Herbarium LE - Specimens

Fam.?

Gia Lai Province, K'Bang District. Secondary and remnants of primary evergreen broad-leaved dry forest with Dacrydium on lava rocks (red soils) at elevation about 1000 m a.s.l., around point 14º15N 108º35E. Tree to 8 m tall along forest trail. Occasional.

6 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4793.

Convallariaceae

Peliosanthes graminea Ridl.?

Dac Lak Province, Lac District, Evergreen broad-leaved forest on very steep slopes along stream. Not common. Flowers dark violet.

Collected in Nguyen Van Canh Garden on 26 March 2026, Nguyen Van Canh, L. Averyanov, Truong Ba Vuong, T. Maisak, AL4551.

Convallariaceae

Peliosanthes weberi (L.Rodr.) N.Tanaka

Cultivated plant originated from Dac Nong Province, Dak Mil District, evergreen forest mixed with dipterocarp forest at elevation of about 300 m a.s.l., common,

Collected in Nguyen Van Canh Garden on 26 March 2026, Nguyen Van Canh, L. Averyanov, Truong Ba Vuong, T. Maisak, AL4547. Flowers light green.

Convallariaceae

Peliosanthes sp.

Dien Bien Prov., Muong Nhe Distr., Muong Nhe Natural Reserve, Leng Su Sin Municipality, A Suoi Voi Village around point about N22º20’16’’ E102º21’35”. Secondary evergreen broad-leaved lowland and submontane forest on mountain stream slopes composed with sandstone at elevations 500–600 m a.s.l. Terrestrial herb on shady steep stream slope. Flower buds dark green with violet tint. Fruits glossy green. Not rare.

18 May 2022, L. Averyanov, Tran Huy Thai, Khang Sinh Nguyen, T. Maisak, VR 1724.

Acanthaceae

“Strobilanthes” sp.

Gia Lai Province, Mang Yang District, Ayune Commune, Kon Ka Kinh National Park. Secondary and primary dry evergreen broad-leaved forest on shale and lava rocks at elevation of 900-1000 m a.s.l. around point 14º13N 108º20E. Erect herb or undershrub to 1.5 m tall on open places along trail. Flowers bright red. Occasional.

2 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4685.

Convallariaceae

Aspidistra sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 800-900 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial rhizomatous herb on shady stream slope. Leaves evenly dark green. Locally very common.

30 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4667.

Living specimens have died before flowering.

Asclepiadaceae

Dischidia sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 800-900 m a.s.l. around the point 12º10’N 107º46’E. Epiphytic herbaceous vine on mossy trees along rocky stream. Not rare.

30 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4665.

Convallariaceae

Tupistra khangii Aver., N.Tanaka & Vislobokov

Plant obtained from local collectors in Son La Province, Thuan Chau District, evergreen broad-leaved forest at elevation of about 900 m a.s.l. Common.

November 2025, Nguyen Van Canh, AL4573.

Cycadaceae

Cycas misholitzii

Dac Lak Province, Lak District, Lieng Buk Sak Waterfall, evergreen broad-leaved forest on very steep slopes along stream. Terrestrial palm-like plant. Sterile samples very common.

August 2025, Nguyen Van Canh, AL4566.

Convallariaceae

Aspidistra hainanensis?

Plant obtained from local collectors in Lao Cai Province, Bat Xat District, evergreen forest at elevation of about 1000 m a.s.l.

20.10.2025, Nguyen Van Canh, AL4567.

Convallariaceae

Aspidistra cylindrica?

Dak Lak Province, Chu Mu Mt., evergreen broad-leaved forest at elevation of about 600 m a.s.l. Occasional.

August 2025, Nguyen Van Canh, AL4569.

Acanthaceae

Phlogacanthus sp.

Dak Lac Province, Yok Don National Park, Dipterocarp forest at elevation of about 200 m a.s.l., flowers dirty purple. Plant cultivated in Canh Garden in Buon Ma Thuot Town. Collected by on 26 March 2026, Nguyen Van Canh, L. Averyanov, Truong Ba Vuong, T. Maisak, AL4554.

Araceae

Thyphonium sp.

Dak Lak Province, Lak District, lithophytic herb on open rocks along stream at elevation of about 600 m a.s.l. Not common.

12 September 2025, Nguyen Van Canh, AL4591.

Myrsinaceae

Ardisia sp.

Dak Lak Province, Lal District, secondary open forest at elevation of about 500 m a.s.l. Shrub to 3 m tall, flowers pink. Common.

February 2025, Nguyen Van Canh, AL4588.

Myrsinaceae

Ardisia sp.

Dak Lak Province, Lal District, secondary open forest at elevation of about 500 m a.s.l. Shrub to 3 m tall, flowers pink. Common.

February 2025, Nguyen Van Canh, AL4588.

Zingiberaceae

Hedychium sp.

Plant obtained from local collectors in Lai Chau Province, evergreen forest at elevation of about 1000 m a.s.l. Terrestrial herb to 1 m tall. Flowers white with yellow. Common.

12 September 2025, Nguyen Van Canh, AL4589.

Convallariaceae

Aspidistra cylindrica?

