Herbarium LE - Specimens

ASTERACEAE

Vernonia

Ha Giang

Vi Xuyen District; Phong Quang

Municipality; Man Village. Collection

on ridge and small limestone peak just

east of house of village head; very

disturbed, secondary vegetation on lower

slopes and on limestone rocks up to

ridgetop.

22°53'11"N 104°56'43"E 437 - 645 m

22°53'23"N 104°56'48"E

Lianescent, scrambling to 10 m. Stem

14 February 2001

D.K. Harder, P.K. Loc, G.E. Schatz &

L.V. Averyanov 6282

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

green with purple; leaves shiny green

above; bracts green to red purple;

florets yellow; pappus silver.

ASTERACEAE

Vernonia

Lao Cai

Van Ban District; Nam Xe Municipality;

Khau Co Pass. Collection in secondary

vegetation E of pass elevating from 1150

m to 1200 m.

22°02'35"N 103°57'23"E 1150 m

Vine, to 7 m, climbing on shrub,

occasional. Stems dull green with light

brown lenticels; inflorescence bracts

green; florets red-purple; stigma white.

27 February 2001

D.K. Harder, N.T. Hiep, L.V. Averyanov &

N.Q. Hieu 6993

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

Page 2

27 February 2001

D.K. Harder, N.T. Hiep, L.V. Averyanov &

N.Q. Hieu 6993

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

ASTERACEAE

Vernonia

Lao Cai

Van Van District; Liem Phu Municipality;

Khuoi Ngoa Village. Collections near

logging camp [Fokienia (Pou Nhou)] on

road through village; collections from

logging camp and along stream.

21°58'14"N 104°19'51"E 606 m

Occasional, lianescent shrub, to 10 m.

Stem green; bracts light green; in fruit

only; pappus silver-tan.

21 February 2001

D.K. Harder, L.V. Averyanov, P.K. Loc &

G.E. Schatz 6548

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

Araliaceae

Aralia sp.

Thanh Hoa Prov., Thuong Xuan Distr., Bat Mot Municipality, Vin village, Xuan Lien Natural Reserve. Remnants of primary broad-leaved evergreen humid forest on rocky, highly eroded, crystalline limestone at elevation 750-800 m a.s.l. around point 19°59’11.4’’N 104°58’31.8’’E. Treelet 3-4 m tall on steep, rather open slope of rocky stream. Not common.

1 November 2013 CPC 6445

L.Averyanov, N.T.Hiep, N.S.Khang

Araliaceae

Schefflera pauciflora R. Vig.

Quang Binh Prov., Minh Hoa Distr., Thuong Hoa municipality, environs of Mo O O O village, around point 17°40’03.6”N 105°55’08.5”E. Dense broad-leaved closed forest along highly eroded rocky limestone ridge at elevation about 500 m a.s.l.

Climbing lithophytic tree to 12 m tall on rocky top of ridge. Flowers yellowish. Occasional.

CPC 3923 29 July 2011

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Vinh.

Araliaceae

Schefflera leucantha ?

Hoa Binh Prov., Tan Lac Distr., Ngo Luong Municipality, Tram Hai Village, ca. 500 m to S of 20°28’15”N 105°17’21”E.

In heavily logged evergreen broad-leaved forest on foot and slope of limestone mts., at elevation about 750-850 m a.s.l. Lithophyte on vertical cliffs. Shrub, ca. 0.4-0.5 m tall. Flowers brown. Rather common.

CPC 1751 30 March 2011

Coll.: N.Q.Hieu, L.Averyanov, N.T.Hiep, P.K.Loc, P.V.The, N.T.Vinh, T.B.Ngan, N.Tap, D.D.Dao, V.T.Ha.

Araliaceae

Schefflera sp.

Lai Chau Prov., Tam Duong Distr., Ho Thau Community, Ho Thau village, around point 22°23’05’’N 103°36’46’’E.

Secondary and primary evergreen closed forest on very steep mountain slopes composed with granite and quartzite at elevation from 1500 to 1900 m a.s.l. Undershrub 1-2 m tall on very steep rocky shady mountain slope. Flowers white to light yellowish. Occasional.

HAL 10174 29 Nov. 2006

Coll.: Nguyen Tien Hiep, L.Averyanov, Pham Van The.

Araliaceae

Schefflera sp.

