

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

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Page 2

Epiphytic. Root brown; raised sporangiabrown.

Pteridophyta J.Y. Bergen & B.M. Davis

Ha Giang

Vi Xuyen District; Phong Quang

Municipality; Man Village. Limestone

peaks and ridges NE of Village.

22°53'21"N 104°56'45"E 595 - 787 m

Epiphytic, occasional, near top of small

hill. Rhizome and roots dark, dull

brown; sporangia light brown in small

light green cups.

15 February 2001

D.K. Harder, P.K. Loc, L.V. Averyanov,

G.E. Schatz & S. Bodine 6379

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

FLORA OF VIETNAM

Polypodiaceae

Asplenium sp. ?

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) along Dak Tour

river on N macroslope of Chu Yang Sinh mt system to main peak (12ш24'N, 108ш26'E). Primary broad-leaved mountain evergreen closed

forest on granites along river slopes at elev. 1000-1200 m. Epiphyte fern with pendent fronds up to 30 cm lg on mossy trees along shady

waterfall sides. Very common.

No VH 6021 05-06.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

FLORA OF VIETNAM

Polypodiaceae s.l.

Asplenium sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of

Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary mossy cloud mixed forest (with Pinus, Fokienia,

Dacrydium and Dacrycarpus) on very steep granite slopes at elev. 2100-2300 m. Epiphyte on mossy trees in shady place. Very common.

VH 6215 09.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

POLYPODIACEAE

Pyrrosia adnascens (Sw.) Ching

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 728, 12.10.1990

POLYPODIACEAE

Taenitis blechnoides (Willd.) Sw.⁣

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 728, 12.10.1990

POLYPODIACEAE

Pyrrosia adnascens (Sw.) Ching

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 987, 19.10.1990

POLYPODIACEAE

Pyrrosia adnascens (Sw.) Ching

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 1062, 20.10.1990

POLYPODIACEAE

Taenitis blechnoides (Willd.) Sw.⁣

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 679, 11.10.1990

FLORA OF VIETNAM

Polypodiaceae s.l.

Pteris sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of

Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary closed wet broad-leaved forest on steep granite slopes

at elev. 1300 m. Terrestrial fern on steep shady slope. Not common.

No VH 6291 07.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

FLORA OF VIETNAM

Polypodiaceae s.l.

Woodwardtia sp. ?

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of

Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary closed wet broad-leaved forest on steep granite slopes

at elev. 1300 m. Terrestrial fern on steep shady slope. Not common.

No VH 6290 07.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

FLORA OF VIETNAM

Polypodiaceae s.l.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of

Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary wet closed mountain broad-leaved forest on granites

along river slopes at elev. 500-900 m. Lithophytic fern on shady mossy river rocks. Common.

No VH 6070 04.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

FLORA OF VIETNAM

Polypodiaceae s.l.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of

Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary closed mountain broad-leaved evergreen forest on very steep granite slopes along river canyon at elev. 1200-1300 m.

Lithophytic fern on wet mossy shady rocks along river. Common.

No VH 6387 12.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

FLORA OF VIETNAM

Polypodiaceae

Asplenium aff.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of

Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary wet closed mountain broad-leaved and mixed forest on

granites along very steep shady slopes of river canyon at elev. 1750 m. Epiphyte at base of large trees on open mossy cliff. Common.

No VH 6318 11.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

FLORA OF VIETNAM

Polypodiaceae s.l.

Dipteris aff.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of

Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary closed wet broad-leaved forest on steep granite slopes

at elev. 1200-1250 m. Lithophytic fern in crevices of open mossy cliffs in river canyon. Not common.

No VH 6276 06.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

FLORA OF VIETNAM

Polypodiaceae s.l.

Dipteris aff.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of

Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary wet closed mountain broad-leaved and mixed forest on

granites along very steep shady slopes of river canyon at elev. 1750 m. Lithophytic and terrestrial fern on shady mossy cliffs. Not common.

No VH 6328 11.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

Polypodiaceae

Angiopteris evecta (G. Forst.) Hoffm.⁣

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 812, 15.10.1990

VIETNAM

MARATTIACEAE

Angiopteris

Lao Cai

Van Ban District; Liem Pho Municipality;

Khuoi Ngoa Village. Collections in

disturbed secondary vegetation and

cultivated tree plantation, along road

between tree [Fokienia (Pou Nhou)]

extraction camp along road to near Khuoi

Ngoa school.

21°58'14"N 104°19'51"E 606 m

Occasional; frond to 2 m; in understory.

