Herbarium LE - Specimens

FLORA OF VIETNAM

FLORA OF LAOS

FLORA OF VIETNAM

FLORA OF VIETNAM

FLORA OF VIETNAM Convallariaceae Aspidistra lubae Aver. et Tillich var. lubae Type Hoa Binh Prov., Lac Son Distr., Tu Do Municipality, Mon village, around point 20°25’29”N 105°19’36”E. Primary broadleaved forest on steep rocky slopes of remnant mountain composed with solid crystalline highly eroded limestone at elevation about 500-700 m a.s.l. Terrestrial herb in shady place. Not rare. CPC 1566a/sp.1 25 March 2011. Coll.: L.Averyanov, T.Maisak, L.Osinovetz et al. Fl. under cultivation at December 2012. Perianth yellowish with dirty violet marking, dark dirty violet inside, stigma white to yellowish, filaments white, anthers light yellowish. Photos. Averyanov L.

FLORA OF VIETNAM Aspidistra heterocarpa Aver., Tillich & V.T. Pham var. echinata T.A.Le, Aver. & Tillich Thua Thien - Hue province, Nam Dong district, Thuong Quang commune, around point 16°07’28.5”N 107°35’52.9”E, primary and secondary evergreen broad-leaved lowland forest at elevation 500-550 m a.s.l., terrestrial herb on shady reclined slope, not common, 10 November 2017, Le Tuan Anh, Nguyen Thi Lien Thuong, LTA101. d-EXSICCATES OF VIETNAMESE FLORA 0295/LTA101 © Le Tuan Anh & L.Averyanov.

FLORA OF VIETNAM Convallariaceae Aspidistra coccigera Aver. et H.-J. Tillich sp.nov. Vietnam, Quang Binh Prov., Minh Hoa Distr., Dan Hoa municipality, Vietnam-Laotian border, around Cha Lo border entrance, point 17º40’52’’N 105º46’00’’E. Broad-leaved primary forest on very steep slopes and along shale ridge at elevation 500-750 m a.s.l. Terrestrial herb on very steep rocky shady slope. Flowers white, tips of petals dark violet-black, stigma brightly red. Locally common. HAL 11817 22 April 2008 photo Coll.: L.Averyanov, P.K.Loc, N.T.Vinh, N.S.Khang

Aspidistra atrata Aver., Tillich & Q.H.Bui⁣

FLORA OF LAOS Convallariaceae Tupistra albiflora K.Larsen Vientiane Prov., Vang Vieng Distr., Phol Xai village, about 8 km to the W of Vang Vieng town, Phar Poon Mt., around point 18°56’11.5”N 102°20’05.0”E. Dry broad-leaved primary and secondary evergreen forest on very steep rocky slopes and on vertical cliffs of remnant highly eroded rocky limestone mountains composed by solid crystalline limestone at elevation 300-900 m. Terrestrial herb with erect stem to 0.3 m tall in shady place. Very common at elevation 700-800 m. LA-VN 431 15 March 2013 Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, P.V.The, S.Lorphengsy. (Fl. 14 Nov. 2016, LA-VN 431 / garden no 13009. Flowers white, tepal apices violet or lilac, anthers pale light yellowish, inflorescence rachis pinkish. Flowers with strange strong smell like chlorophorm). Flowered 6 Dec. 2017 LA-VN 431 / garden no 13009 Photos L. Averyanov.

FLORA OF VIETNAM Convallariaceae Tupistra wattii Dien Bien Prov., Tua Chua Distr., Sin Chai Municipality, around point 22°03’38”N 103°19’56”E. Primary humid evergreen broad-leaved forest on very steep rocky slopes and on tops of remnant mountain composed with highly eroded marble-like solid crystalline limestone at elevation 1350-1500 m a.s.l. Terrestrial and lithophytic herb to 0.5 m tall on shady rocky slope. Flowers light green. Very common. CPC 900 13 December 2010 Coll.: L.Averyanov, P.K.Loc, P.V.The, N.T.Vinh.

FLORA OF VIETNAM. Fam. Convallariaceae. Tupistra wattii (C.B. Clarke) Hook.f. ???. Northern Vietnam, Lao Cai Prov., Van Ban Distr., to the west of Nam. Xay village (around point 21°58'09"N, 104°00'55"E). Primary. broad-leaved evergreen montane forest with bamboo along tops of. granite/quartzite ridge at elev. 1800 m a.s.l. Creeping terrestrial. herb on open wet slope. Flowers greenish to dull orange. Not common. No HAL 2135   07 March 2002. . Collectors: L.Averyanov, P.K.Loc, D.T.Doan (FFI Project).

