Herbarium LE - Specimens

Moraceae

Streblus sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Tree to 10 m tall in shady place. Fruits dull green. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4324 (NHC124).

Polypodiaceae s.l.

Leptochilus sp.?

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Creeping lithophytic fern in shady place. Not rare.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4323 (NHC123).

Euphorbiaceae

Antidesma ghaesembilla Gaertn.?

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Treelet or shrub to 3 m tall in rather open place. Flowers greenish. Common.

Anacardiaceae

Allospondias lakonensis (Pierre) Stapf

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Tree about 15 m tall in rather open place. Flowers white. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4320 (NHC121).

Simaroubaceae

Eurycoma longifolia Jack

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Treelet to 4 m tall in rather open place. Flowers purplish. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4319 (NHC120).

Apocynaceae

Tabernaemontana pauciflora Blume

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Treelet to 4 m tall in rather open place. Flowers white. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4318 (NHC119).

Orchidaceae

Tropidia angulosa (Lindl.) Blume

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Terrestrial herb to 0.8 m tall in shady place. Not common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4317 (NHC118).

Gnetaceae

Gnetum latifolium?

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Woody vine to 30 m long, with stem to 20 cm in diameter. Not rare.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4316 (NHC117).

Rubiaceae

Psychotria sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Terrestrial undershrub to 1.5 m tall. Flowers olive greenish. Not rare.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4315 (NHC116).

Selaginellaceae

Selaginella sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Terrestrial herb to 0.4 m tall in rather open place. Locally common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4314 (NHC115).

Hypoxidaceae

Curculigo sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Terrestrial rosulate herb in rather open place. Flowers yellow. Locally common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4313 (NHC114).

Zingiberaceae

Alpinia oblongifolia Hayata

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Terrestrial herb to 1 m tall in rather open place. Flowers white, median tepal with 2 red stripes. Locally common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4312 (NHC113).

Aquifoliaceae

Ilex purpurea?

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Treelet 6-8 m tall in open place. Flowers dirty violet. Occasional.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4311 (NHC112).

Rubiaceae

Randia sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Treelet 5-6 m tall in open place. Flowers green. Occasional.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4310 (NHC111).

Euphorbiaceae

Aporosa dioica (Roxb.) Müll.Arg.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Tree 6-8 m tall in open place. Young fruits light green. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4309 (NHC110).

Clusiaceae

Garcinia sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Tree 6-8 m tall in open place. Flowers yellow. Occasional.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4308 (NHC109).

Apocynaceae

Melodinus suaveolens (Hance) Champ. ex Benth.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Scandent shrub or vine to 5 m long with milky juice in open place. Flowers white. Locally common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4307 (NHC108).

Caprifoliaceae

Lonicera sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Littoral vine to 5 long along sea shore. Flowers white, later pale yellow. Not common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4306 (NHC107).

Verbenaceae

Clerodendron sp.?

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Littoral climbing shrub with scandent shoot to 3 long along sea shore. Flowers white. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4305 (NHC106).

Goodeniaceae

Scaevola taccada (Gaertn.) Roxb.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Littoral shrub to 3 m tall along sea shore. Flowers white. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4304 (NHC105).

Rhizophoraceae?

Bruguiera gymnorhiza (L.) Lam. ex Savigny

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Mangrove scrub in flooded littoral zone along sea shore. Treelet to 3 m tall. Old flowers reddish, fruits dark green. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4303 (NHC104).

Rhizophoraceae

Ceriops tagal (Perr.) C.B.Rob.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Mangrove scrub in flooded littoral zone along sea shore. Treelet or shrub to 2 m tall. Flower buds yellowish with red. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4302 (NHC103).

Convallariaceae

Aspidistra quadrangularis Aver.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Terrestrial rhizomatous herb in shady place. Young fruits light green, quadrangular. Locally common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4301 (NHC102).

= Aspidistra arnautovii var. catbaensis (Tillich) Tillich, Taiwania 59(1): 7 (2014).

