Herbarium LE - Specimens

Rubiaceae

Ixora sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Undershrub to shrum 0.5-1.5 m tall in open place along trail. Flowers bright red. Very common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4375. Photo only.

Pandanaceae

Pandanus tectorius Parkinson

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Sublittoral palm-like shrub to 3 m tall along rocky seashore line among open limestone and sandstone boulders. Fruits orange to orange-red. Very common

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4374. Photo only.

Myrsinaceae

Aegiceras corniculatum (L.) Blanco

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Littoral mangrove shrub to 1.5 m tall in flooded littoral zone. Flowers white. Very common. Main dominant of mangrove plant community.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4349 (NHC149).

Theaceae

Camellia oleifera C.Abel

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Treelet 7-8 m tall and about 20 cm DBH along trail. Flower buds light green, flowers white. Not common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4343 (NHC143).

Sapindaceae

Nephelium lappaceum L.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Tree to 15 m tall along trail. Fruits green. Not common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4342 (NHC142).

Verbenaceae

Clerodendrum japonicum (Thunb.) Sweet

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Undershrub to 2 m tall in open rocky place along trail. Flowers bright red. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4341 (NHC141).

Euphorbiaceae

Antidesma fordii Hemsl.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Treelet 4-6 m tall in open place along trail. Flowers light greenish. Not rare.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4340 (NHC140).

Myrsinaceae

Ardisia sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Treelet to 2.5 m tall in open place along trail. Flowers pink-purple. Occasional.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4339 (NHC139).

Vitaceae

Cayratia sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Herbaceous vine to 3 m long in open place along trail. Young fruits green. Not rare.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4338 (NHC138).

Fabaceae s.l. (Caesalpiniaceae)

Peltophorum pterocarpum (DC.) Backer ex K.Heyne

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Tree to 20 m tall and 0.8 m DBH. Flowers bright yellow. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4337 (NHC137).

Rubiaceae

Ixora finlaysoniana Wall.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Shrub to 1.5 m tall in open place along trail. Flowers white. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4336 (NHC136).

Sapindaceae

Xerospermum noronhianum (Blume) Blume

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Tree to 20 m tall. Flowers white to yellowish. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4335 (NHC135).

Pandaceae

Microdesmis caseariifolia Planch. ex Hook.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Treelet to 6-7 m tall in open place along trail. Flowers bright yellow. Not common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4334 (NHC134).

Rutaceae

Micromelum falcatum (Lour.) Tanaka

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Shrub or small treelet to 3 m tall in open place along trail. Fruits orange to red. Very common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4333 (NHC133).

Apocynaceae

Tabernaemontana bufalina Lour.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Shrub to 3 m tall in open place along trail. Flowers white, fruits orange. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4332 (NHC132).

Annonaceae

Uvaria littoralis (Blume) Blume

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Woody vine to 10 m long. Flowers bright red. Not common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4331 (NHC131).

Fabaceae

Baphia capparidifolia Baker

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Scandent shrub with shoots to 3 m long in open place. Flowers white. Not common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4330 (NHC130).

Fabaceae s.l. (Mimosaceae)

Entada phaseoloides (L.) Merr.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Woody vine to 25 m long on shady slope. Not rare.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4329 (NHC129).

Oleaceae

Myxopyrum smilacifolium (Wall.) Blume

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Woody vine to 5 m long in open places along trail. Medicinal plant. Locally common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4328 (NHC128).

Rosaceae

Rubus leucanthus Hance

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Climbing shrub with shoots to 2 m long in open places along trail. Flowers white. Very common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4327 (NHC127).

Sauraiaceae

Saurauia tristyla DC.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Treelet to 6 m tall in open wet rocky place. Flowers light pink. Occasional.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4326 (NHC126).

Fabaceae s.l. (Caesalpiniaceae)

Cassia sp.?

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Woody vine to 10 m long in open place. Flowers bright yellow. Not cmmon.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4325 (NHC125).

Moraceae

Streblus sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Tree to 10 m tall in shady place. Fruits dull green. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4324 (NHC124).

Polypodiaceae s.l.

Leptochilus sp.?

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 100-150 m a.s.l. along forest trail. Creeping lithophytic fern in shady place. Not rare.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4323 (NHC123).

Euphorbiaceae

Antidesma ghaesembilla Gaertn.?

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Treelet or shrub to 3 m tall in rather open place. Flowers greenish. Common.

Anacardiaceae

Allospondias lakonensis (Pierre) Stapf

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Tree about 15 m tall in rather open place. Flowers white. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4320 (NHC121).

Simaroubaceae

Eurycoma longifolia Jack

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Treelet to 4 m tall in rather open place. Flowers purplish. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4319 (NHC120).

Apocynaceae

Tabernaemontana pauciflora Blume

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Treelet to 4 m tall in rather open place. Flowers white. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4318 (NHC119).