FLORA OF VIETNAM Fam. Convallariaceae Tupistra sp. N. Vietnam, Prov. Cao Bang, Distr. Tra Linh, Subdistr. Quoc Toan, evergreen forest on slopes of remnant limestone mountains near Thang Heng lake in environs of Thang Hen and Lung Tao villages at 600-700 m alt. Herb with leaves up to 1.5 m lg. Flowers black. Very common at 600-700 m alt. No VH 2463 04.01.1996 International Botanical Expedition of the U.S.A. National Geographic Society (grant No 5094-93) Participants: L. Averyanov, N.T. Ban, A. Budantzev, L. Budantzev, N.T. Hiep, D.D. Huyen, P.K. Loc, G. Yakovlev

FLORA OF VIETNAM. Fam. Convallariaceae. Tupistra sp. N. Vietnam, prov. Lai Chau, distr. Sin Ho, municipality Ta Ngau about. 30 km of Lai Chau town to the N (N 22°18' E 103°15'). Secondary open. wet mossy evergreen broadleaved forest with remnants of primary. elements on rocky steep slope of remnant karst limestone ridge at. elev. about 1650 m. Terrestrial and lithophyte forb up to 0.6 m hg on. shady steep slope. Very common. No NTH 2789   25 May 1999. International Botanical Expedition of the U.S.A. National Geographic. Society (grant No 6300-98). . Collectors: N.T.Hiep, P.H.Hoang, Averyanov L.

FLORA OF LAOS Convallariaceae Tupistra muricata(Gagnep.) N.Tanaka Vientiane Prov., Vang Vieng Distr., Nathong village, about 5 km to the W of Vang Vieng town, Tham Kang Mt., around point 18°55’51.4”N 102°23’50.6”E. Dry broad-leaved primary and secondary evergreen forest on very steep rocky slopes and on vertical cliffs of remnant highly eroded rocky limestone mountains composed by solid crystalline limestone at elevation 300-400 m. Terrestrial herb with erect stem to 5 cm tall on shady slope. Leaves uniform green, to 1 m long, petiole 30-50 cm long, leaf blade elliptic to 50 cm long 15 cm wide. Rare. LA-VN 410a 14 March 2013 (living only) Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, P.V.The, S.Lorphengsy Fl. in cult. in October – November 2015. Flowers with chloroform smell, white, tepals violet at apices.

FLORA OF LAOS Convallariaceae Tupistra muricata (Gagnep.) N.Tanaka Luang Prabang prov., Ngoy distr., Nong Khiew village, Pha Deng Mountain around point 20°34'14.3''N 102°37'31.8''E. Secondary semideciduous and evergreen dry forest on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. 700-825 m a.s.l. Terrestrial herb on steep rocky shady slope. Fruits dirty violet. Not common. LA-VN 1904 31 March 2017 Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.Maisak, Vichith Lamxay, Koudkeo Phommachanh, Phizar Sayalath, Keooudone Souvannakhoummane.

FLORA OF VIETNAM Convallariaceae Aspidistra lubae Aver. et Tillich var. lancifolia Aver. et Tillich Type Hoa Binh Prov., Lac Son Distr., Tu Do Municipality, Mon village, around point 20°25’29”N 105°19’36”E. Primary broadleaved forest on steep rocky slopes of remnant mountain composed with solid crystalline highly eroded limestone at elevation about 500-700 m a.s.l. Terrestrial herb in shady place. Not rare. CPC 1566a/sp.4 25 March 2011. Coll.: L.Averyanov, T.Maisak, L.Osinovetz et al. Fl. under cultivation at January 2013. Perianth white with dirty purple-violet marking, dark dirty violet inside, stigma white to light yellowish, filaments white, anthers light yellowish. Photos. Averyanov L.

FLORA OF LAOS

FLORA OF VIETNAM Convallariaceae Aspidistra coccigera Aver. et H.-J. Tillich sp.nov. Vietnam, Quang Binh Prov., Minh Hoa Distr., Dan Hoa municipality, Vietnam-Laotian border, around Cha Lo border entrance, point 17º40’52’’N 105º46’00’’E. Broad-leaved primary forest on very steep slopes and along shale ridge at elevation 500-750 m a.s.l. Terrestrial herb on very steep rocky shady slope. Flowers white, tips of petals dark violet-black, stigma brightly red. Locally common. HAL 11817 22 April 2008 photo Coll.: L.Averyanov, P.K.Loc, N.T.Vinh, N.S.Khang

FLORA OF VIETNAM

All drawings © L. Averyanov

FLORA OF VIETNAM. Convallariaceae. Tupistra sp. Dien Bien Prov., Tua Chua Distr., Tua Thang Municipality, Ta Xi Lang village, around point 21°57'17"N 103°25'31"E. Highly degraded primary and secondary evergreen broadleaved forest on very steep slopes and on tops of remnant mountains composed with solid limestone at elevation 850-950 m a.s.l. Lithophytic herb to 20 cm tall on shady mossy rocks. Flowers green to yellow. Occasional. CPC 2284 13 April 2011. Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, N.T.Vinh.

