Herbarium LE - Specimens

All photos © - L. Averyanov

All photos © - L. Averyanov

FLORA OF LAOS

All photos © - L. Averyanov

FLORA OF LAOS

All photos © - L. Averyanov

FLORA OF LAOS

FLORA OF LAOS

All photos © - L. Averyanov & Nguyen Sinh Khang

FLORA OF LAOS

FLORA OF LAOS

FLORA OF LAOS

All photos © - L. Averyanov

FLORA OF LAOS

FLORA OF LAOS

FLORA OF LAOS

All photos © - L. Averyanov

FLORA OF LAOS

All photos © - L. Averyanov

FLORA OF LAOS

All photos © - L. Averyanov, N.S. Khang

FLORA OF LAOS Hin Nam No Nature Protected Area Asclepiadaceae Hoya aphylla Aver. & K.S. Nguyen sp. nov. Laos, Khammoune prov., Boualapha distr., Ban Dou village, around Khuay cave, at 17°30'35.1"N, 105°51'28.7"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 250-390 m a.s.l. Erect terrestrial and lithophytic herb, later herbaceous or semi-woody vine on rocky slope. Flowers white to light yellowish, corona pale brownish-pink. Not common. 10 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 890. d-EXSICCATES OF VIETNAMESE FLORA 0318 / AL 890 © L.Averyanov & K.S. Nguyen

FLORA OF LAOS Hin Nam No Nature Protected Area Melanthiaceae Chionographis actinomorpha Aver. & N. Tanaka Laos, Khammouane prov., Boualapha distr., Nong Seng village, southern sandstone slopes at summit of Phou Chuang (Poujeuang) Mt. around point 17°35'00.3"N, 105°47'35.6"E. Primary evergreen mixed and coniferous dense wind-formed forest at elev. 1000-1400 m a.s.l. Terrestrial and lithophytic rosulate herb in open mossy place on mountain top. Flowers white. Occasional. 4 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 619.

FLORA OF LAOS Hin Nam No Nature Protected Area Euphorbiaceae Euphorbia thymifolia L. Laos, Khammoune prov., Boualapha distr., Vang Maner village, Palai mountain area, around point 17°30'13.6"N, 105°49'23.9"E, 520 m a.s.l. Primary dry evergreen and semideciduous broad-leaved forest on sandy alluvial terrace and slopes of river valley between karstic limestone hills at 180-220 m a.s.l. Prostrate herb on open alluvial sandy deposits along river. Flowers white. Occasional. 8 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 812.

FLORA OF LAOS Hin Nam No Nature Protected Area Euphorbiaceae Euphorbia thymifolia L. Laos, Khammoune prov., Boualapha distr., Vang Maner village, Palai mountain area, around point 17°30'13.6"N, 105°49'23.9"E, 520 m a.s.l. Primary dry evergreen and semideciduous broad-leaved forest on sandy alluvial terrace and slopes of river valley between karstic limestone hills at 180-220 m a.s.l. Prostrate herb on open alluvial sandy deposits along river. Flowers white. Occasional. 8 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 812.

FLORA OF LAOS Convallariaceae Aspidistra khangii Aver, el Tillich Herbarium specimen prepared 8 September 2016 by L. Averyanov, LA-VN 104a from cultivated plant collected in Laos, Attapeu province, Saysettha district, Dong Am Pham National Protected Area, to the east of Paosamphanmixay village, around point 14°48'24.8"N, 107°24'23.2" E, primary broad-leaved evergreen forest at ca 900 m a.s.l., 28 Feb 2013, L. Averyanov et al., LA-VN 104 (plant used for preparation of the species type voucher herbarium specimens.

FLORA OF LAOS Convallariaceae Aspidistra khangii Aver. et Tillich, sp.nov. Type Attapeu Prov., Saysettha Distr., Dong Am Pham National Protected Area, to the E of Paosamphanmixay village, around point 14°48’24.8”N 107°24’23.2”E. Primary broad-leaved evergreen lowland dry forest on steep shaly hill slopes at elevation 700-920 m a.s.l. Terrestrial herb with ascending stem and erect leaves to 1.2 m long along mountain ridge in shady places. Very common to abundant. Old flowers dull brownish. LA-VN 104 28 February 2013 Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, P.V.The, K.Inkhavilay, S.Lorphengsy

FLORA OF LAOS Convallariaceae Aspidistra khangii Aver. et Tillich, sp.nov. Type Attapeu Prov., Saysettha Distr., Dong Am Pham National Protected Area, to the E of Paosamphanmixay village, around point 14°48’24.8”N 107°24’23.2”E. Primary broad-leaved evergreen lowland dry forest on steep shaly hill slopes at elevation 700-920 m a.s.l. Terrestrial herb with ascending stem and erect leaves to 1.2 m long along mountain ridge in shady places. Very common to abundant. Old flowers dull brownish. LA-VN 104 28 February 2013 Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, P.V.The, K.Inkhavilay, S.Lorphengsy

FLORA OF LAOS Hin Nam No Nature Protected Area Convallariaceae Aspidistra medusa Aver., K.S. Nguyen & Tillich Laos, Khammoune prov., Boualapha distr., Vang Maner village, Palai mountain area, around point 17°30'13.6"N, 105°49'23.9"E, 520 m a.s.l. Primary dry evergreen and semideciduous broad-leaved forest on sandy alluvial terrace and slopes of river valley between karstic limestone hills at 180-220 m a.s.l. Terrestrial herb with short underground rhizome on shady slope along river. Leaves uniform grassy green, flowers entirely white, stigma near margin sometime with light violet tint, fruits dark olive-green. Locally common. Flower buds (to be said) edible after boiling and cooking with local sauce. 8 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 813. d-EXSICCATES OF VIETNAMESE FLORA 0321 / AL 813 © L.Averyanov & K.S. Nguyen.

FLORA OF LAOS Hin Nam No Nature Protected Area Convallariaceae Aspidistra medusa Aver., K.S. Nguyen & Tillich Laos, Khammoune prov., Boualapha distr., Vang Maner village, Palai mountain area, around point 17°30'13.6"N, 105°49'23.9"E, 520 m a.s.l. Primary dry evergreen and semideciduous broad-leaved forest on sandy alluvial terrace and slopes of river valley between karstic limestone hills at 180-220 m a.s.l. Terrestrial herb with short underground rhizome on shady slope along river. Leaves uniform grassy green, flowers entirely white, stigma near margin sometime with light violet tint, fruits dark olive-green. Locally common. Flower buds (to be said) edible after boiling and cooking with local sauce. 8 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 813. d-EXSICCATES OF VIETNAMESE FLORA 0321 / AL 813 © L.Averyanov & K.S. Nguyen.

FLORA OF LAOS Hin Nam No Nature Protected Area Araliaceae Trevesia palmata (Roxb. ex Lindl.) Vis. Laos, Khammoune prov., Boualapha distr., Vang Maner village, around point 17°29'39.3"N, 105°50'24.6"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 350-430 m a.s.l. Treelet to 6 m tall on shady rocky slope. Flowers yellow, fruits green. Common. 9 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 882.

FLORA OF LAOS Hin Nam No Nature Protected Area Araliaceae Schefflera sp. Laos, Khammoune prov., Boualapha distr., Nong Seng village, Chanh Daeng mountain, around point 17°31'11.1"N, 105°46'41.4"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 400-600 m a.s.l. Treelet 2-3 m tall on open rocky slope near mountain top. Fruits red, later black. Common. 11 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 1032.

FLORA OF LAOS Hin Nam No Nature Protected Area Araliaceae Schefflera elliptica (Blume) Harms Laos, Khammouane prov., Boualapha distr., Nong Seng village, southern sandstone slopes of Phou Chuang (Poujeuang) Mt. around point 17°34'13.3"N 105°47'57.5"E. Primary evergreen broad-leaved dry forest at elev. 700-1000 m a.s.l. Treelet or shrub 3-4 m tall on open rocks. Common. 3 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 503.

FLORA OF LAOS Hin Nam No Nature Protected Area Araliaceae Trevesia palmata (Roxb. ex Lindl.) Vis. Laos, Khammouane prov., Boualapha distr., Nong Seng village, southern sandstone slopes of Phou Chuang (Poujeuang) Mt. around point 17°34'13.3"N 105°47'57.5"E. Primary evergreen broad-leaved dry forest at elev. 700-1000 m a.s.l. Treelet to 6 m tall on steep rocky slope. Common. 3 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 561.

FLORA OF LAOS Hin Nam No Nature Protected Area Araliaceae Schefflera wallichiana (Wight & Arn.) Harms ? Laos, Khammoune prov., Boualapha distr., Nong Seng village, Chanh Daeng mountain, around point 17°31'11.1"N, 105°46'41.4"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 400-600 m a.s.l. Treel to 10 m tall on open rocks near mountain top. Flowers yellow. Common. 11 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 1037.

