Herbarium LE - Specimens

FLORA OF VIETNAM

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM Ericaceae Rhododendron saxicola Sleumer Ha Giang prov., Quan Ba distr., Tung Vai commune, Thang village, around point 23º03ʹ04.2ʺN 104º51ʹ09.7ʺE, steep rocky slopes near mountain top composed with eroded stratified highly eroded limestone at elevation 1200-1450 m a.s.l. Primary evergreen broad-leaved very humid forest. Tree 3-4 m tall on rocky mountain top. Leaves purple below. Common. 24 April 2018, L. Averyanov, Nguyen Sinh Khang, Nguyen Tien Hiep, Nguyen Quang Hieu, Chuong Quang Ngan, T. Maisak, VR 834.

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM Rubiaceae Mussaenda glabra Vahl Ha Giang prov., Quan Ba distr., Tung Vai commune, Thang village, around point 23º03ʹ04.2ʺN 104º51ʹ09.7ʺE, steep rocky slopes near mountain top composed with eroded stratified highly eroded limestone at elevation 1200-1450 m a.s.l. Primary evergreen broad-leaved very humid forest. Scandent shrub 3-4 m tall in open place. Inflorescence bracts white, flowers yellow. Common. 24 April 2018, L. Averyanov, Nguyen Sinh Khang, Nguyen Tien Hiep, Nguyen Quang Hieu, Chuong Quang Ngan, T. Maisak, VR 852.

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM Polypodiaceae s.l. Ha Giang prov., Quan Ba distr., Tung Vai commune, Thang village, around point 23º03ʹ04.2ʺN 104º51ʹ09.7ʺE, steep rocky slopes near mountain top composed with eroded stratified highly eroded limestone at elevation 1200-1450 m a.s.l. Primary evergreen broad-leaved very humid forest. Creeping epiphyte on mossy tree. Common. 24 April 2018, L. Averyanov, Nguyen Sinh Khang, Nguyen Tien Hiep, Nguyen Quang Hieu, Chuong Quang Ngan, T. Maisak, VR 827.

All photos © L. Averyanov & K.S. Nguyen

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM Taxaceae Amentotaxus argotaenia (Hance) Pilg. Ha Giang prov., Quan Ba distr., Tung Vai commune, Thang village, around point 23º03ʹ04.2ʺN 104º51ʹ09.7ʺE, steep rocky slopes near mountain top composed with eroded stratified highly eroded limestone at elevation 1200-1450 m a.s.l. Primary evergreen broad-leaved very humid forest. Trre to 15 m tall. Fructification brightly red, juicy. Occasional. 24 April 2018, L. Averyanov, Nguyen Sinh Khang, Nguyen Tien Hiep, Nguyen Quang Hieu, Chuong Quang Ngan, T. Maisak, VR 812a.

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM Polygalaceae Polygala sp. Ha Giang prov., Quan Ba distr., Tung Vai commune, Thang village, around point 23º03ʹ04.2ʺN 104º51ʹ09.7ʺE, steep rocky slopes near mountain top composed with eroded stratified highly eroded limestone at elevation 1200-1450 m a.s.l. Primary evergreen broad-leaved very humid forest. Undershrub to 1.5 m tall in open place. Not common. 24 April 2018, L. Averyanov, Nguyen Sinh Khang, Nguyen Tien Hiep, Nguyen Quang Hieu, Chuong Quang Ngan, T. Maisak, VR 805.

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM Begoniaceae Begonia sp.nov. Ha Giang Prov., Bac Me District., Phieng Luong commune, Phieng Luong village, So Long mountaine. Primary evergreen broad- leaf forest on limestone at elevation 1000-1100 m a.s.l. Grows in soil limestone humus , on slope and top of mountaine at elevation 1135 m a.s.l. around point 220 38’ 27.1’’N;1050 18’ 48.1’’E, 1097 m a.s.l. Terrestrial herb, succulent, fruit pink 06 April 2014, CPC 7291 Coll.: N. Q. Hieu, N.T.Hiep, N.N. Hung, T.Q. Ngan, T.P.Tuan, G.M. Pao

FLORA OF VIETNAM

FLORA OF VIETNAM

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM Begoniaceae Begonia “arachna” sp. nov.? Cao Bang Prov., Bao Lac Distr., Hong An municipality, Mi Lung village. Primary broad-leaved and mixed humid evergreen forest (with Podocarpus, Pinus, Fokienia and Tsuga) on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation about 1500-1550 m a.s.l. around point 22°49’15.4’’N 105°49’53’’E. Lithophytic herb on vertical mossy cliffs and boulders. Leaves emerald green, below with purple nerves. Flowers light pink, anther and style yellow, ovary purple. Very common. 21 November 2014 CPC 7568 photos Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz

FLORA OF VIETNAM Begoniaceae Begonia sp. Cao Bang Prov., Nguyen Binh Distr., Yen Lac Municipality, Chi Doi village, around point 22°45'34"N, 105°51'48.7"E. Primary fractionaly logged coniferous forest with Pseudotsuga sinensis along highly eroded rocky limestone ridge at elevation about 1400 m a.s.l. Lithophytic herb on shady cliffs on miffle part of mountain slope. Leaves dark green above, light green with purple veins below. Flowers pink, anthers yellow. Very common. 5 October 2013 CPC 5421 L.Averyanov, N.T.Hiep, L.M.Tuan, N.S.Khang, T.Maisak, L.Osinovets Photos

FLORA OF VIETNAM Begoniaceae Begonia khangii sp.nov.? Nghe An Prov., Ky Son Distr., Na Ngoi Municipality, eastern slopes of Phu Xai Lai Leng Mountain. Primary very humid broad-leaved forest on steep mountain slopes composed of sandstone and gray shale at elevation 2100-2200 m a.s.l. around point 19°11’59.3’’N 104°11’23.9’’E. Lithophytic herb on mossy rocks. Flowers pink, anthers and stigmas yellow. Leaves below with dull dirty purple spots. Very common. 25 October 2013 CPC 6281 Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, L.M.Tuan, N.A.Trang, L.H.Dan

FLORA OF LAOS Begoniaceae Begonia sp. Laos, Khammouane prov., Thakhek. Отвесная стена известнякового останца на высоте 1-2 м, пещера Thamnangene. 157 м над ур. м., 17°26.634N, 105°56.872E. Е.Константинов K245. Гербарный сбор/фото 03.11.2013.

FLORA OF LAOS Hin Nam No Nature Protected Area Convallariaceae Peliosanthes cryptogamica sp. nov. ? (P. inaperta Aver. & N. Tanaka aff.) Laos, Khammoune prov., Boualapha distr., Nong Seng village, Vang Phae Lay mountain, around point 17°31'02.7"N, 105°47'17.2"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 300-530 m a.s.l. Terrestrial and lithophytic herb on steep shady rocky slope. Very common to abundant. Usual co-dominant of forest herbaceous stratum. Flowers dark green, cleistogamous (?). 11 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 909.

FLORA OF VIETNAM

FLORA OF VIETNAM Plants of Phong Nha - Ke Bang National Park Fam. Pentaphragmaceae Pentaphragma sp. Vietnam, Quang Binh Prov., Minh Hoa Distr., Thuong Hoa Municipality, about 1 km to N of Ban On village (17º40'21''N, 105º58'00E). Logged primary closed evergreen broad-leaved lowland forest along stream and foothills of rocky ridge composed with solid marble-like highly eroded crystalline limestone at elevation 340-400 m a.s.l. Terrestrial herb on steep slope. Flowers dull pinkish-violet. Rare. No HAL 5929 19 January 2005. Collectors: L.Averyanov, P.K.Loc, P.V.The, A.Averyanova, N.T.Vinh, N.Q.Vinh, N.T.Binh.

FLORA OF VIETNAM

FLORA OF VIETNAM Rubiaceae Ophiorrhiza sp. Ha Giang prov., Quan Ba distr., Tung Vai commune, Thang village, around point 23º03ʹ04.2ʺN 104º51ʹ09.7ʺE, steep rocky slopes near mountain top composed with eroded stratified highly eroded limestone at elevation 1200-1450 m a.s.l. Primary evergreen broad-leaved very humid forest. Epiphyte on mossy tree. Common. 24 April 2018, L. Averyanov, Nguyen Sinh Khang, Nguyen Tien Hiep, Nguyen Quang Hieu, Chuong Quang Ngan, T. Maisak, VR 815.

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM Convallariaceae Tupistra fungilliformis F.T. Wang & S. Yun Liang Ha Giang prov., Quan Ba distr., Tung Vai commune, Thang village, around point 23º03ʹ04.2ʺN 104º51ʹ09.7ʺE, steep rocky slopes near mountain top composed with eroded stratified highly eroded limestone at elevation 1200-1450 m a.s.l. Primary evergreen broad-leaved very humid forest. Epiphyte on mossy tree. Common. 24 April 2018, L. Averyanov, Nguyen Sinh Khang, Nguyen Tien Hiep, Nguyen Quang Hieu, Chuong Quang Ngan, T. Maisak, VR 811.

