Herbarium LE - Specimens

Anthericaceae

Chlorophytum sp.

Bac Kan Province, Ba Be District, Nam Mau Commune, Nam Dai Village. Dry evergreen broad-leaved forest on steep rocky slopes composed by white to gray crystalline marble-like limestone around point 22º25’20’’N 105º35’03’E at elev. of about 650 m a.s.l. Ascending herb to 0.5 m tall on rocky shady wet slope. Not common.

18 October 2023. Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, Nguyen Van Ly, T. Maisak, AL2302.

Apiaceae

Sanicula elata D.Do

Lam Dong Province, Lac Duong District, Bidoup Nui Ba National Par, southern slopes of Bidoup Mountain about 5 km to the south from Bidoup peak around point 12º04.076’N 108º38.927E. Open grassy Pinus kesiya forest and woodlands at elevation about 1900 m a.s.l. Erect terrestrial herb to 0.4 m tall. Not common.

08 November 2023, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, T. Maisak, AL2506.

Apiaceae

Hydrocotyle nepalensis Hook.⁣

Lam Dong Province, Lac Duong District, Bidoup Nui Ba National Par, southern slopes of Bidoup Mountain about 5 km to the south from Bidoup peak around point 12º04.076’N 108º38.927E. Open grassy Pinus kesiya forest and woodlands at elevation about 1900 m a.s.l. Creeping terrestrial herb in open wet place. Common.

08 November 2023, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, T. Maisak, AL2504.

Apiaceae

Hydrocotyle tonkinensis Tardieu⁣

Bac Kan Province, Ba Be District, Nam Mau Commune, Nam Dai Village. Dry evergreen broad-leaved forest on steep rocky slopes composed by white to gray crystalline marble-like limestone around point 22º25’20’’N 105º35’03’E at elev. of about 650 m a.s.l. Terrestrial creeping herb in wet open place. Common.

18 October 2023. Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, Nguyen Van Ly, T. Maisak, AL2305.

Araceae

Typhonium hangiae V.D.Nguyen, D.D.Nguyen & V.C.Nguyen

Isotype

Quang Nam Province, Nong Son District, evergreen broad-leaved forest at elevation of 200-300 m a.s.l. along stream. Spathe dull brown. Not common.

21 January 2022, Nguyen Thi Hang 22.226. AL2474.

Araceae

Acorus gramineus Aiton

Lam Dong Province, Di Linh District, Brah Yang Mountain, primary and old secondary evergreen broad-leaved and mixed forest (with Pinus khasya) on granite around point 11º27.669ʹ N 108º03.543ʹ E at elevation of 1000-1100 m a.s.l.

Lithophytic rhizomatous herb on wet rocks along small forest stream. Locally very common.

14 November 2023, L. Averyanov, Nguyen Van Canh, T. Maisak, AL2691.

Araliaceae

Dendropanax sp.

Lam Dong Province, Lac Duong District, Bidoup Nui Ba National Par, southern slopes of Bidoup Mountain about 5 km to the south from Bidoup peak around point 12º04.076’N 108º38.927E. Primary broad-leaved evergreen forest on steep granite slopes at elevation 2000–2200 m a.s.l. Treelet about 1.5 m tall. Not common.

08 November 2023, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, T. Maisak, AL2532 (BD 039).

Araliaceae

Heteropanax sp.

Lam Dong Province, Lac Duong District, Bidoup Nui Ba National Par, southern slopes of Bidoup Mountain about 5 km to the south from Bidoup peak around point 12º04.076’N 108º38.927E. Primary broad-leaved evergreen forest on steep granite slopes at elevation 2000–2200 m a.s.l. Tree 15-20 m tall. Flowers yellowish. Occasional.

08 November 2023, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, T. Maisak, AL2533 (BD 040).

Asclepiadaceae

Hoya thuathienhuensis T.B.Tran, Rodda & Simonsson

Quang Nam Province, Nong Son District, evergreen forest at elevation at about 300 m a.s.l. February 2022, Nguyen Van Canh,

Herbarium collected in Nguyen Van Canh Garden at 3 November, by Nguyen Van Canh, L. Averyanov, T. Maisak, AL2425.

Asclepiadaceae

Hoya sp. (H. sp. nov., H. multiflora aff.)

Dak Lak Province, M’Drak District, Chu Mu Mountain, evergreen broad-leaved forest at elevation of about 900 m a.s.l. Flowers greenish. Very rare.

April 2023, Nguyen Van Canh, AL2470.

Asclepiadaceae

Hoya fusca Wall.