Nghe An Prov., Ky Son Distr., Na Ngoi Municipality, eastern slopes of Phu Xai Lai Leng Mountain. Primary very humid broad-leaved forest on steep mountain slopes composed of sandstone and gray shale at elevation about 2300 m a.s.l. around point 19°12’23.3’’N 104°11’33.7’’E. Tree 3-4 m tall (probably to 20 m tall) on steep slope. Flowers yellowish. Common.

24 October 2013 CPC 6225

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, L.M.Tuan, N.A.Trang, L.H.Dan

Araliaceae

Schefflera sp.

Hoa Binh Prov., Lac Son Distr., Tu Do Municipality, Coi Gao village, around point 20°26’07”N 105°17’19”E. Secondary rich forest on steep rocky slopes of limestone mountain at elevation about 600 m a.s.l. Shrub to 3 m tall. Flowers yellowish. Not common.

CPC 1491 24 March 2011

Coll.: N.Q.Hieu, L.Averyanov, N.T.Hiep, P.K.Loc, P.V.The, N.T.Vinh, T.B.Ngan, N.Tap, D.D.Dao, V.T.Ha.

Araliaceae

Schefflera pauciflora R. Vig.

Quang Binh Prov., Minh Hoa Distr., Thuong Hoa municipality, environs of Mo O O O village, around point 17°39’11.6”N 105°54’53”E. Primary broad-leaved closed forest on very steep rocky slope of remnant mountain composed with highly eroded solid crystalline limestone at elevation 400-500 m a.s.l. Epiphytic vine to 15 m long. Not common.

CPC 3672 24 July 2011

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Vinh.

Araliaceae

Schefflera sp.

Vientiane Prov., Vang Vieng Distr., Na Po village, about 12 km to the W of Vang Vieng town, Pa Nang Oua Mt., around point 18°55’44.9”N 102°20’20.1”E. Dry broad-leaved primary and secondary evergreen and semideciduous forest on very steep rocky slopes and on vertical cliffs of remnant highly eroded rocky limestone mountains composed by solid crystalline limestone at elevation 700-1000 m. Treelet on rocky slope near mountain top. Occasional.

LA-VN 522 17 March 2013

Coll.: N.T.Hiep, L.Averyanov, K.Chantthavongsa, N.S.Khang, P.V.The, S.Lorphengsy

Araliaceae

Schefflera sp.

Hoa Binh Prov., Lac Son Distr., Tu Do Municipality, Cho village, around point 20°25’21”N 105°18’01”E. Secondary forest along stream in alluvial valley between limestone hills at elevation about 230 m a.s.l. Scandent shrub to 4 m tall. Fruits green. Occasional.

CPC 1427 24 March 2011

Coll.: N.Q.Hieu, L.Averyanov, N.T.Hiep, P.K.Loc, P.V.The, N.T.Vinh, T.B.Ngan, N.Tap, D.D.Dao, V.T.Ha.

Fam. Asteraceae

Vernonia sp. ?

N. Vietnam, prov. Cao Bang, distr. Bao Lac, municipality Yen Lac, vicinities of Yen Lac village (N 22ш44' E 105ш50'), about 44 km to NWW

from Cao Bang town. Primary open mixed semididuous mossy forest on steep mt slopes near tops of limestone remnant karst ridge at

1200-1300 m alt. Herb vine up to 8 n lg in open place. Not common.

No CBL 301 15 Nov. 1998

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6300-98)

Collectors: Averyanov L., N.T.Hiep, P.K.Loc, N.X.Tam.

Plants of Phong Nha - Ke Bang National Park

Fam. Asteraceae

Vernonia sp.

Vietnam, Quang Binh Prov., Bo Trach Distr., Hung Trach Municipality, Khe Me locality, around point 17є27'51''N, 106є22'46''E at elev. 900-1000 m a.s.l. Slightly logged primary closed broad-leaved evergreen forest on slopes of mountains composed with shale and sandstone.

Hemi-woody vine 15-20 m long. Flowers light violet, fragrant. Not common.

No HAL 6247 29 January 2005. Photo.

Collectors: L.Averyanov, P.K.Loc, P.V.The, A.Averyanova, N.T.Vinh, N.Q.Vinh, N.T.Binh.

Fam. Asteraceae

Cynareae sp.

S. Vietnam, Prov. Kon Tum, Distr. Dak Gley, about 3-4 km to the W of Dak Gley town. Secondary mixed (with Pinus khesya) dry forest with bamboos rich in primary elements at 650-800. Herb up to 0.5 m high. Flowers violet. Rare.