20 February 2001

D.K. Harder, P.K. Loc, L.V. Averyanov,

G.E. Schatz & S. Bodine 6527

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

----------------------------------------

Page 2

Pinnae deep green, lighter below;

marginal sporangia dull brown; clear

green sap from cut rachis.

FLORA OF VIETNAM

Marattiaceae

Angiopteris sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of

Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary closed wet broad-leaved and mixed forest (with Pinus,

Fokienia, Dacrydium and Dacrycarpus) on steep granite slopes at elev. 1200-1300 m. Terrestrial and lithophytic fern with fronds up to 4 m lg

on very steep rocky shady slope of river canyon along river. Not rare.

No VH 6238 08.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

VIETNAM

MARATTIACEAE

Angiopteris

Lao Cai

Van Ban District; Nam Xe Municipality.

Collection in small forest patch near

road between Nam Xe Village and Khau Co

Pass.

22°02'26"N 103°59'21"E 612 m

Occasional, fronds to 2.5 m. Pinnae dark

green, lighter below, sporangia brown;

short stem to 0.5 m.

24 February 2001

D.K. Harder & et al. 6795

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

Polypodiaceae s.l.

Belvisia sp.

Dien Bien Prov., Muong Nhe Distr., Muong Nhe Natural Reserve, Sin Thau Municipality, Ta Mieu Village around point about N 22º24’02’’ E 102º08’38”. Old humid secondary evergreen broad-leaved montane forest of very steep mountain slopes composed with sandstone at elevations 1800–1850 m a.s.l. Epiphyte on mossy tree on mountain top. Leaves almost red. Locally common.

14 May 2022, L. Averyanov, Tran Huy Thai, Khang Sinh Nguyen, Cuong Huu Nguyen, T. Maisak, Bac Cam Khoa, VR 1627.

FLORA OF VIETNAM

Polypodiaceae s.l.

Dipteris sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of

Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary wet closed mountain broad-leaved and mixed forest on

granites along very steep shady slopes of river canyon at elev. 1750 m. Lithophytic and terrestrial fern on shady mossy cliffs. Rare.

No VH 6329 11.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

VLORA OF VIETNAM

Pteris ensiformis Burm. f.⁣

Khanh Hoa Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 115, 29.10.1990

VLORA OF VIETNAM

Pteris ensiformis Burm. f.⁣

Khanh Hoa Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 115, 29.10.1990

VLORA OF VIETNAM

Pteris ensiformis Burm. f.⁣

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 600, 10.10.1990

VLORA OF VIETNAM

Lindsaea javanensis Blume⁣.

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 810, 15.10.1990

FLORA OF VIETNAM

Polypodiaceae s.l.

Adianthum sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of

Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary closed wet broad-leaved forest on steep granite slopes

at elev. 1200-1250 m. Lithophytic and terrestrial fern on shady very steep slope of river canyon. Common.

No VH 6270 06.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

VLORA OF VIETNAM

Lindsaea javanensis Blume⁣

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 629, 10.10.1990

VLORA OF VIETNAM

Lindsaea orbiculata (Lam.) Mett.⁣

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 808, 15.10.1990

VLORA OF VIETNAM

Lindsaea orbiculata (Lam.) Mett.⁣

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 1099, 20.10.1990

VLORA OF VIETNAM

Lindsaea orbiculata (Lam.) Mett.⁣

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 604, 10.10.1990

VLORA OF VIETNAM

Lindsaea orbiculata (Lam.) Mett.⁣

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 809, 15.10.1990

VLORA OF VIETNAM

Lindsaea orbiculata (Lam.) Mett.⁣

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 630, 10.10.1990

VLORA OF VIETNAM

Adiantum caudatum L.⁣

Quang Ngai Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 455, 07.10.1990

VLORA OF VIETNAM

Adiantum caudatum L.⁣

Quang Ngai Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 455, 07.10.1990

VLORA OF VIETNAM

Adiantum caudatum L.⁣

Quang Ngai Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 455, 07.10.1990

VLORA OF VIETNAM

Adiantum caudatum L.⁣

Khanh Hoa Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 234, 01.10.1990

Lygodium microphyllum (Cav.) R.Br.⁣ ?

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 732, 15.10.1990

VLORA OF VIETNAM

Lygodium microphyllum (Cav.) R.Br.⁣

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 501, 09.10.1990

VLORA OF VIETNAM

Lygodium microphyllum (Cav.) R.Br.⁣?

Quang Ngai Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 456, 07.10.1990

VLORA OF VIETNAM

Lygodium microphyllum (Cav.) R.Br.⁣?

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 1098, 20.10.1990

VLORA OF VIETNAM

Lygodium microphyllum (Cav.) R.Br.⁣?

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 732, 15.10.1990