Dak Lak Province, Lak District, evergreen forest at elevation of about 1200 m a.s.l., not common. Flowers light yellow green.

21 May 2023 by Nguyen Van Canh, AL2463.

Bac Kan Province, Ba Be District, Ba Be National Park. Dry evergreen broad-leaved secondary forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone along lake shore on point 22º24’58’’N 105º36’48’E at elev. of about 250 m a.s.l.

29 October 2023, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, Nguyen Van Ly, T. Maisak; Nguyen Van Ly 20231029 155.

Asteraceae

Gnaphalium sp.

Lam Dong Province, Lac Duong District, Bidoup Nui Ba National Par, southern slopes of Bidoup Mountain about 5 km to the south from Bidoup peak around point 12º04.076’N 108º38.927E. Open grassy Pinus kesiya forest and woodlands at elevation about 1700–1900 m a.s.l. Erect terrestrial herb to 0.4 m tall. Very common.

08 November 2023, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, T. Maisak, AL2554.

Balanophoraceae

Balanophora indica?

Bac Kan Province, Ba Be District, Nam Mau Commune, Nam Dai Village. Dry evergreen broad-leaved forest on steep rocky slopes composed by white to gray crystalline marble-like limestone around point 22º25’20’’N 105º35’03’E at elev. of about 650 m a.s.l. Achlorophyllous ground red parasite on steep shady rocky slope. Not common. Male specimen.

18 October 2023. Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, Nguyen Van Ly, T. Maisak, AL2316 (same 2317).

Balanophoraceae

Balanophora indica?

Bac Kan Province, Ba Be District, Nam Mau Commune, Nam Dai Village. Dry evergreen broad-leaved forest on steep rocky slopes composed by white to gray crystalline marble-like limestone around point 22º25’20’’N 105º35’03’E at elev. of about 650 m a.s.l. Achlorophyllous ground red parasite on steep shady rocky slope. Not common. Male specimen.

18 October 2023. Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, Nguyen Van Ly, T. Maisak, AL2316 (same 2317).

Balsaminaceae

Impatiens sp.

Bac Kan Province, Ba Be District, Nam Mau Commune, Nam Dai Village. Dry evergreen broad-leaved forest on steep rocky slopes composed by white to gray crystalline marble-like limestone around point 22º25’20’’N 105º35’03’E at elev. of about 650 m a.s.l. Lithophytic and terrestrial herb among shady rocks. Leaves dark green, with white medial nerve, flowers white outside, dark purple inside. Locally common.

17 October 2023. Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, Nguyen Van Ly, T. Maisak, AL2266.

Balsaminaceae

Impatiens sp.

Bac Kan Province, Ba Be District, Ba Be National Park. Dry evergreen broad-leaved secondary forest on cliffs composed by white crystalline marble-like highly eroded limestone along lake shore on point 22º24’58’’N 105º36’48’E at elev. of about 160 m a.s.l. Erect lithophytic herb to 0.5 m tall on exposed rocks. Flowers yellow. Locally very common.

21 October 2023, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, Nguyen Van Ly, T. Maisak, AL2349.

Balsaminaceae

Impatiens coerulea sp. nov.?

Dak Lak Province, Lac District, southern part of Nam Kar Nature Reserve around point 12º16’10’’N 108º05’24’’E at elevations of about 600 m a.s.l. Open steep rocky slope in secondary evergreen broad-leaved forest with bamboo along road.

Lithophytic herb with fleshy stem to 0.8 m tall among large rocks. Flowers bluish with yellow. Locally very common.

05 November 2023, L. Averyanov, Nguyen Van Canh, T. Maisak, AL2481.

Balsaminaceae

Impatiens coerulea sp. nov.?

Dak Lak Province, Lac District, southern part of Nam Kar Nature Reserve around point 12º16’10’’N 108º05’24’’E at elevations of about 600 m a.s.l. Open steep rocky slope in secondary evergreen broad-leaved forest with bamboo along road.

Lithophytic herb with fleshy stem to 0.8 m tall among large rocks. Flowers bluish with yellow. Locally very common.

05 November 2023, L. Averyanov, Nguyen Van Canh, T. Maisak, AL2481.

Balsaminaceae

Impatiens coerulea sp. nov.?

Dak Lak Province, Lac District, southern part of Nam Kar Nature Reserve around point 12º16’10’’N 108º05’24’’E at elevations of about 600 m a.s.l. Open steep rocky slope in secondary evergreen broad-leaved forest with bamboo along road.