Orchidaceae

Tropidia angulosa (Lindl.) Blume

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Terrestrial herb to 0.8 m tall in shady place. Not common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4317 (NHC118).

Gnetaceae

Gnetum latifolium?

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Woody vine to 30 m long, with stem to 20 cm in diameter. Not rare.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4316 (NHC117).

Rubiaceae

Psychotria sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Terrestrial undershrub to 1.5 m tall. Flowers olive greenish. Not rare.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4315 (NHC116).

Selaginellaceae

Selaginella sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Terrestrial herb to 0.4 m tall in rather open place. Locally common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4314 (NHC115).

Hypoxidaceae

Curculigo sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Terrestrial rosulate herb in rather open place. Flowers yellow. Locally common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4313 (NHC114).

Zingiberaceae

Alpinia oblongifolia Hayata

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Terrestrial herb to 1 m tall in rather open place. Flowers white, median tepal with 2 red stripes. Locally common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4312 (NHC113).

Aquifoliaceae

Ilex purpurea?

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Treelet 6-8 m tall in open place. Flowers dirty violet. Occasional.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4311 (NHC112).

Rubiaceae

Randia sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Treelet 5-6 m tall in open place. Flowers green. Occasional.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4310 (NHC111).

Euphorbiaceae

Aporosa dioica (Roxb.) Müll.Arg.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Tree 6-8 m tall in open place. Young fruits light green. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4309 (NHC110).

Clusiaceae

Garcinia sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Tree 6-8 m tall in open place. Flowers yellow. Occasional.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4308 (NHC109).

Apocynaceae

Melodinus suaveolens (Hance) Champ. ex Benth.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Scandent shrub or vine to 5 m long with milky juice in open place. Flowers white. Locally common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4307 (NHC108).

Caprifoliaceae

Lonicera sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Littoral vine to 5 long along sea shore. Flowers white, later pale yellow. Not common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4306 (NHC107).

Verbenaceae

Clerodendron sp.?

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Littoral climbing shrub with scandent shoot to 3 long along sea shore. Flowers white. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4305 (NHC106).

Goodeniaceae

Scaevola taccada (Gaertn.) Roxb.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Littoral shrub to 3 m tall along sea shore. Flowers white. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4304 (NHC105).

Rhizophoraceae?

Bruguiera gymnorhiza (L.) Lam. ex Savigny

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Mangrove scrub in flooded littoral zone along sea shore. Treelet to 3 m tall. Old flowers reddish, fruits dark green. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4303 (NHC104).

Rhizophoraceae

Ceriops tagal (Perr.) C.B.Rob.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Mangrove scrub in flooded littoral zone along sea shore. Treelet or shrub to 2 m tall. Flower buds yellowish with red. Common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4302 (NHC103).

Convallariaceae

Aspidistra quadrangularis Aver.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Terrestrial rhizomatous herb in shady place. Young fruits light green, quadrangular. Locally common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4301 (NHC102).

Convallariaceae

Peliosanthes sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone slope at elevation about 50 m a.s.l. along forest trail. Terrestrial clustering herb in shady place. Young fruits light green. Locally common.

16 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4300 (NHC101).

Orchidaceae

Anoectochilus roxburghii (Wall.) Lindl.

Hanoi Capital Region, Ba Vi National Park, humid mossy primary and secondary evergreen broad-leaved forest on steep mountain slopes composed by solid shale at elevation of 1100-1300 m a.s.l. along trail to peak of 1296 m. Terrestrial or lithophytic herb on shady slope. Leaves drk green with yellowish to pink nerves. Rare.

11 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4297 (NHC099).

Orchidaceae

Epipogium roseum (D.Don) Lindl.

Hanoi Capital Region, Ba Vi National Park, humid mossy primary and secondary evergreen broad-leaved forest on steep mountain slopes composed by solid shale at elevation of 1100-1300 m a.s.l. along trail to peak of 1296 m. Terrestrial tuberiferous achlorophyllous white to light yellowish herb on shady slope. Rare.

11 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4296 (NHC098).

Polypodiaceae s.l.

Microsorus sp.?

Hanoi Capital Region, Ba Vi National Park, humid mossy primary and secondary evergreen broad-leaved forest on steep mountain slopes composed by solid shale at elevation of 1100-1300 m a.s.l. along trail to peak of 1296 m. Creeping epiphytic vine on tall mossy tree. Common.

11 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4295 (NHC097).

Thymellaeaceae

Wikstroemia indica (L.) C.A. Mey.

Hanoi Capital Region, Ba Vi National Park, humid mossy primary and secondary evergreen broad-leaved forest on steep mountain slopes composed by solid shale at elevation of 1100-1300 m a.s.l. along trail to peak of 1296 m. Shrub or small treelet to 1.5 m tall on open rocky steep slope. Fruits orange red to red. Not common.

11 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4294 (NHC096).