No VH 2041 26.11.1995

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 5094-93)

Participants: L. Averyanov, N.T. Ban, A. Budantzev, L. Budantzev, N.T. Hiep, D.D. Huyen, P.K. Loc, G. Yakovlev

Cirsium lineare (Thunb.) Schult-Bip.

S. Vietnam, Prov. Lam Dong, distr. Lac Duong, municipalite Da Chay, vicinities Klong Lanh village, 32 km to NNE from Dalat city (N 12°

08', E 108° 39'). Grassy-sedges swampy Pinus kesiya forest on high terrace of river valley at the base of hills at 1450 m alt. in 6-7 km

from Bi Dup main peak to the W. Herb up to 2 m hg. Flowers lilac. Rare.

No VH 3972 14.04.1997

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 5803-96)

Collectors: L. Averyanov, N.Q. Binh

Fam. Asteraceae

Cirsium sp.

N. Vietnam, prov. Cao Bang, distr. Bao Lac, municipality Ca Thanh, about 3 km to SW from Yen Lac village (N 22ш44' E 105ш50'), about 42

km to NWW from Cao Bang town. Secondary open shrubs on alluvial foothills of limestone remnant ridge at 1300 m alt. Herb up to 1.5 m

hg. Not rare.

No CBL 387 18 Nov. 1998

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6300-98)

Collectors: Averyanov L., P.K.Loc, N.X.Tam.

Fam. Asteraceae

Cirsium sp.

N. Vietnam, prov. Cao Bang, distr. Nguyen Binh, municipality Ca Thanh, vicinities Ca Lu village, about 7-8 km to SE from Yen Lac village (N

22ш44' E 105ш50'), about 38 km to NWW from Cao Bang town. Open secondary grassland slopes on alluvial walley and foothills of karst

remnant limestone ridge at 1100-1150 m alt. herb up to 1.5 m hg. Flowers pink-violet. Not common.

No CBL 565 21 Nov. 1998

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6300-98)

Collectors: Averyanov L., P.K.Loc, N.X.Tam.

Fam. Asteraceae

Cirsium sp.

N. Vietnam, prov. Cao Bang, distr. Nguyen Binh, municipality Nguyen Binh (N 22ш37' E 105ш52'), about 41 km to W from Cao Bang town.

Elevation from Le A pass to Na Ni mt. peak at 1400-1900 m alt. Open wet steep slopes on soils developed on granite and shale rocks.

Herb up to 1.5 m hg. Flowers greyish-white. Not rare.

No CBL 019 11-12 Nov. 1998

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6300-98)

Collectors: Averyanov L., N.Q.Binh, N.T.Hiep, P.K.Loc, N.X.Tam.

Asteraceae

Hemistepta lyrata Bunge.

Ha Nam Prov.

LX-VN 042-a, 29.03.1984

Det. Sennikov A.

Asteraceae

Hemistepta lyrata Bunge.

Lang Son Prov.

3233, 08.04.1961

Det. Sennikov A.

Asteraceae

Hemistepta lyrata Bunge.

Ninh Binh Prov.

Avrorin N., Avrorin 10231

10.05.1963

Det. Sennikov A.

Fam. Asteraceae

Hemistepta lyrata

N. Vietnam, prov. Cao Bang, distr. Nguyen Binh, municipality Thai Hoc, about 5-6 km to ENE of Nguyen Binh town (N 22ш39' E 105ш57').

Secondary open shrubs on steep slopes of karst remnant limestone mesae at 500-550 m elev. Terrestrial herb in wet place along road. Flowers

pale lilac-violet. Common. Commonest weed in the area.

No CBL 1624 21 April 1999

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6300-98)

Collectors: P.K.Loc, P.H.Hoang, Averyanov L.

Asteraceae

Eupatorium odoratum L.⁣

Khanh Hoa Prov.

Averyanov L., Kudryavtzeva E.

Averyanov 1291, 17.04.1987

Asteraceae

Eupatorium odoratum L.⁣

Kien Giang Prov.

Averyanov L., Kudryavtzeva E.

Averyanov 1157, 12.04.1987

Asteraceae

Eupatorium odoratum L.⁣

Ba Ria-Vung Tau Prov.

Averyanov L., Kudryavtzeva E.