FLORA OF VIETNAM. Fam. Convallariaceae. Tupistra sp. N. Vietnam, prov. Cao Bang, distr. Bao Lac, municipality Dinh Phung,. about 5 km to NW from Yen Lac village (N 22°44' E 105°50'), about 50. km to NWW from Cao Bang town. Old secondary closed broadleaved wet. cloud forest on steep slopes and cliffs along top of Nam Linh. limestone ridge at 1500-1580 m alt. Epilite and terrestrial herb up to. 0.6 m hg. Flowers green. Common. No CBL 468   19 Nov. 1998. International Botanical Expedition of the U.S.A. National Geographic. Society (grant No 6300-98). . Collectors: Averyanov L., P.K.Loc, N.X.Tam.

Tupistra sp.

FLORA OF VIETNAM. Convallariaceae. Campylandra theana Aver., sp.nov. Paratype. Quang Binh Prov., Tuyen Hoa Distr., Lam Hoa municipality, Hung village, around point 17°55'41"N 105°49'43"E. Highly degraded primary and secondary evergreen broadleaved forest on very steep rocky slopes of remnant limestone hills composed with highly eroded marble-like crystalline limestone at elev. 200-300 m a.s.l. Lithophytic herb on vertical shady cliff. Flowers dark dirty purple-violet, anthers dull yellow. Not common. CPC 2616 1 May 2011 Photos. Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, P.V.The, N.T.Vinh.

Текст этикетки FLORA OF LAOS Convallariaceae Tupistra muricata (Gagnep.) N.Tanaka Vientiane Prov., Kasi Distr., Tam Tai village, about 10 km to the NW of Kasi town (19°12'57"N 102°14'59"E). Remnants of primary and secondary broad-leaved evergreen forest on very steep rocky slopes of remnant mountain composed with highly eroded solid crystalline limestone at elevation 500-650 m. Terrestrial herb with erect thick stem to 0.5 m tall on shady rocky slope. Old flowers almost black. Not rare. LA-VN 756 24 March 2013 Coll.: L.Averyanov, N.S.Khang, S.Lorphengsy Flowered in October 2014. Flowers pure white. Photos.

FLORA OF LAOS Convallariaceae Tupistra muricata(Gagnep.) N.Tanaka Vientiane Prov., Vang Vieng Distr., Nathong village, about 5 km to the W of Vang Vieng town, Tham Kang Mt., around point 18°55°51.4°N 102°23°50.6°E. Dry broad-leaved primary and secondary evergreen forest on very steep rocky slopes and on vertical cliffs of remnant highly eroded rocky limestone mountains composed by solid crystalline limestone at elevation 300-400 m. Terrestrial herb with erect stem to 5 cm tall on shady slope. Leaves uniform green, to 1 m long, petiole 30-50 cm long, leaf blade elliptic to 50 cm long 15 cm wide. Rare. LA-VN 410a 14 March 2013 (living only) Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, P.V.The, S.Lorphengsy Fl. in cult. in October ° November 2015. Flowers with chloroform smell, white, tepals violet at apices.

FLORA OF LAOS. Convallariaceae. Tupistra muricata (Gagnep.) N.Tanaka. Vientiane prov., Kasi distr., Si Sang Vone village, around point 19°06'10.9"N 102°10'46.1"E. Secondary broad-leaved evergreen dry forest on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. 600-700 m a.s.l. Terrestrial and lithophytic herb on shady steep rocky slope. Locally common. Fruits globular, dirty dark green. Common. LA-VN 1786 28 March 2017. Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.Maisak, Khamla Inkhavilay, Hongthong Southyvohanh, Pinmany Sisavath, Chanthala Cheuaysamphanh.

FLORA OF VIETNAM Convallariaceae Aspidistra lubae Aver. et Tillich var. lancifolia Aver. et Tillich Type Hoa Binh Prov., Lac Son Distr., Tu Do Municipality, Mon village, around point 20°25’29”N 105°19’36”E. Primary broadleaved forest on steep rocky slopes of remnant mountain composed with solid crystalline highly eroded limestone at elevation about 500-700 m a.s.l. Terrestrial herb in shady place. Not rare. CPC 1566a/sp.4 25 March 2011. Coll.: L.Averyanov, T.Maisak, L.Osinovetz et al.