All photos © L. Averyanov & K.S. Nguyen

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM Begoniaceae Begonia sp. Cao Bang Prov., Thong Nong Distr., Yen Son Municipality, Nhieu Lung village, around point 22°46'07.9"N, 105°53'59.4"E. Primary fractionaly logged coniferous forest with Pseudotsuga sinensis along highly eroded rocky limestone ridge at elevation 1150-1250 m a.s.l. Lithophytic rosulate herb on vertical cliffs in shady humid place in rocky valley. Leaves dark green with light green palmate markings above, light green with purple nerves below. Flowers pink, anther yellow. Locally very common. 8 October 2013 CPC 5472 L.Averyanov, N.T.Hiep, L.M.Tuan, N.S.Khang, T.Maisak, L.Osinovets

FLORA OF VIETNAM

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM Rubiaceae Ophiorrhiza sp. Ha Giang prov., Quan Ba distr., Tung Vai commune, Thang village, around point 23º03ʹ04.2ʺN 104º51ʹ09.7ʺE, steep rocky slopes near mountain top composed with eroded stratified highly eroded limestone at elevation 1200-1450 m a.s.l. Primary evergreen broad-leaved very humid forest. Herb to 0.5 m tall in open place. Common. 24 April 2018, L. Averyanov, Nguyen Sinh Khang, Nguyen Tien Hiep, Nguyen Quang Hieu, Chuong Quang Ngan, T. Maisak, VR 806.

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM Begoniaceae Begonia sp. Tuyen Quang Prov., Na Hang Distr., Sinh Long Municipality, Khuoi Phin village, around point 22°38'21.9"N, 105°20'43.5"E. Primary evergreen mixed humid forest on rocky limestone on highly eroded karstic hills composed with gray solid marble-like limestone at elevation 1100-1200 m a.s.l. Lithophytic herb on vertical cliffs. Leaves above uniform green, below brightly red. Flowers pink. Abundant. 29 September 2013 CPC 5317 L.Averyanov, N.T.Hiep, L.M.Tuan, N.S.Khang, D.V.Cuong, N.V.Truong.T.Maisak, L.Osinovets

Northern Vietnam, with no exact location. 2014, Huong, XL 518

FLORA OF VIETNAM Begoniaceae Begonia khangii sp.nov.? Nghe An Prov., Ky Son Distr., Na Ngoi Municipality, eastern slopes of Phu Xai Lai Leng Mountain. Primary very humid broad-leaved forest on steep mountain slopes composed of sandstone and gray shale at elevation 2100-2200 m a.s.l. around point 19°11’59.3’’N 104°11’23.9’’E. Lithophytic herb on mossy rocks. Flowers pink, anthers and stigmas yellow. Leaves below with dull dirty purple spots. Very common. 25 October 2013 CPC 6281 Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, L.M.Tuan, N.A.Trang, L.H.Dan

FLORA OF VIETNAM

FLORA OF LAOS Begoniaceae Begonia sp. Vientiane Prov., Vang Vieng Distr., Na Po village, about 12 km to the W of Vang Vieng town, Pa Nang Oua Mt., around point 18°55’44.9”N 102°20’20.1”E. Dry broad-leaved primary and secondary evergreen and semideciduous forest on very steep rocky slopes and on vertical cliffs of remnant highly eroded rocky limestone mountains composed by solid crystalline limestone at elevation 700-1000 m. Creeping lithophyte on shady cliff near mountain top. Leaves dark green to olive-brown with white spots above, dull violet below with unclear white spots, petiole purple. Locally common. LA-VN 521 17 March 2013 Coll.: N.T.Hiep, L.Averyanov, K.Chantthavongsa, N.S.Khang, P.V.The, S.Lorphengsy

FLORA OF VIETNAM Peliosanthes convallarioides Aver. & N.Tanaka Quang Binh Prov., Minh Hoa Distr., Dan Hoa Municipality, Bai Dinh village around point 17°45’28”N 105°46’14”E, very steep rocky slopes of highly eroded marble-like solid remnant limestone mountains, 600-700 m, terrestrial rosulate herb on shady very steep rocky slope, leaves dark glossy green with whitish longitudinal wide stripe, flowers greenish, often with dirty violet tint, fruits blue, not rare, 4 February 2009, L. Averyanov et al., HAL 12209. Herbarium prepared at 19.04.2019, L. Averyanov, HAL 12209a. d-EXSICCATES OF VIETNAMESE FLORA 0354 / HAL 12209a © L. Averyanov.

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM Rubiaceae Ophiorrhiza sp. Ha Giang prov., Quan Ba distr., Tung Vai commune, Thang village, around point 23º03ʹ04.2ʺN 104º51ʹ09.7ʺE, steep rocky slopes near mountain top composed with eroded stratified highly eroded limestone at elevation 1200-1450 m a.s.l. Primary evergreen broad-leaved very humid forest. Terrestrial and lithophytic herb 1 m tall. Very common on rocky mountain top. 24 April 2018, L. Averyanov, Nguyen Sinh Khang, Nguyen Tien Hiep, Nguyen Quang Hieu, Chuong Quang Ngan, T. Maisak, VR 809.

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM Begoniaceae Begonia “arachna” sp.nov.? Ha Giang Prov., Bac Me Distr., Phieng Luong municipality, around Phieng Luong village. Primary broad-leaved humid evergreen forest on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation 1100-1150 m a.s.l. around point 22°38’07.1’’N 105°19’22.1’’E. Lithophytic rosulate herb on shady vertical cliffs. Flowers light pink to almost white, ant hers and stigma yellow, Fruits purple. Leaves green, dark red-purple below, petioles dull purple. Very common. 13 November 2014 CPC 7391 Photos. Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz

FLORA OF VIETNAM Begoniaceae Begonia sp. Cao Bang Prov., Thong Nong Distr., Yen Son Municipality, Nhieu Lung village, around point 22°46'07.9"N, 105°53'59.4"E. Primary fractionaly logged coniferous forest with Pseudotsuga sinensis along highly eroded rocky limestone ridge at elevation 1150-1250 m a.s.l. Lithophytic rosulate herb on vertical cliffs in shady humid place in rocky valley. Leaves dark green with light green palmate markings above, light green with purple nerves below. Flowers pink, anther yellow. Locally very common. 8 October 2013 CPC 5472 L.Averyanov, N.T.Hiep, L.M.Tuan, N.S.Khang, T.Maisak, L.Osinovets

FLORA OF VIETNAM Podocarpaceae Podocarpus pilgeri Foxw. Ha Giang prov., Quan Ba distr., Tung Vai commune, Thang village, around point 23º03ʹ04.2ʺN 104º51ʹ09.7ʺE, steep rocky slopes near mountain top composed with eroded stratified highly eroded limestone at elevation 1200-1450 m a.s.l. Primary evergreen broad-leaved very humid forest. Tree to 10 m tall near rocky mountain top. Common. 24 April 2018, L. Averyanov, Nguyen Sinh Khang, Nguyen Tien Hiep, Nguyen Quang Hieu, Chuong Quang Ngan, T. Maisak, VR 800.

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM Lamiaceae Scutellaria sichourensis C.Y. Wu & H.W. Li Ha Giang prov., Quan Ba distr., Tung Vai commune, Thang village, around point 23º03ʹ04.2ʺN 104º51ʹ09.7ʺE, steep rocky slopes near mountain top composed with eroded stratified highly eroded limestone at elevation 1200-1450 m a.s.l. Primary evergreen broad-leaved very humid forest. Terrestrial herb to 1 m tall among open mossy rocks. Flowers purple-violet. Not rare. 24 April 2018, L. Averyanov, Nguyen Sinh Khang, Nguyen Tien Hiep, Nguyen Quang Hieu, Chuong Quang Ngan, T. Maisak, VR 801.

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM Begoniaceae Begonia sp. Cao Bang Prov., Nguyen Binh Distr., Yen Lac Municipality, Chi Doi village, around point 22°45'34"N, 105°51'48.7"E. Primary fractionaly logged coniferous forest with Pseudotsuga sinensis along highly eroded rocky limestone ridge at elevation about 1400 m a.s.l. Lithophytic herb on shady cliffs on miffle part of mountain slope. Leaves dark green above, light green with purple veins below. Flowers pink, anthers yellow. Very common. 5 October 2013 CPC 5421 L.Averyanov, N.T.Hiep, L.M.Tuan, N.S.Khang, T.Maisak, L.Osinovets Photos

Northern Vietnam, with no exact location. 2014, Huong, XL 518

FLORA OF VIETNAM Begoniaceae Begonia sp. Nghe An Prov., Ky Son Distr., Na Ngoi Municipality, Phu Kha village, eastern slopes of Phu Xai Lai Leng mountain system. Primary broad-leaved humid forest on steep granite slopes at elevation about 2350 m a.s.l. around point 19°12’21.2’’N 104°11’28.9’’E. Tuberiferous herb 5-10 cm tall. Rare. 23 October 2013 CPC 6072 Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, L.M.Tuan, N.A.Trang, L.H.Dang