Lithophytic herb with fleshy stem to 0.8 m tall among large rocks. Flowers bluish with yellow. Locally very common.

05 November 2023, L. Averyanov, Nguyen Van Canh, T. Maisak, AL2481.

Balsaminaceae

Impatiens coerulea sp. nov.?

Dak Lak Province, Lac District, southern part of Nam Kar Nature Reserve around point 12º16’10’’N 108º05’24’’E at elevations of about 600 m a.s.l. Open steep rocky slope in secondary evergreen broad-leaved forest with bamboo along road.

Lithophytic herb with fleshy stem to 0.8 m tall among large rocks. Flowers bluish with yellow. Locally very common.

05 November 2023, L. Averyanov, Nguyen Van Canh, T. Maisak, AL2481.

Balsaminaceae

Impatiens sp.

Lam Dong Province, Lac Duong District, Bidoup Nui Ba National Par, southern slopes of Bidoup Mountain about 5 km to the south from Bidoup peak around point 12º04.076’N 108º38.927E. Primary broad-leaved evergreen mossy forest along main ridge at elevation 2100–2200 m a.s.l. Terrestrial herb to 0.5 m tall in wet open place. Flowers purple. Locally common.

08 November 2023, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, T. Maisak, AL2547 (BD 054).

Balsaminaceae

Impatiens sp.

Bac Kan Province, Ba Be District, Nam Mau Commune, Nam Dai Village. Dry evergreen broad-leaved forest on steep rocky slopes composed by white to gray crystalline marble-like limestone around point 22º25’20’’N 105º35’03’E at elev. of about 650 m a.s.l. Erect terrestrial herb in rocky open place along stream. Flowers yellow. Locally common.

18 October 2023. Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, Nguyen Van Ly, T. Maisak, AL2309.

Balsaminaceae

Impatiens sp.

Bac Kan Province, Ba Be District, Nam Mau Commune, Nam Dai Village. Dry evergreen broad-leaved forest on steep rocky slopes composed by white to gray crystalline marble-like limestone around point 22º25’20’’N 105º35’03’E at elev. of about 650 m a.s.l. Erect terrestrial herb in rocky open place along stream. Flowers yellow. Locally common.

18 October 2023. Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, Nguyen Van Ly, T. Maisak, AL2309.

Balsaminaceae

Impatiens sp.

Bac Kan Province, Ba Be District, Nam Mau Commune, Nam Dai Village. Dry evergreen broad-leaved forest on steep rocky slopes composed by white to gray crystalline marble-like limestone around point 22º25’20’’N 105º35’03’E at elev. of about 650 m a.s.l. Erect herb in rocky open wet place along stream. Flowers yellow. Locally common.

18 October 2023. Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, Nguyen Van Ly, T. Maisak, AL2311.

Begoniaceae

Begonia sp.

Wild collected plant obtained by Nguyen Van Canh from street market of northern Vietnam in 2021.

Herbarium collected in Nguyen Van Canh Garden at 3 November, by Nguyen Van Canh, L. Averyanov, T. Maisak, AL2427.

Begoniaceae

Begonia sp.

Wild collected plant obtained by Nguyen Van Canh from street market of northern Vietnam in 2021.

Herbarium collected in Nguyen Van Canh Garden at 3 November, by Nguyen Van Canh, L. Averyanov, T. Maisak, AL2428.

Begoniaceae

Begonia sp.

Wild collected plant obtained by Nguyen Van Canh from street market of northern Vietnam in 2021.

Herbarium collected in Nguyen Van Canh Garden at 3 November, by Nguyen Van Canh, L. Averyanov, T. Maisak, AL2426.

Begoniaceae

Begonia sp.

Wild collected plant obtained by Nguyen Van Canh from street market of northern Vietnam in 2021.

Herbarium collected in Nguyen Van Canh Garden at 3 November, by Nguyen Van Canh, L. Averyanov, T. Maisak, AL2429.

Begoniaceae

Begonia kimlongii V.C.Nguyen, T.S.Hoang & C.W.Lin

Dak Lak Province, M’Drak District, Chu Mu Mountain, evergreen forest at elevation of about 600 m a.s.l.

Herbarium collected in Nguyen Van Canh Garden at 3 November, by Nguyen Van Canh, L. Averyanov, T. Maisak, AL2430.

Begoniaceae

Begonia sp.

Dak Lak Province, Ea’Hleo District, Dipterocarp forest at elevation of about 300 m a.s.l., March 2021, Nguyen Van Canh.

Herbarium collected in Nguyen Van Canh Garden at 3 November, by Nguyen Van Canh, L. Averyanov, T. Maisak, AL2431.

Begoniaceae

Begonia jubar V.T.Pham & C.W.Lin

Dak Lak Province, Lac District, southern part of Nam Kar Nature Reserve around point 12º16’10’’N 108º05’24’’E at elevations of about 600 m a.s.l. Open steep rocky slope in secondary evergreen broad-leaved forest with bamboo along road.

Lithophytic herb among large rocks. Flowers pink. Locally very common. Collected in locus classicus.

05 November 2023, L. Averyanov, Nguyen Van Canh, T. Maisak, AL2477.

Begoniaceae

Begonia jubar V.T.Pham & C.W.Lin

Dak Lak Province, Lac District, southern part of Nam Kar Nature Reserve around point 12º16’10’’N 108º05’24’’E at elevations of about 600 m a.s.l. Open steep rocky slope in secondary evergreen broad-leaved forest with bamboo along road.

Lithophytic herb among large rocks. Flowers pink. Locally very common. Collected in locus classicus.

05 November 2023, L. Averyanov, Nguyen Van Canh, T. Maisak, AL2477.

Begoniaceae

Begonia sp.

Bac Kan Province, Ba Be District, Nam Mau Commune, Nam Dai Village. Dry evergreen broad-leaved forest on steep rocky slopes composed by white to gray crystalline marble-like limestone around point 22º25’20’’N 105º35’03’E at elev. of about 650 m a.s.l. Lithophytic herb on shady vertical cliff. Leaves uniform light green. Flowers white. Locally common.

17 October 2023. Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, Nguyen Van Ly, T. Maisak, AL2265.

Begoniaceae

Begonia sp.

Lam Dong Province, Lac Duong District, Bidoup Nui Ba National Par, southern slopes of Bidoup Mountain about 5 km to the south from Bidoup peak around point 12º04.076’N 108º38.927E. Primary broad-leaved evergreen forest on steep granite slopes at elevation 2000–2200 m a.s.l. Terrestrial herb to 1 m tall in wet open place. Leaves uniform green. Flowers white to light pink. Common.

08 November 2023, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, T. Maisak, AL2541 (BD 048).

Begoniaceae

Begonia sp.

Lam Dong Province, Lac Duong District, Bidoup Nui Ba National Par, southern slopes of Bidoup Mountain about 5 km to the south from Bidoup peak around point 12º04.076’N 108º38.927E. Primary broad-leaved evergreen forest on steep granite slopes at elevation 2000–2200 m a.s.l. Terrestrial herb to 1 m tall in wet open place. Leaves uniform green. Flowers white to light pink. Common.

08 November 2023, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, T. Maisak, AL2541 (BD 048).

Boraginaceae

Ehretia asperula Zoll. & Moritzi

Bac Kan Province, Ba Be District, Nam Mau Commune, Nam Dai Village. Dry evergreen broad-leaved forest on steep rocky slopes composed by white to gray crystalline marble-like limestone around point 22º25’20’’N 105º35’03’E at elev. of about 650 m a.s.l. Woody vine to 4-5 m long in rather open slope. Fruits dull orange. Not common.

17 October 2023. Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, Nguyen Van Ly, T. Maisak, AL2270.

Campanulaceae

Campanula sp. (vel Adenophora sp.)?

Lam Dong Province, Lac Duong District, Bidoup Nui Ba National Par, southern slopes of Bidoup Mountain about 5 km to the south from Bidoup peak around point 12º04.076’N 108º38.927E. Open grassy Pinus kesiya forest and woodlands at elevation about 1900 m a.s.l. Erect terrestrial herb to 30 cm tall. Flowers blue. Occasional.

08 November 2023, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, T. Maisak, AL2502 (BD 009).

Cardiopteridaceae

Cardiopteris quinqueloba Hassk.⁣

Lam Dong Prov.

AL 2496a, 08.11.2023

Averyanov L. et al.

Cardiopteridaceae

Cardiopteris quinqueloba (Hassk.) Hassk.

Bac Kan Province, Ba Be District, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’26’’N 105º36’12’E at elev. of 600-800 m a.s.l. Herbaceous vine 3-4 m long with milky juice in open rocky place. Not common.

30 October 2023, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, Nguyen Van Ly, T. Maisak, AL2375.

Cardiopteridaceae

Cardiopteris quinqueloba (Hassk.) Hassk.

Bac Kan Province, Ba Be District, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’26’’N 105º36’12’E at elev. of 600-800 m a.s.l. Herbaceous vine 3-4 m long with milky juice in open rocky place. Not common.

30 October 2023, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, Nguyen Van Ly, T. Maisak, AL2375.

Chloranthaceae

Sarcandra sp.

Lam Dong Province, Lac Duong District, Bidoup Nui Ba National Par, southern slopes of Bidoup Mountain about 5 km to the south from Bidoup peak around point 12º04.076’N 108º38.927E. Primary broad-leaved evergreen forest on steep granite slopes at elevation about 2000 m a.s.l. Shrub to 2 m tall in rather open place. Not rare.

08 November 2023, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, T. Maisak, AL2513 (BD 020).

Commelinaceae

Pollia sp.?

Bac Kan Province, Ba Be District, Nam Mau Commune, Nam Dai Village. Dry evergreen broad-leaved forest on steep rocky slopes composed by white to gray crystalline marble-like limestone around point 22º25’20’’N 105º35’03’E at elev. of about 650 m a.s.l. Terrestrial herb in humid valley between limestone hills. Rare.

18 October 2023. Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, Nguyen Van Ly, T. Maisak, AL2275.

Commelinaceae

Floscopa scandens Lour.

Bac Kan Province, Ba Be District, Nam Mau Commune, Nam Dai Village. Dry evergreen broad-leaved forest on steep rocky slopes composed by white to gray crystalline marble-like limestone around point 22º25’20’’N 105º35’03’E at elev. of about 650 m a.s.l. Terrestrial ascending herb in wet place and along stream. Flowers pink. Very common.

17 October 2023. Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, Nguyen Van Ly, T. Maisak, AL2267.

Commelinaceae

Cyanotis sp.?

Lam Dong Province, Lac Duong District, Bidoup Nui Ba National Par, southern slopes of Bidoup Mountain about 5 km to the south from Bidoup peak around point 12º04.076’N 108º38.927E. Open grassy Pinus kesiya forest and woodlands at elevation about 1700–1900 m a.s.l. Prostrate herb in open places. Flowers blue. Very common.

08 November 2023, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, T. Maisak, AL2552.

Commelinaceae

Commelina sp. s.l.

Bac Kan Province, Ba Be District, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’26’’N 105º36’12’E at elev. of 600-800 m a.s.l. Erect lithophytic herb with erect stems to 3 m long, branching and creeping in apical part in shady rocks along ridge edge. Not ripe fruits glossy green. Not common.

30 October 2023, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, Nguyen Van Ly, T. Maisak, AL2395.

Commelinaceae

Commelina sp. s.l.

Bac Kan Province, Ba Be District, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’26’’N 105º36’12’E at elev. of 600-800 m a.s.l. Erect lithophytic herb with erect stems to 3 m long, branching and creeping in apical part in shady rocks along ridge edge. Not ripe fruits glossy green. Not common.

30 October 2023, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, Nguyen Van Ly, T. Maisak, AL2395.

Commelinaceae

Pollia sp.

Bac Kan Province, Ba Be District, Ba Be National Park. Dry evergreen broad-leaved secondary forest on steep rocky slopes composed by white crystalline marble-like highly eroded limestone near Dau Dang Waterfall of Nang River around point 22º27’09’’N 105º34’16’E at elev. of about 150 m a.s.l. Terrestrial erect herb to 1 m tall in wet open place. Flowers white, fruits bluish. Very common.

20 October 2023, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, Nguyen Van Ly, T. Maisak, AL2323.

Commelinaceae

Bac Kan Province, Ba Be District, Nam Mau Commune, Nam Dai Village. Dry evergreen broad-leaved forest on steep rocky slopes composed by white to gray crystalline marble-like limestone around point 22º25’20’’N 105º35’03’E at elev. of about 650 m a.s.l. Terrestrial ascending herb in wet open place. Not rare.

18 October 2023. Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, Nguyen Van Ly, T. Maisak, AL2307.

Cycadaceae

Cycas brachycantha K.D.Hill, H.T.Nguyen & P.K.Loc

Male plant cultivated in Ba Be National park originated from Bac Kan Province, Ba Be District, Ba Be National Park. Rocky outcrops in primary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º24’42’’N 105º38’06’E at elev. of about 700 m a.s.l. Erect palm-like tree 0.4-0.8 m tall. In observation of year 2001 & 2002 locally very common. Presently almost extinct due to overcollection for sale and cultivation as an ornamental plant.

30 October 2023, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, Nguyen Van Ly, T. Maisak, AL2399.