FLORA OF VIETNAM Convallariaceae Aspidistra lubae Aver. et Tillich var. lancifolia Aver. et Tillich Type Hoa Binh Prov., Lac Son Distr., Tu Do Municipality, Mon village, around point 20°25’29”N 105°19’36”E. Primary broadleaved forest on steep rocky slopes of remnant mountain composed with solid crystalline highly eroded limestone at elevation about 500-700 m a.s.l. Terrestrial herb in shady place. Not rare. CPC 1566a/sp.4 25 March 2011. Coll.: L.Averyanov, T.Maisak, L.Osinovetz et al. Fl. under cultivation at November 2013. Perianth white with dirty purple-violet marking, dark dirty violet inside, stigma white to light yellowish, filaments white, anthers light yellowish. Photos. P. Efimov.

FLORA OF LAOS Convallariaceae Aspidistra laotica A Aver. et Tillich Type Vientiane Prov., Vang Vieng Distr., Nam Pe village, about 12 km to the W of Vang Vieng town, around point 18°58’41.0”N 102°18’54.1”E. Dry broad-leaved primary and secondary evergreen and semideciduous forest on very steep rocky slopes and on vertical cliffs of remnant highly eroded rocky limestone mountains composed by solid crystalline limestone at elevation 500-1000 m. Terrestrial herb in shady rocky place. Leaves dark uniform green. Flower buds yellowish-white, purple stripped. Fruits globular, 1 cm in diam., dirty purple, rugose. Very common at 300-500 m. LA-VN 554 18 March 2013 Coll.: N.T.Hiep, L.Averyanov, K.Chantthavongsa, N.S.Khang, P.V.The, S.Lorphengsy Flowered under cultivation at June 2013. Tube white; free parts of tepals black; anthers dull yellowish, stigma dark brown, style white; ovary and pedicel light greenish. d-EXSICCATES OF VIETNAMESE FLORA 0216/LA-VN 554 & 554a (LE) © L.Averyanov

FLORA OF VIETNAM Aspidistra dodecandra (Gagnep.) Tillich (Evrardiella dodecandra Gagnep.) Khanh Hoa province, Khanh Vinh district, evergreen broad-leaved submontane forest at elevation 850 m. 28 September - 17 October 2017, P.V. The et al. Received by L. Averyanov, T. Maisak, AL 288. Herbarium sheet, living plant, photos.

FLORA OF VIETNAM Convallariaceae Aspidistra bifolia Aver., Tillich et K.S.Nguyen Type Son La Prov., Moc Chau Distr., Chieng Son comm., about 1 km to NE from Chieng Son Village, around point 20°46’02.0N 104°37’03.1E. Remnants of primary broad-leaved evergreen forest on tops of hills composed with karstic highly eroded yellow-white limestone at elevation 1150-1250 m a.s.l. Terrestrial herb with long underground rhizome in shady rocky place. Flowers white with fine purple marks. Fruits white later dirty purple-brownish wit green tint. Locally very common. 27 September 2016 CPC 8057 Photos (living plants) Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.V.Maisak, N.T.Son. d-EXSICCATES OF VIETNAMESE FLORA 0263/CPC 8057 © @ L.Averyanov

FLORA OF VIETNAM

FLORA OF VIETNAM. Fam. Convallariaceae. Tupistra sp. N.Vietnam, Cao Bang Prov., Trung Khanh Distr., Ngoc Khe Municipality, Pac Nga village, around point 22°54'59"N, 106°31'44"E, at elevation 700-800 m a.s.l. Closed secondary and remnants of primary broad-leaved evergreen forest on very steep rocky slopes of rocky ridges and mesae composed with light gray solid marble-like highly eroded crystalline limestone. Lithophytic herb on mossy exposed rocks. Not common. No HAL 5696 12 June 2004. . Collectors: L.Averyanov, P.K.Loc, P.V.The, N.T.Vinh.

FLORA OF VIETNAM

FLORA OF VIETNAM Convallariaceae Tupistra theana Aver. Paratype Quang Binh Prov., Minh Hoa Distr., Thuong Hoa municipality, around point 17°41’28”N 105°53’42.7”E. Primary closed evergreen broad-leaved forest on rocky very steep cliffy slope of remnant mountain composed with solid crystalline limestone at elevation about 700 m a.s.l. near to the top of mountain. Lithophyte herb on vertical shady cliffs. Fruits dark green, 1 or 2-seeded. Locally common. CPC 3952 5 August 2011 Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Vinh, N.V.Tap, P.V.The, L.T.Kien.