Dak Lak Province, Chu Mu Mountain, evergreen broad-leaved forest at elevation of about 600 m a.s.l. Terrestrial rhizomatous herb. Flowers dull yellow, or yellow with purple. Common.

August 2025, Nguyen Van Canh, AL4590.

Cycadaceae

Cycas misholitzii

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 800-900 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial plant with thick underground caudex and suberect leaves to 2 m long on shady slope near stream. Occasional.

30 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4645.

Oleaceae

Jasminum sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 800-900 m a.s.l. around the point 12º10’N 107º46’E. Scandent shrub with vine stem to 3 m long among open shrubs on exposed stream rocks. Flowers white. Common.

30 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4644.

Fabaceae

Entada sp.?

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 800-900 m a.s.l. around the point 12º10’N 107º46’E. Woody vine to 10 m long among open shrubs on exposed stream rocks. Flowers yellowish. Not common.

30 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4643.

Fabaceae

Entada sp.?

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 800-900 m a.s.l. around the point 12º10’N 107º46’E. Woody vine to 10 m long among open shrubs on exposed stream rocks. Flowers yellowish. Not common.

30 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4643.

Selaginellaceae

Selaginella sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Wetland thicket in marshland depression with domination of Pandanus and Livistona along stream. Terrestrial sedge to 1 m tall on shady slope. Common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4595.

Convallariaceae

Aspidistra cylindrica?

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial rhizomatous herb with thick hypogeous rhizome on shady stream slopy. Occasional.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4596.

Selaginellaceae

Selaginella sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial herb to 0.5 m tall on shady stream slope. Common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4598.

Araceae

Pothos chinensis (Raf.) Merr.

Gia Lai Province, Mang Yang District, Ayune Commune, Kon Ka Kinh National Park. Secondary and primary dry evergreen broad-leaved forest on shale and lava rocks at elevation of 900-1000 m a.s.l. around point 14º13N 108º20E. Semiwoody creeping epiphytic vine to 15 m long on tall tree. Locally common.

2 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4692.

Rubiaceae

Mussaenda sp.

Gia Lai Province, K'Bang District. Highly degraded secondary evergreen broad-leaved dry forest at elevation 600-800 m a.s.l., around point 14º26N 108º22E. Scandent shrub with shoots to 3 m long. Inflorescence bracts white, flowers ywllow to yellow-orange. Very common.

7 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4759.

Convallariaceae

Peliosanthes griffithii Baker?

Cultivated plant originated from Dak Nong Province, Nam Nung Nature Reserve, evergreen broad-leaved forest at elevation of about 800 m a.s.l.

Selaginellaceae

Selaginella sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial herb on shady stream slope. Not common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4593.

Ericaceae (Monotropaceae)

Monotropa callistoma Ze Wei, Li J.Liu & Bing Liu

Plant obtained from local collectors in Lao Cai Province, Bat Xat District, evergreen broad-leaved forest at elevation of about 1800 m a.s.l. Not common. Flowers white with yellow mouth.

26 March 2026, Nguyen Van Canh, AL4546.

Pandanaceae

Pandanus sp.

Gia Lai Province, Mang Yang District, Ayune Commune, Kon Ka Kinh National Park. Secondary and primary dry evergreen broad-leaved forest on shale and lava rocks at elevation of 900-1000 m a.s.l. around point 14º13N 108º20E. Palm-like few branching treelet to 4 m tall in shady wet place. Common.

3 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4713.

Gesneriaceae

Petrocosmea sp.

Lao Cai Prov., Sapa Distr., the territory of Hoang Lien National Park allied to Sapa Town, route to Ham Rong mountain, around point N 22º20’02’’ E 103º50’55”, elevation 1600-1680 m a.s.l. Humid mossy secondary forest and scrub on very steep slopes of mountain composed with solid marble-like highly eroded limestone on the territory of town botanical garden. Lithophytic rosulate herb in crevices of vertical shady mossy limestone cliff. Not common.

26 May 2022, L. Averyanov, Tran Huy Thai, Khang Sinh Nguyen, T. Maisak, VR 1821.

Orchidaceae

Oberonia sp.

Lam Dong Province, Bidoup Nui Ba National Park, humid primary evergreen broad-leaved and secondary mixed (with Pinus kesiya) forest at elevation of 1200-2100 m a.s.l.

14-26 January 2026, A. Petrov, AL4534 (living plant only).

Flowered in culture on 30 July 2026, L. Averyanov, AL4534.

Vitaceae

Tetrastigma sp.?

Lai Chau Province, Muong Te District, Mu Ca Commune. Old rich secondary evergreen broad-leaved forest on stratified shale at elevation of about 540 m a.s.l. around point 22º29’40’’N 102º24’15’’E. Small treelet or woody vine to 1 m long. Not common.

25 April 2026, Nguyen Tuan Duong, Nguyen Chi Thanh, Nguyen Tuan Cuong, Nguyen Tien Hiep, Nguyen Huu Cuong, L. Averyanov, T. Maisak, АL5019.

Convallariaceae

Aspidistra sp. (“macrocarpa”)

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 1000-1200 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial rhizomatous herb in shady place. Not rare.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4640.

Convallariaceae

Aspidistra sp. (“macrocarpa”)

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 1000-1200 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial rhizomatous herb in shady place. Not rare.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4640.

Polypodiaceae s.l.

Vittaria sp.

Xiangkhouang Prov., Peak Distr., Oran village, around point 19°38’53”N 103°23’12”E. Primary and secondary broad-leaved evergreen dry forest on shale sandstone slopes and along ridge edge at elev. about 1750-1850 m a.s.l. Epiphyte on mossy tree in shady, humid place. Not common.

LA-VN 901 2 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

Polypodiaceae s.l.

Tectaria sp.

Cao Bang Prov., Bao Lac Distr., Hong An municipality, Mi Lung village. Primary broad-leaved and mixed humid evergreen forest on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation 1400-1450 m a.s.l. around point 22°49’23.4’’N 105°49’54.5’’E. Terrestrial fern in shady humid places among boulders. Fronds to 1.5 m long. Common.

19 November 2014 CPC 7501

Coll.: L.Averyanov, N.T.Hiep, N.S.Khang, T.Maisak, L.Osinovetz

Polypodiaceae s.l.

Tectaria sp.?

Houphan Prov., Hem Distr., Khon Ngua village, around point 19°55'53.2''N 103°16'24.2''E.Secondary broad-leaved evergreen open dry forest with bamboo on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. 700-950 m a.s.l.

Lithophytic fern on shady cliff near mountain top. Common.

LA-VN 1490 15 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham, Somchanh Bounphanmy.

Polypodiaceae s.l.

Thelypteris sp.

Houphan Prov., Hem Distr., Nacoc village, Pu He mountain around point 20°10'36.7''N 103°11'39.2''E. Secondary broad-leaved evergreen open dry forest with bamboo on very steep rocky mountain slopes composed by shaly limestone at elev. 700-900 m a.s.l. Terrestrial fern in open wet place along rocky mountain stream. Common.

LA-VN 1370 12 April 2015

Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.

Polypodiaceae

Pteris sp.

Ha Giang Prov., Vi Xuyen Distr., Tung Ba municipality, Khau Ca NR. Primary broad-leaved humid evergreen forest on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation 700-1150 m a.s.l. around point 22°50’39.4’’N 105°07’50.3’’E. Terrestrial herb 0.5 m in tall, common.

08 May 2015 CPC 7773

Coll.: N.Q. Hieu, N.T.Hiep, T.B. Ngan, C.Q. Ngan, D.K. Harder, C. P. Tuy, G.C.Pao

VIETNAM

Polipodiaceae

Thelypteris sp.

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 1070, 20.10.1990

VIETNAM

Polipodiaceae

Thelypteris sp.

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 1070, 20.10.1990

Polypodiaceae s.l.

Tectaria sp.

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 816, 15.10.1990

Polypodiaceae s.l.

Tectaria sp.

Quang Ngai Prov.

Thai Tran Bai

412, 07.10.1990

VIETNAM

Polipodiaceae

Thelypteris sp.

Quang Ninh Prov.

Kudryavtzeva E., Ogureeva G.

Kudryavtzeva 551, 09.10.1990

Polypodiaceae s.l.

Thelipteris sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12º30'N, 108º30'E) to main peak of

Chu Yang Sinh mt system (12º24'N, 108º26'E) along Dak Tour river (N slope). Primary mossy cloud mixed forest (with Pinus, Fokienia,

Dacrydium and Dacrycarpus) on very steep granite slopes at elev. 2200-2400 m. Terrestrial fern on steep mossy cliff. Common.

No VH 6207 09.05.2000

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 6383-98)

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

Polypodiaceae

Vittaria sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12º30'N, 108º30'E) along Dak Tour river on N macroslope of Chu Yang Sinh mt system to main peak (12º24'N, 108º26'E). Primary broad-leaved mountain evergreen closed forest on granites along river slopes at elev. 1000-1200 m. Epiphyte with pendent fronds up to 1.2 m lg in shady river canyon. Not rare.

No VH 6025 05-06.05.2000

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 6383-98)

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

Polypodiaceae s.l.

Vittaria sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12º30'N, 108º30'E) to main peak of

Chu Yang Sinh mt system (12º24'N, 108º26'E) along Dak Tour river (N slope). Primary wet closed mountain broad-leaved and mixed forest on

granites along very steep shady slopes of river canyon at elev. 1700 m. Epiphyte on vertical mossy shady cliffs along river. Not common.

No VH 6349 11.05.2000

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 6383-98)

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh

Polypodiaceae s.l.

Vittaria sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12º30'N, 108º30'E) to main peak of

Chu Yang Sinh mt system (12º24'N, 108º26'E) along Dak Tour river (N slope). Primary wet closed mixed forest (with Pinus, Fokienia,

Dacrydium and Dacrycarpus) along ridge at elev. 1600 m. Epiphyte in shady place. Not rare.

No VH 6185 09.05.2000

International Botanical Expedition of the U.S.A. National Geographic Society

Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh