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LE 01318702

LE 01318702

Leptochilus sinovietnamica Liang Zhang, N.T.Lu & Li Bing Zhang⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: Tien Hiep Nguyen; Averyanov, Leonid Vladimirovich Field collecting number: LA-VN1487. Collection date: 15.04.2015.
Administrative regions: LA - Houaphan.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Laos
Original label text:
FLORA OF LAOS
Polypodiaceae s.l.
Colysis sp.

Houphan Prov., Hem Distr., Khon Ngua village, around point 19°55'53.2''N 103°16'24.2''E. Secondary broad-leaved evergreen open dry forest with bamboo on very steep rocky mountain slopes composed by solid marble like highly eroded limestone at elev. 700-950 m a.s.l.
Terrestrial fern on steep shady rocky slope. Common.
LA-VN 1487 15 April 2015
Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham, Somchanh Bounphanmy.
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318702 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318702
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LE 01318659

LE 01318659

Cyrtomium sp.⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: Tien Hiep Nguyen; Averyanov, Leonid Vladimirovich; et al. Field collecting number: LA-VN933. Collection date: 03.04.2015.
Administrative regions: LA - Xiangkhouang.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Laos
Original label text:
FLORA OF LAOS
Polypodiaceae s.l.
Cyrtomium sp.

Xiangkhouang Prov., Peak Distr., Khoang village, Pa Khao mt., around point 19°33’02.2’N 103°39’56.3’E. Primary and secondary broad-leaved evergreen dry forest on solid rocky highly eroded limestone at elev. 1200-1450 m a.s.l. Terrestrial and lithophytic fern on very steep slopes and cliffs. Not rare.
LA-VN 933 3 April 2015
Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318659 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318659
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LE 01318658

LE 01318658

Leptochilus sinovietnamica Liang Zhang, N.T.Lu & Li Bing Zhang⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: Tien Hiep Nguyen; Averyanov, Leonid Vladimirovich; et al. Field collecting number: LA-VN1342. Collection date: 11.04.2015.
Administrative regions: LA - Houaphan.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Laos
Original label text:
FLORA OF LAOS
Polypodiaceae s.l.
Colysis sp.

Houphan Prov., Hem Distr., Na Puok village, Pu Tham Nhon mountain around point 20°10'00.0''N 103°24'37.0''E. Primary and secondary broad-leaved evergreen dry forest on very steep rocky mountain slopes composed by solid marble-like highly eroded limestone at elev. 1050-1150 m a.s.l. Terrestrial fern in shady slope near mountain base. Common.
LA-VN 1342 11 April 2015
Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318658 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318658
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LE 01318622

LE 01318622

Grypothrix megacuspis (Baker) S.E.Fawc. & A.R.Sm.⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6662. Collection date: 22.02.2001.
Administrative regions: VN - Tinh Lao Cai.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
THELYPTERIDACEAE
Cyclosorus

Lao Cai
Van Ban District; Liem Phu Municipality;
Khuoi Ngoa Village. Collections on
ridges just west of village.
21°59'31"N 104°19'05"E 323 - 1050 m
In shady wet place. Rachis shiny
black-purple, rhizome and rachis hairs
dark brown-black; rachis shiny
purple-black.
22 February 2001
D.K. Harder, P.K. Loc, L.V. Averyanov,
G.E. Schatz & S. Bodine 6662
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318622 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318622
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LE 01318603

LE 01318603

Abacopteris sp.⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: Averyanov, Leonid Vladimirovich; et al. Field collecting number: VH5392. Collection date: 18.04.2000.
Administrative regions: VN - Kon Tum.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
FLORA OF VIETNAM
Polypodiaceae
Cyclosurus sp.

S. Vietnam, Kontum Prov., Kon Plong Distr., Hieu Municipality, Mang La forest enterprise (14ш39'N, 108ш25'E). Primary broad-leaved wet
evergreen forest on sandstone at elev. 1100-1200 m. Terrestrial fern with fronds up to 2 m lg on steep shady wet slope along forest stream.
Very common.
No VH 5392 19.04.2000
International Botanical Expedition of the U.S.A. National Geographic
Society (grant No 6383-98)
___________________________________________________________
Collectors: L.Averyanov, P.K.Loc, P.H.Hoang, D.X.Du, N.T.Vinh
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318603 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318603
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LE 01318600

LE 01318600

Gymnosphaera podophylla (Hook.) Copel⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: Averyanov, Leonid Vladimirovich; et al. Field collecting number: VH6384. Collection date: 12.05.2000.
Administrative regions: VN - Tinh GJak Lak.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
FLORA OF VIETNAM
Polypodiaceae s.l.
Cyathaea sp. ?

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of
Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary closed mountain broad-leaved evergreen forest on very steep granite slopes along river canyon at elev. 1400 m. Woody fern 2-3 m hg with fronds up to 2-2.5 m lg in shady wet river side slope.
Not rare.
No VH 6384 12.05.2000
International Botanical Expedition of the U.S.A. National Geographic
Society (grant No 6383-98)
___________________________________________________________
Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318600 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318600
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LE 01318599

LE 01318599

Sphaeropteris brunoniana (Hook.) R.M.Tryon⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: Averyanov, Leonid Vladimirovich; et al. Field collecting number: VH6385. Collection date: 12.05.2000.
Administrative regions: VN - Tinh GJak Lak.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
FLORA OF VIETNAM
Polypodiaceae s.l.
Cyathaea sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of
Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary closed mountain broad-leaved evergreen forest on very steep granite slopes along river canyon at elev. 1400 m. Woody fern 6 m hg with fronds up to 4-5 m lg in shady wet river side slope. Not
rare.
No VH 6385 12.05.2000
International Botanical Expedition of the U.S.A. National Geographic
Society (grant No 6383-98)
___________________________________________________________
Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318599 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318599
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LE 01318590

LE 01318590

Lepisorus nudus (Hook.) Ching.⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: Averyanov, Leonid Vladimirovich; et al. Field collecting number: VH6204. Collection date: 09.05.2000.
Administrative regions: VN - Tinh GJak Lak.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
FLORA OF VIETNAM
Polypodiaceae s.l.
Cyclosurus sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of
Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary mossy cloud mixed forest (with Pinus, Fokienia,
Dacrydium and Dacrycarpus) on very steep granite slopes at elev. 2440 m. Creeping epiphyte. Not common.
No VH 6204 09.05.2000
International Botanical Expedition of the U.S.A. National Geographic
Society (grant No 6383-98)
___________________________________________________________
Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318590 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318590
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LE 01318578

LE 01318578

Pseudocyclosorus tylodes (Kunze) Ching.⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: Averyanov, Leonid Vladimirovich; et al. Field collecting number: VH5382. Collection date: 18.04.2000.
Administrative regions: VN - Kon Tum.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
FLORA OF VIETNAM
Polypodiaceae
Athyrium sp.

S. Vietnam, Kontum Prov., Kon Plong Distr., Hieu Municipality, Mang La forest enterprise (14ш39'N, 108ш25'E). Primary broad-leaved wet
evergreen forest on sandstone at elev. 1100-1200 m. Terrestrial fern in open wet slope. Common.
No VH 5382 18.04.2000
International Botanical Expedition of the U.S.A. National Geographic
Society (grant No 6383-98)
___________________________________________________________
Collectors: L.Averyanov, P.K.Loc, P.H.Hoang, D.X.Du, N.T.Vinh
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318578 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318578
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LE 01318561

LE 01318561

Cyrtomium hemionitis Christ⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6181. Collection date: 11.02.2001.
Administrative regions: VN - Tinh Ha Giang.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
DRYOPTERIDACEAE
Cyrtomium

Ha Giang
Quan Ba District; Bat Dai Son
Municipality; Bat Dai Son Provincial
Protected Area (2000); ridge N of Bat
Dai Son border station.
23°09'11"N 104°59'06"E 1287 m
Rare, in deep shade, on rocks. Hair on
rachis and lower shoots brown; sporangia
dark brown.
11 February 2001
D.K. Harder, N.T. Hiep, L.V. Averyanov &
P.K. Loc 6181
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318561 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318561
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LE 01318559

LE 01318559

Abacopteris nudata (Roxb.) S.E.Fawc. & A.R.Sm.⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6395. Collection date: 15.02.2001.
Administrative regions: VN - Tinh Ha Giang.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
THELYPTERIDACEAE
Cyclosorus

Ha Giang
Vi Xuyen District; Phong Quang
Municipality; Man Village. Limestone
peaks and ridges NE of Village.
22°53'21"N 104°56'45"E 595 - 787 m
Occasional, in understory in shade.
Rachis brown; sporangia dark brown.
15 February 2001
D.K. Harder, P.K. Loc, L.V. Averyanov,
G.E. Schatz & S. Bodine 6395
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318559 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318559
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LE 01318555