FLORA OF VIETNAM. Convallariaceae. Tupistra wattii. Dien Bien Prov., Muong Cha Distr., Hua Ngai Municipality, Ha La Chu Village, around point 21°53'46"N 103°10'17"E. Highly degraded primary evergreen broadleaved humid forest on very steep slopes of remnant mountains composed with solid limestone at elevation 1100-1400 m a.s.l. Terrestrial and lithophytic herb to 0.5 m tall on shady rocky slope. Flowers dull orange. Common. CPC 1937 07 April 2011. Coll.: L.Averyanov, P.K.Loc, N.Q.Hieu, N.T.Vinh.

FLORA OF LAOS Convallariaceae Tupistra muricata (Gagnep.) N.Tanaka Vientiane prov., Kasi distr., Si Sang Vone village, around point 19°06'10.9''N 102°10'46.1''E. Secondary and primary evergreen dry forest on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. 600-750 m a.s.l. Terrestrial herb on shady rocky slope. Fruits globular, dirty green to dirty violet. Locally common. LA-VN 1822 28 March 2017 Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.Maisak, Khamla Inkhavilay, Hongthong Southyvohanh, Pinmany Sisavath, Chanthala Cheuaysamphanh.

FLORA OF LAOS. Convallariaceae. Tupistra muricata (Gagnep.) N.Tanaka. Vientiane prov., Vang Vieng distr., Oa Hom village, around point 19°06'52.4"N 102°22'35.8"E. Secondary broad-leaved evergreen and semideciduous dry forest with on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. 550-650 m a.s.l. Terrestrial herb on shady rocky steep slope. Fruits globular, dirty green with violet tint. Locally common. LA-VN 1517 22 March 2017. Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, C.Q.Ngan, T.Maisak, Khamla Inkhavilay, Hongthong Southyvohanh, Pinmany Sisavath, Chanthala Cheuaysamphanh.

FLORA OF VIETNAM Convallariaceae Aspidistra lubae Aver. et Tillich var. lancifolia Aver. et Tillich Type Hoa Binh Prov., Lac Son Distr., Tu Do Municipality, Mon village, around point 20°25’29”N 105°19’36”E. Primary broadleaved forest on steep rocky slopes of remnant mountain composed with solid crystalline highly eroded limestone at elevation about 500-700 m a.s.l. Terrestrial herb in shady place. Not rare. CPC 1566a/sp.4 25 March 2011. Coll.: L.Averyanov, T.Maisak, L.Osinovetz et al.

FLORA OF VIETNAM Aspidistra heterocarpa Aver., Tillich & V.T. Pham var. echinata T.A.Le, Aver. & Tillich Thua Thien - Hue province, Nam Dong district, Thuong Quang commune, around point 16°07’28.5”N 107°35’52.9”E, primary and secondary evergreen broad-leaved lowland forest at elevation 500-550 m a.s.l., terrestrial herb on shady reclined slope, not common, 10 November 2017, Le Tuan Anh, Nguyen Thi Lien Thuong, LTA101. d-EXSICCATES OF VIETNAMESE FLORA 0295/LTA101 © Le Tuan Anh & L.Averyanov.

FLORA OF VIETNAM Convallriaceae Aspidistra elatior Blume var. vietnamensis Aver. et H.-J. Tillich var. nov. Type Quang Nam Prov., Dong Giang Distr., Ma Cooih Municipality around point 15°58’11”N 107°40’15”E. Remnants of primary closed evergreen broad-leaved forests on very steep slopes of hill composed with solid highly eroded limestone at elevation 450-500 m a.s.l. Terrestrial herb on steep shady slope. Leaves with numerous small yellow spots. Locally very common. HAL 12116a 9 January 2009 Coll.: L.Averyanov, P.K.Loc, P.V.The, N.T.Vinh Flowered under cultivation at 10 July 2011. Flowers odorless, dark dirty purple, tepal apices dull whitish. Photos. L.Averyanov, HAL 12116. Type.

FLORA OF VIETNAM Aspidistra dodecandra (Gagnep.) Tillich (Evrardiella dodecandra Gagnep.) Khanh Hoa province, Khanh Vinh district, evergreen broad-leaved submontane forest at elevation 850 m. 28 September - 17 October 2017, P.V. The et al. Received by L. Averyanov, T. Maisak, AL 288. Herbarium sheet, living plant, photos.