Convallariaceae

Peliosanthes sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Terrestrial clustering herb in shady place. Young fruits light green. Locally common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4300 (NHC101).

Orchidaceae

Anoectochilus roxburghii (Wall.) Lindl.

Hanoi Capital Region, Ba Vi National Park, humid mossy primary and secondary evergreen broad-leaved forest on steep mountain slopes composed by solid shale at elevation of 1100-1300 m a.s.l. along trail to peak of 1296 m. Terrestrial or lithophytic herb on shady slope. Leaves drk green with yellowish to pink nerves. Rare.

11 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4297 (NHC099).

Orchidaceae

Epipogium roseum (D.Don) Lindl.

Hanoi Capital Region, Ba Vi National Park, humid mossy primary and secondary evergreen broad-leaved forest on steep mountain slopes composed by solid shale at elevation of 1100-1300 m a.s.l. along trail to peak of 1296 m. Terrestrial tuberiferous achlorophyllous white to light yellowish herb on shady slope. Rare.

11 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4296 (NHC098).

Polypodiaceae s.l.

Microsorus sp.?

Hanoi Capital Region, Ba Vi National Park, humid mossy primary and secondary evergreen broad-leaved forest on steep mountain slopes composed by solid shale at elevation of 1100-1300 m a.s.l. along trail to peak of 1296 m. Creeping epiphytic vine on tall mossy tree. Common.

11 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4295 (NHC097).

Thymellaeaceae

Wikstroemia indica (L.) C.A. Mey.

Hanoi Capital Region, Ba Vi National Park, humid mossy primary and secondary evergreen broad-leaved forest on steep mountain slopes composed by solid shale at elevation of 1100-1300 m a.s.l. along trail to peak of 1296 m. Shrub or small treelet to 1.5 m tall on open rocky steep slope. Fruits orange red to red. Not common.

11 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4294 (NHC096).

Myrsinaceae

Ardisia primulaefolia Gardn. et Champ.

Hanoi Capital Region, Ba Vi National Park, humid mossy primary and secondary evergreen broad-leaved forest on steep mountain slopes composed by solid shale at elevation of 1100-1300 m a.s.l. along trail to peak of 1296 m. Adpressed to ground rosulate herb on mossy vertical cliff near mountain top. Flowers buds white. Not common.

11 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4293 (NHC095).

Podocarpaceae

Podocarpus neriifolius D.Don

Hanoi Capital Region, Ba Vi National Park, humid mossy primary and secondary evergreen broad-leaved forest on steep mountain slopes composed by solid shale at elevation of 1100-1300 m a.s.l. along trail to peak of 1296 m. Tree about 15 m tall on steep shady rocky slope. Occasional.

11 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4292 (NHC094).

Cyperaceae

Carex baviensis Franch.?

Hanoi Capital Region, Ba Vi National Park, humid mossy primary and secondary evergreen broad-leaved forest on steep mountain slopes composed by solid shale at elevation of 1100-1300 m a.s.l. along trail to peak of 1296 m. Terrestrial and lithophytic sedge in open mossy places along forest trail. Common near mountain top.

11 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4291 (NHC093).

Convallariaceae

Aspidistra bogneri Tillich

Hanoi Capital Region, Ba Vi National Park, humid mossy primary and secondary evergreen broad-leaved forest on steep mountain slopes composed by solid shale at elevation of 1100-1300 m a.s.l. along trail to peak of 1296 m. Terrestrial rhizomatous herb with erect rigid leaves to 1.5 m long on steep shady slope. Flowers dark purple-violet to almost black. Not rare.

11 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4290 (NHC092).

Selaginellaceae

Selaginella sp

Hanoi Capital Region, Ba Vi National Park, humid mossy primary and secondary evergreen broad-leaved forest on steep mountain slopes composed by solid shale at elevation of 1100-1300 m a.s.l. along trail to peak of 1296 m. Lithophytic and terrestrial herb with pendulous shoots to 0.8 m long on very steep shady rocky slope near mountain top. Locally common.

11 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4289 (NHC091).