LE 01318555

Cyrtomium sp.⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6356. Collection date: 15.02.2026.
Administrative regions: VN - Tinh Ha Giang.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
DRYOPTERIDACEAE
Cyrtomium

Ha Giang
Vi Xuyen District; Phong Quang
Municipality; Man Village. Limestone
peaks and ridges NE of Village.
22°53'21"N 104°56'45"E 595 - 787 m
Occasional, among rocks. Hairs on
rhizome and rachis brown, shiny;
sporangia brown; rachis green, shiny.
15 February 2001
D.K. Harder, P.K. Loc, L.V. Averyanov,
G.E. Schatz & S. Bodine 6356
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318555 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318555
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LE 01318548

LE 01318548

Gymnosphaera henryi (Baker) S.R.Ghosh⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6298. Collection date: 17.02.2001.
Administrative regions: VN - Tinh Ha Giang.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
CYATHEACEAE
Cyathea

Ha Giang
Vi Xuyen District; Phong Quang
Municipality; Man Village. Collection
on ridge and small limestone peak just
east of house of village head; very
disturbed, secondary vegetation on lower
slopes and on limestone rocks up to
ridgetop.
22°53'11"N 104°56'43"E 437 - 645 m
22°53'23"N 104°56'48"E
Occasional. On stem to 2 m; rachis dull
14 February 2001
D.K. Harder, P.K. Loc, G.E. Schatz &
L.V. Averyanov 6298
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
----------------------------------------
Page 2
brown; fronds to 2 m long; pinnae dark
green above; sporangia brown.
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318548 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318548
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LE 01318546

LE 01318546

Selliguea obtusa (Ching) S.G.Lu, Hovenkamp & M.G.Gilbert.⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH7489. Collection date: 08.03.2001.
Administrative regions: VN - Tinh Son La.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
POLYPODIACEAE
Crypsinus

Son La
Moc Chau District; Long Luong
Municipality; Sang Cai Village.
Collections in primary, wet,
broad-leaved, evergreen and
semi-deciduous, mossy forest along
limestone ridge situated between
1400-1500 m elevation.
20°45'51"N 104°51'26"E 1300 - 1500 m
Epiphyte fern on mossy trees. Common.
8 March 2001
D.K. Harder, P.K. Loc, L.V. Averyanov &
N.X. Tam 7489
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318546 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318546
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LE 01318543

LE 01318543

Abacopteris sp.⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6607. Collection date: 21.02.2001.
Administrative regions: VN - Tinh Lao Cai.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
THELYPTERIDACEAE
Cyclosorus

Lao Cai
Van Ban District; Liem Phu Municipality;
Khuoi Ngoa Village.
21°58'20"N 104°20'16"E 527 - 1120 m
21°58'08"N 104°21'00"E
Occasional, in shade. Rachis green,
shiny; sporangia brown.
21 February 2001
D.K. Harder, L.V. Averyanov & P.K. Loc
6607
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318543 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318543
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LE 01318541

LE 01318541

Cyclosorus sp.⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6624. Collection date: 21.02.2001.
Administrative regions: VN - Tinh Lao Cai.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
THELYPTERIDACEAE
Cyclosorus

Lao Cai

Van Ban District; Liem Phu Municipality;
Khuoi Ngoa Village.
21°58'20"N 104°20'16"E 527 - 1120 m
21°58'08"N 104°21'00"E
Occasional, in shade. Pinnae dull dark
green; rachis shiny green to darker
brown at base; sporangia brown.
21 February 2001
D.K. Harder, L.V. Averyanov & P.K. Loc
6624
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318541 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318541
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LE 01318539

LE 01318539

Cyrtomium sp.⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH7499. Collection date: 08.03.2001.
Administrative regions: VN - Tinh Son La.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
Pteridophyta J.Y. Bergen & B.M. Davis

Son La
Moc Chau District; Long Luong
Municipality; Sang Cai Village.
Collections in primary, wet,
broad-leaved, evergreen and
semi-deciduous, mossy forest along
limestone ridge situated between
1400-1500 m elevation.
20°45'51"N 104°51'26"E 1300 - 1500 m
Terrestrial and lithophytic fern. Not
8 March 2001
D.K. Harder, P.K. Loc, L.V. Averyanov &
N.X. Tam 7499
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318539 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318539
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LE 01318532

LE 01318532

Cheilanthes tenuifolia (Burm.f.) Sw.⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: Kudryavtzeva, Ekaterina Petrovna; Ogureeva G. Field collecting number: Kudryavtzeva144. Collection date: 29.09.1990.
Administrative regions: VN - Tinh Khanh Hoa.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318532 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318532
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LE 01318531

LE 01318531

Cheilanthes velutina (Tardieu & C.Chr.) Fraser-Jenk.⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: Kudryavtzeva, Ekaterina Petrovna; Ogureeva G. Field collecting number: Kudryavtzeva266. Collection date: 03.10.1990.
Administrative regions: VN - Tinh Khanh Hoa.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318531 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318531
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LE 01318524

LE 01318524

Polystichum sp.⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6141. Collection date: 11.02.2001.
Administrative regions: VN - Tinh Ha Giang.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
DRYOPTERIDACEAE
Cyrtomium

Ha Giang
Quan Ba District; Bat Dai Son
Municipality; Bat Dai Son Provincial
Protected Area (2000); valley just W of
Bat Dai Son border station; valley of
slash and burn agriculture, then higher
up, degraded secondary vegetation in
shade.
23°09'04"N 104°59'53"E 1000 m
Occasional, lithophyitc. Rachis
11 February 2001
D.K. Harder, P.K. Loc, L.V. Averyanov,
G.E. Schatz & S. Bodine 6141
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
----------------------------------------
Page 2
dark-brown; few sporangia dark brown to
black.
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318524 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318524
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LE 01318520

LE 01318520

Alsophila costularis Baker.⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6525. Collection date: 20.02.2001.
Administrative regions: VN - Tinh Lao Cai.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
CYATHEACEAE
Cyathea

Lao Cai
Van Ban District; Liem Pho Municipality;
Khuoi Ngoa Village. Collections in
disturbed secondary vegetation and
cultivated tree plantation, along road
between tree [Fokienia (Pou Nhou)]
extraction camp along road to near Khuoi
Ngoa school.
21°58'14"N 104°19'51"E 606 m
Occasional, in secondary understory.
20 February 2001
D.K. Harder, P.K. Loc, L.V. Averyanov,
G.E. Schatz & S. Bodine 6525
Supported by National Science Foundation
and National Geographic Society
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318520 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318520
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LE 01318514

LE 01318514

Trigonospora tenera (Roxb.) Mazumdar⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6576. Collection date: 21.02.2001.
Administrative regions: VN - Tinh Lao Cai.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
THELYPTERIDACEAE
Cyclosorus

Lao Cai

Van Van District; Liem Phu Municipality;
Khuoi Ngoa Village. Collections near
logging camp [Fokienia (Pou Nhou)] on
road through village; collections from
logging camp and along stream.
21°58'14"N 104°19'51"E 606 m
Occasional, in shade, roots stiff
brown-black; sporangia black to dark
brown.
21 February 2001
D.K. Harder, L.V. Averyanov, P.K. Loc &
G.E. Schatz 6576
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318514 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318514
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LE 01318504

LE 01318504

Sphaeropteris brunoniana (Hook.) R.M.Tryon⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6530. Collection date: 20.02.2001.
Administrative regions: VN - Tinh Lao Cai.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
CYATHEACEAE
Cyathea

Lao Cai
Van Ban District; Liem Pho Municipality;
Khuoi Ngoa Village. Collections in
disturbed secondary vegetation and
cultivated tree plantation, along road
between tree [Fokienia (Pou Nhou)]
extraction camp along road to near Khuoi
Ngoa school.
21°58'14"N 104°19'51"E 606 m
Occasional, tree fern to 2 m tall. Stem
20 February 2001
D.K. Harder, P.K. Loc, L.V. Averyanov,
G.E. Schatz & S. Bodine 6530
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318504 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318504
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LE 01318502

LE 01318502

Lepisorus ovatus (C.Presl) R.Kr.Singh & V.K.Rawat⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6278. Collection date: 12.02.2026.
Administrative regions: VN - Tinh Ha Giang.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
PTERIDACEAE
Coniogramme

Ha Giang
Quan Ba District; Sin Suoi Ho School and
mountain range to the south. Collection
in degraded forest along N slope and
along limestone ridge; limestone
substrate and exposed outcrops in open
and shaded habitat.
23°06'57"N 105°01'48"E 1100 m
Occasional, on lower slopes. Rachis with
dark brown-black hairs, rachis dark
12 February 2001
D.K. Harder, P.K. Loc, N.T. Hiep, George
Schatz & L.V. Averyanov 6278
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
----------------------------------------
Page 2
green; sporangia brown.
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318502 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318502
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LE 01318498