FLORA OF LAOS Hin Nam No Nature Protected Area Araceae Arisaema roxburghii Kunth Laos, Khammoune prov., Boualapha distr., Vang Maner village, Palai mountain area, around point 17°30'13.6"N, 105°49'23.9"E, 520 m a.s.l. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 300-500 m a.s.l. Terrestrial herb among shady rocks. Spathe white mottled with gray, apex green. Occasional. 8 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 770.

FLORA OF LAOS Hin Nam No Nature Protected Area Araceae Homalomena occulta (Lour.) Schott Laos, Khammoune prov., Boualapha distr., Vang Maner village, Palai mountain area, around point 17°30'13.6"N, 105°49'23.9"E, 520 m a.s.l. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 300-500 m a.s.l. Terrestrial herb on open rocks. Spathe green. Occasional. 8 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 759.

FLORA OF LAOS Hin Nam No Nature Protected Area Araceae Scindapsus sp.? Laos, Khammoune prov., Boualapha distr., Vang Maner village, Palai mountain area, around point 17°30'13.6"N, 105°49'23.9"E, 520 m a.s.l. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 300-500 m a.s.l. Creeping epiphytic vine in shady place. Not common. 8 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 752.

FLORA OF LAOS Hin Nam No Nature Protected Area Araceae Aglaonema costatum N.E. Br. Laos, Khammoune prov., Boualapha distr., Vang Maner village, Palai mountain area, around point 17°30'13.6"N, 105°49'23.9"E, 520 m a.s.l. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 300-500 m a.s.l. Terrestrial creeping herb in shady place, Inflorescence white. Very common. 8 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 750.

FLORA OF LAOS Hin Nam No Nature Protected Area Araceae Amorphophallus carneus Ridl. Laos, Khammoune prov., Boualapha distr., Ban Dou village, around Khuay cave, at 17°30'35.1"N, 105°51'28.7"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 250-390 m a.s.l. Terrestrial ephemeroid herb on shady rocky slope. Flowers and spathe white. Common. 10 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 892.

FLORA OF LAOS Hin Nam No Nature Protected Area Araceae Hapaline benthamiana Schott Laos, Khammoune prov., Boualapha distr., Vang Maner village, Palai mountain area, around point 17°30'13.6"N, 105°49'23.9"E, 520 m a.s.l. Primary dry evergreen and semideciduous broad-leaved forest on sandy alluvial terraces and slopes of river valley between karstic limestone hills at 180-220 m a.s.l. Terrestrial tuberiferous herb on flattened sandy terrace and slopes along river valley, commonly in rather open grassy places. Spathe and inflorescence almost white, flowers very fragrant. Locally very common. 8 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 814.

FLORA OF LAOS Hin Nam No Nature Protected Area Araceae Amorphophallus pilosus Hett. ? Laos, Khammoune prov., Boualapha distr., Vang Maner village, around point 17°29'39.3"N, 105°50'24.6"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 350-430 m a.s.l. Ephemeroid tuberiferous herb in open place. Inflorescence to 1.4 m tall. Spathe pale greenish, dirty purple inside near base, flowers and axis extension yellow. Not common. 9 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 830.

FLORA OF LAOS Hin Nam No Nature Protected Area Araceae Arisaema roxburghii Kunth Laos, Khammouane prov., Boualapha distr., Nong Seng village, southern sandstone slopes at summit of Phou Chuang (Poujeuang) Mt. around point 17°35'18.1"N, 105°47'47.5"E. Primary evergreen mixed and coniferous dense wind-formed forest at elev. 1000-1400 m a.s.l. Terrestrial herb in shady humid place. Locally common. 5 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 650a.

FLORA OF LAOS Hin Nam No Nature Protected Area Araceae Hapaline benthamiana Schott Laos, Khammoune prov., Boualapha distr., Nong Seng village, Vang Phae Lay mountain, around point 17°31'02.7"N, 105°47'17.2"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 300-530 m a.s.l. Terrestrial and lithophytic tuberiferous herb on rocky slope at hill base. Spathe and flowers white, fragrant. Very common. 11 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 942.

FLORA OF LAOS Hin Nam No Nature Protected Area Araceae Alocasia sp. Laos, Khammoune prov., Boualapha distr., Vang Maner village, around point 17°29'39.3"N, 105°50'24.6"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 350-430 m a.s.l. Terrestrial rhizomatous herb among shady rocks on shady rocky slope. Spathe dark glossy green. Not rare. 9 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 851.

FLORA OF LAOS Hin Nam No Nature Protected Area Araceae Arisaema roxburghii Kunth Laos, Khammoune prov., Boualapha distr., Nong Seng village, Chanh Daeng mountain, around point 17°31'11.1"N, 105°46'41.4"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 400-600 m a.s.l. Terrestrial tuberiferous herb among shady rocks. Spathe gray-white mottled outside, white inside, apex green, apex green. Occasional. 11 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 992.

FLORA OF LAOS Hin Nam No Nature Protected Area Araceae Aglaonema sp. Laos, Khammoune prov., Boualapha distr., Ban Dou village, around Khuay cave, at 17°30'35.1"N, 105°51'28.7"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 250-390 m a.s.l. Erect terrestrial herb to 0.5 m tall on shady rocky slope. Common. 10 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 902.

FLORA OF LAOS Hin Nam No Nature Protected Area Araceae Alocasia sp. Laos, Khammouane prov., Boualapha distr., Nong Seng village, southern sandstone slopes of Phou Chuang (Poujeuang) Mt. around point 17°34'13.3"N 105°47'57.5"E. Primary evergreen broad-leaved dry forest at elev. 700-1000 m a.s.l. Terrestrial herb in rocky shady place. Spathe dark green. Occasional. 3 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 518.

FLORA OF LAOS Hin Nam No Nature Protected Area Araceae Aglaonema costatum N.E. Br. Laos, Khammouane prov., Boualapha distr., Nong Seng village, southern sandstone slopes of Phou Chuang (Poujeuang) Mt. around point 17°34'13.3"N 105°47'57.5"E. Primary evergreen broad-leaved dry forest at elev. 700-1000 m a.s.l. Terrestrial creeping herb in rocky shady place. Spathe grassy green to almost white. Common. 3 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 519.

FLORA OF LAOS Hin Nam No Nature Protected Area Araceae Aglaonema ovatum Engl. Laos, Khammouane prov., Boualapha distr., Nong Seng village, southern sandstone slopes of Phou Chuang (Poujeuang) Mt. around point 17°34'13.3"N 105°47'57.5"E. Primary evergreen broad-leaved dry forest at elev. 700-1000 m a.s.l. Terrestrial herb to 1 m tall in shady place. Spathe light green. Very common. 3 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 526.

FLORA OF LAOS Hin Nam No Nature Protected Area Aquifoliaceae Ilex memecylifolia Champ. ex Benth. Laos, Khammouane prov., Boualapha distr., Nong Seng village, southern sandstone slopes at summit of Phou Chuang (Poujeuang) Mt. around point 17°35'18.1"N, 105°47'47.5"E. Primary evergreen mixed and coniferous dense wind-formed forest at elev. 1000-1400 m a.s.l. Treelet or shrub 2-3 m tall on mountain top. Flowers buds light violet. Locally common. 5 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 665a.

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KOMAROV BOTANICAL INSTITUTE

FLORA OF LAOS Hin Nam No Nature Protected Area Convallariaceae Tupistra siphonantha N.Tanaka, Vislobokov & D.P.Hannon Laos, Khammoune prov., Boualapha distr., Nong Seng village, Chanh Daeng mountain, around point 17°31'11.1"N, 105°46'41.4"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 400-600 m a.s.l. Lithophytic herb with short thick plagiotropic rhizome on exposed rocks near mountain top. Very common at elevation 450-600 m a.s.l. 11 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 983. Flowered 05 Nov 2018. Flowers white, photos, L. Averyanov

FLORA OF LAOS Convallariaceae Tupistra muricata (Gagnep.) N.Tanaka Xiangkhouang Prov., Kham Distr., Huad village, Phou tat Vinh mt., around point 19°32'31.6'N 103°39'40.4'E. Primary and secondary broad-leaved evergreen and semideciduous dry forest on very steep rocky slopes of limestone highly eroded mountains composed by solid marble like limestone at elev. 1200-1400 m a.s.l. Terrestrial herb on mountain slope. Locally very common. Fruits almost black. LA-VN 1041 4 April 2015. Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham. Living specimen: LA-VN 1041 / garden no 15171a, flowered 6 Dec. 2017 Coll. in garden in 6 Dec. 2017, by L. Averyanov, photos.