Averyanov 756, 03.04.1987

Asteraceae

Eupatorium odoratum L.⁣

Ba Ria-Vung Tau Prov.

Averyanov L., Kudryavtzeva E.

Averyanov 709, 02.04.1987

Fam. Asteraceae

Eupatorium odoratum

Quang Nam Prov.

Averyanov L., Kudryavtzeva E.

Averyanov 388, 28.03.1987

Fam. Asteraceae

Eupatorium odoratum

Khanh Hoa Prov.

Averyanov L., Kudryavtzeva E.

Averyanov 1291, 17.04.1987

Fam. Asteraceae

Eupatorium odoratum

Hai Phong Prov.

LX-VN 3368, 23.12.1986

Fam. Asteraceae

Eupatorium odoratum

N. Vietnam, prov. Cao Bang, distr. Thach An, municipality Thuy Hung, vicinities of Na Lit village (N 22ш30' E 106ш27'), about 27 km to SE

from Cao Bang town. Secondary closed evergreen broadleaved forest between remnant limestone ridges on alluvial valley along stream at

about 100 m alt. Terrestrial erect and climbing herb up to 3 m hg in open place. Flowers dull violet. Commonest very aggressive weed all

over the area.

No CBL 870 04 Dec. 1998

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6300-98)

Collectors: Averyanov L., N.Q.Binh, P.K.Loc, N.X.Tam.

Fam. Asteraceae

Eupatorium sp.

N. Vietnam, prov. Cao Bang, distr. Ha Lang, municipality An Lac, vicinities of Phia Deng village (N 22ш43' E 106ш36'), about 36 km to E

from Cao Bang town. Secondary broadleaved remnant forest along stream on alluvial valley between limestone ridges at 500 m alt. Epilite herb

or shrublet up to 1 m hg on open stream limestone rocks. Flowers pale violet. Not common, but sporadically abundant.

No CBL 779 29 Nov. 1998

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6300-98)

Collectors: Averyanov L., P.K. Loc.

Asteraceae

Chromolaena odorata (L.) R.M. King & H. Rob.

Lang Son Prov.

Daniel Atha, Tien Hiep Nguyen et al.

NY-HN 066, 09.12.2005

Det. Tran T.Phuong Anh at 09.12.2005

Polypodiaceae s.l.

Davallia sp.

Xiangkhouang Prov., Peak Distr., Khoang village, Pa Khao mt., around point 19°33'02.2'N 103°39'56.3'E. Primary and secondary broad-leaved evergreen dry forest on solid rocky highly eroded limestone at elev. 1200-1450 m a.s.l. Creeping epiphyte on tall mossy tree. Common.

LA-VN 1001 3 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

FLORA OF LAOS

Polypodiaceae s.l.

Microlepia sp.

Xiangkhouang Prov., Peak Distr., Oran village, around point 19°38’53”N 103°23’12”E. Primary and secondary broad-leaved evergreen dry forest on shale sandstone slopes and along ridge edge at elev. about 1750-1850 m a.s.l. Terrestrial fern in shady places. Common.

LA-VN 873 2 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

FLORA OF LAOS

Polypodiaceae s.l.

Microlepia sp.?

Xiangkhouang Prov., Peak Distr., Oran village, around point 19°38’53”N 103°23’12”E. Primary and secondary broad-leaved evergreen dry forest on shale sandstone slopes and along ridge edge at elev. about 1750-1850 m a.s.l. Terrestrial fern in shady wet place. Common.

LA-VN 892 2 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

Polypodiaceae s.l.

Lematosorus sp.??

Cao Bang Prov., Bao Lac Distr., Hong An municipality, Mi Lung village. Primary broad-leaved and mixed humid evergreen forest (with Podocarpus, Pinus, Fokienia and Tsuga) on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation about 1500-1550 m a.s.l. around point 22°49’15.4’’N 105°49’53’’E. Creeping epiphyte on old mossy tree on rocky mossy mountain slope near mountain top. Occasional.

21 November 2014 CPC 7580

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz

Polypodiaceae s.l.

Lepisorus sp.

Cao Bang Prov., Bao Lac Distr., Dinh Phung municipality, Ban Meo village. Secondary broad-leaved evergreen forest on steep stream slope composed by shale stratified limestone at elevation 800-900 m a.s.l. around point 22°44’32.0’’N 105°46’29.1’’E. Lithophytic fern on large mossy stream boulders. Not common.