LE 01318498

Isotype of Polystichum hagiangense Li Bing Zhang, N.T.Lu & Liang Zhang,⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6162. Collection date: 11.02.2001.
Administrative regions: VN - Tinh Ha Giang.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
DRYOPTERIDACEAE
Cyrtomium

Ha Giang
Quan Ba District; Bat Dai Son
Municipality; Bat Dai Son Provincial
Protected Area (2000); ridge N of Bat
Dai Son border station.
23°09'11"N 104°59'06"E 1287 m
Rare, on sheer limestone cliff. At base,
hair on fronds brown; sporangia darker
brown to black.
11 February 2001
D.K. Harder, N.T. Hiep, L.V. Averyanov &
P.K. Loc 6162
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318498 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318498
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LE 01318492

LE 01318492

Polystichum sp.⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6356. Collection date: 15.02.2001.
Administrative regions: VN - Tinh Ha Giang.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
DRYOPTERIDACEAE
Cyrtomium

Ha Giang
Vi Xuyen District; Phong Quang
Municipality; Man Village. Limestone
peaks and ridges NE of Village.
22°53'21"N 104°56'45"E 595 - 787 m
Occasional, among rocks. Hairs on
rhizome and rachis brown, shiny;
sporangia brown; rachis green, shiny.
15 February 2001
D.K. Harder, P.K. Loc, L.V. Averyanov,
G.E. Schatz & S. Bodine 6356
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318492 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318492
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LE 01318491

LE 01318491

Abacopteris nudata (Roxb.) S.E.Fawc. & A.R.Sm.⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6806. Collection date: 24.02.2001.
Administrative regions: VN - Tinh Lao Cai.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
THELYPTERIDACEAE
Cyclosorus

Lao Cai
Van Ban District; Nam Xe Municipality.
Collection in small forest patch near
road between Nam Xe Village and Khau Co
Pass.
22°02'26"N 103°59'21"E 612 m
Occasional, on slope near stream. Rachis
dull green; midrib of pinnae
yellow-brown; sporangia dull
brown-black.
24 February 2001
D.K. Harder & et al. 6806
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318491 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318491
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LE 01318487

LE 01318487

Leptochilus vietnamensis Liang Zhang, N.T.Lu & Li Bing Zhang⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH7442. Collection date: 07.03.2001.
Administrative regions: VN - Tinh Son La.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
POLYPODIACEAE
Colysis

Son La
Moc Chau District; Van Ho Municipality,
Hua Tat Village. Collections on
limestone ridge from degraded forests to
cultivation on lower slopes to primary
broadleaved, closed wet forest along
limestone ridge and slopes near tops.
20°46'13"N 104°46'12"E 1100 - 1300 m
Epiphyte. Not common.
7 March 2001
D.K. Harder, P.K. Loc, L.V. Averyanov &
N.X. Tam 7442
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318487 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318487
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LE 01318485

LE 01318485

Gymnosphaera sp.⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: Averyanov, Leonid Vladimirovich; et al. Field collecting number: VH6250. Collection date: 07.05.2000.
Administrative regions: VN - Tinh GJak Lak.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
FLORA OF VIETNAM
Polypodiaceae s.l.
Cyathaea sp. ?

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of
Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary closed wet broad-leaved and mixed forest (with Pinus,
Fokienia, Dacrydium and Dacrycarpus) on steep granite slopes at elev. 1200-1550 m. Woody fern up to 4 m hg and fronds up to 2.5 m lg on
shady very steep slope of river canyon. Not common.
No VH 6250 07.05.2000
International Botanical Expedition of the U.S.A. National Geographic
Society (grant No 6383-98)
___________________________________________________________
Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318485 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318485
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LE 01318474

LE 01318474

Cyclosorus sp.⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6724. Collection date: 22.02.2001.
Administrative regions: VN - Tinh Lao Cai.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
THELYPTERIDACEAE
Cyclosorus

Lao Cai
Van Ban District; Liem Phu Municipality;
Khuoi Ngoa Village. Collections on
ridges just west of village.
21°59'31"N 104°19'05"E 323 - 1050 m
Frequent, in shade all along ridge in
shade. Rachis light brown to dark green;
sporangia brown to orange brown.
22 February 2001
D.K. Harder, P.K. Loc, L.V. Averyanov,
G.E. Schatz & S. Bodine 6724
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318474 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318474
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LE 01318452

LE 01318452

Glaphyropteridopsis erubescens (Wall. ex Hook.) Ching⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH7132. Collection date: 02.03.2001.
Administrative regions: VN - Tinh Son La.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
Pteridophyta J.Y. Bergen & B.M. Davis

Son La
Yen Chau District; Muong Lum
Municipality; Lum Village. Collections
on ridge S of village. Collected with
P.K. Loc, N.T. Hiep, L.V. Averyanov,
G.E. Schatz, S. Bodine & N.Q. Hieu.
21°00'47"N 104°29'30"E 900 - 1134 m
Occasional, terrestrial. Frond to 1.5 m;
lower rachis dark brown; sporangia dark
brown; lower-most pinnae pendulous.
2 March 2001
D.K. Harder & et al. 7132
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318452 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318452
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LE 01318451

LE 01318451

Sphaeropteris brunoniana (Hook.) R.M.Tryon⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6302. Collection date: 14.02.2001.
Administrative regions: VN - Tinh Ha Giang.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
CYATHEACEAE
Cyathea

Ha Giang
Vi Xuyen District; Phong Quang
Municipality; Man Village. Collection
on ridge and small limestone peak just
east of house of village head; very
disturbed, secondary vegetation on lower
slopes and on limestone rocks up to
ridgetop.
22°53'11"N 104°56'43"E 437 - 645 m
22°53'23"N 104°56'48"E
Occasional, in understory. Rachis light
14 February 2001
D.K. Harder, P.K. Loc, G.E. Schatz &
L.V. Averyanov 6302
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
----------------------------------------
Page 2
yellow, green to brown; sporangia brown.
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318451 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318451
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LE 01318448

LE 01318448

Dryopteris sp.⁣ det. Cheng-Wei Chen at 14.02.2026

Collectors: D.K. Harder; et al. Field collecting number: DKH6889. Collection date: 26.02.2001.
Administrative regions: VN - Tinh Lao Cai.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
THELYPTERIDACEAE
Cyclosorus

Lao Cai
Van Ban District; Nam Xe Municipality;
Nam Xi Tan Village. Collection in forest
on steep W-facing slope along stream;
just above village; on N side of road
between Nam Xe and Khau Co Pass at near
40 km to Than Uyen mark.
22°02'31"N 103°58'32"E 700 - 1113 m
Occasional, in shade. Rachis darker
brown-black; sporangia brown; rachis
26 February 2001
D.K. Harder, P.K. Loc, N.T. Hiep, G.E.
Schatz, S. Bodine & N.Q. Hieu 6889
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
----------------------------------------
Page 2
hairs dark brown-black.
Record creation: 2026-02-17, Galina Savich, PhotoScan D2
Citation: Specimen LE 01318448 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318448
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LE 01323861

LE 01323861

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-17, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01323861 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01323861
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LE 01323860

LE 01323860

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-17, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01323860 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01323860
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LE 01323859

LE 01323859

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-17, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01323859 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01323859
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LE 01323858

LE 01323858

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-17, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01323858 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01323858
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LE 01323856

LE 01323856

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-17, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01323856 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01323856
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LE 01323857

LE 01323857

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-17, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01323857 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01323857
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LE 01323855

LE 01323855

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-17, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01323855 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01323855
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LE 01323854

LE 01323854

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-17, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01323854 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01323854
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LE 01323853

LE 01323853

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-17, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01323853 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01323853
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LE 01323852

LE 01323852

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-17, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01323852 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01323852
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LE 01323850

LE 01323850

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-17, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01323850 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01323850
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LE 01323851

LE 01323851

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-17, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01323851 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01323851
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LE 01323849

LE 01323849

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-17, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01323849 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01323849
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LE 01323848

LE 01323848

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-17, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01323848 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01323848
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LE 01323847

LE 01323847

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-17, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01323847 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01323847
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LE 01323846

LE 01323846

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-17, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01323846 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01323846
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LE 01323845

LE 01323845

Groups of specimens: Образцы сектора Средней Азии; Типовые образцы сектора Средней Азии
Record creation: 2026-02-17, Mariya Sheludyakova, PhotoScan D2
Citation: Specimen LE 01323845 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01323845
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