24 November 2014 CPC 7610

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz

Polypodiaceae

Gen. sp.

Ha Giang Prov., Vi Xuyen Distr., Tung Ba municipality, Khau Ca NR. Primary broad-leaved humid evergreen forest on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation 700-1150 m a.s.l. around point 22°50’39.4’’N 105°07’50.3’’E. Terrestrial herb 0.5-1-1,5 m in tall, common.

08 May 2015 CPC 7772

Coll.: N.Q. Hieu, N.T.Hiep, T.B. Ngan, C.Q. Ngan, D.K. Harder, C. P. Tuy, G.C.Pao

Polypodiaceae s.l.

Lepisorus sp.

Xiangkhouang Prov., Kham Distr., Huad village, Phou tat Vinh mt., around point 19°32'31.6'N 103°39'40.4'E. Primary and secondary broad-leaved evergreen and semideciduous dry forest on very steep rocky slopes of limestone highly eroded mountains composed by solid marble like limestone at elev. 1200-1400 m a.s.l. Creeping epiphyte on tall old mossy tree along ridge edge. Occasional.

LA-VN 1094 4 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

FLORA OF LAOS

Polypodiaceae s.l.

Lepisorus sp.?

Xiangkhouang Prov., Peak Distr., Oran village, around point 19°38’53”N 103°23’12”E. Primary and secondary broad-leaved evergreen dry forest on shale sandstone slopes and along ridge edge at elev. about 1750-1850 m a.s.l. Creeping epiphytic vine 3-5 m long in shady place. Common.

LA-VN 888 2 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

FLORA OF LAOS

Polypodiaceae s.l.

Davallia sp.

Houphan Prov., Xon Distr., Nong Hua Pa Noi village, around point 20°13'02.1''N 103°24'23.3''E. Primary and secondary broad-leaved evergreen forest on very steep rocky mountain slopes composed by shaly limestone at elev. 1100-1200 m a.s.l. Epiphyte on tall trees. Flowers buds white-yellowish. Locally very common.

LA-VN 1230 11 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

FLORA OF LAOS

Polypodiaceae s.l.

Dryopteris sp.?

Houphan Prov., Hem Distr., Na Puok village, Pu Tham Nhon mountain around point 20°10'00.0''N 103°24'37.0''E. Primary and secondary broad-leaved evergreen dry forest on very steep rocky mountain slopes composed by solid marble-like highly eroded limestone at elev. 1050-1150 m a.s.l. Terrestrial fern in shady slope near mountain base. Very common.

LA-VN 1341 11 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

DRYOPTERIDACEAE

Diplazium

Son La

Moc Chau District; Van Ho Municipality,

Hua Tat Village. Collections on

limestone ridge from degraded forests to

cultivation on lower slopes to primary

broadleaved, closed wet forest along

limestone ridge and slopes near tops.

20°46'13"N 104°46'12"E 1100 - 1300 m

Lithophytic fern on vertical shady

cliffs. Not rare.

7 March 2001

D.K. Harder, P.K. Loc, L.V. Averyanov &

N.X. Tam 7441

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

FLORA OF VIETNAM

Polypodiaceae s.l.

Drynaria sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) along Dak Tour

river on N macroslope of Chu Yang Sinh mt system. Primary broad-leaved mountain evergreen forest on granites along river slopes at elev.

500-550 m. Lithophytic and epiphytic fern with fronds up to 1.5 m lg in open places along river. Very common.

No VH 5963 03.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

FLORA OF VIETNAM

Polypodiaceae s.l.

Drynaria sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) along Dak Tour

river on N macroslope of Chu Yang Sinh mt system. Primary broad-leaved mountain evergreen forest on granites along river slopes at elev.

500-550 m. Lithophytic and epiphytic fern with fronds up to 1.5 m lg in open places along river. Very common.

No VH 5963 03.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

FLORA OF VIETNAM

Polypodiaceae s.l.

Drynaria sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) along Dak Tour

river on N macroslope of Chu Yang Sinh mt system. Primary broad-leaved mountain evergreen forest on granites along river slopes at elev.

500-550 m. Lithophytic and epiphytic fern with fronds up to 1.5 m lg in open places along river. Very common.

No VH 5963 03.05.2000

International Botanical Expedition of the U.S.A. National Geographic

Society (grant No 6383-98)

___________________________________________________________

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh