Herbarium LE - Specimens

Orchidaceae

Cypripedium lentiginosum P.J.Cribb & S.C.Chen

Northwest Vietnam, Ha Giang Province, 23 June 2023, Hoang Duc Minh, Nguyen Sinh Khang, AL2218.

Orchidaceae

Cypripedium lentiginosum P.J.Cribb & S.C.Chen

Northwest Vietnam, Ha Giang Province, 23 June 2023, Hoang Duc Minh, Nguyen Sinh Khang, AL2218.

Meliaceae

Chukrasia tabularis Wight & Arn.

Bac Kan Province, Ba Be District, Ba Be National Park. Dry evergreen broad-leaved secondary forest on steep rocky slope composed by white crystalline marble-like highly eroded limestone along lake shore on point 22º25’46’’N 105º36’58’E at elev. of about 200 m a.s.l. Tree to 25 m tall. Flowers yellowish green, very fragrant.

28 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2956.

Zingiberaceae

Amomum sp.?

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Secondary evergreen broad-leaved forest on steep rocky slope composed by white crystalline marble-like highly eroded limestone around point 22º27’36’’N 105º39’18’E at elevation of about 250 m a.s.l. Terrestrial herb to 1.5 m tall in shady slope. Flowers white, lip with yellow spot. Locally common.

26 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2916.

Convallariaceae

Aspidistra opaca Tillich

Khanh Hoa Province, Khanh Vĩnh District, eastern slopes of Chu Mu Mountains at elevation 350­450 m a.s.l. Dry secondary evergreen broad-leaved forest on very steep granite and shale slopes. Terrestrial herb with thick hypogeous rhizome on sandy slope along small rocky stream. Old flowers dark violet to almost black, stigma yellowish, fruits dirty green, spiny in apical half. Locally common. 26 February 2023, L. Averyanov, Nguyen Van Canh, AL 2159. Living plants and herbarium.

Asclepiadaceae

Hoya graveolens Kerr

Dak Lak Province, Ea’Hleo District, evergreen broad-leaved forest at elevation of about 300 m a.s.l., lithophytic vine, obtained from local collectors in 2014.

10 May 2024, Nguyen Van Canh garden, AL3139.

Solanaceae

Solanum sp.

Dak Lak Province, Lak District, around Lieng Bok Sak Waterfall at elevation about 400 m a.s.l. Dry evergreen broad-leaved forest along rocky stream on granite. Erect terrestrial herb to 1.5 m tall in open place along road. Flowers white, not ripe fruits pale green. Common.

11 May 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3145.

Araceae

Typhonium sp.?

Plant cultivated in Ly Ngoc Quan private garden, Lam Dong Province, Lam Ha District, Phi To Village as a weed. Photo and herbarium prepared in 11 May 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3153.

Gesneriaceae

Petrocodon sp.

Plant obtained from local collector in the north of VN. Cultivated in Loc Bao Sang private garden, Lam Dong Province, Di Linh District. Flowers dark blue.

Photo prepared in 12 May 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3159.

Gesneriaceae

Primulina sp.

Plant obtained from local collector in the north of VN. Cultivated in Loc Bao Sang private garden, Lam Dong Province, Di Linh District. Flowers large, white, outside with light purple markings.

Photo prepared in 12 May 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3160.

Myrsinaceae

Ardisia sp. nov.?

Plant obtained from local collector in Lam Dong Province, Di Linh District area. Cultivated in Loc Bao Sang private garden, Lam Dong Province, Di Linh District.

Herbarium and photo prepared in 12 May 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3165.

Convallariaceae

Aspidistra sp.

Dak Lak Province, Lak District, Krong No Municipality, around point 12.1408 N 108.1325 E. Dry evergreen broad-leaved secondary forest on hill slopes composed with granite at elevation 800–900 m a.s.l. Terrestrial herb on shady slope along rocky stream. Rare.

24 November 2022, L. Averyanov, Nguyen Van Canh, T. Maisak, Truong Ba Vuong, et al., AL 1786.

Gesneriaceae

Primulina sp.

Yen Bai Province, Luc Yen District, To Mau Municipality, Ngoi Tham Village, remnants of primary evergreen broad-leaved forest on limestone mountain at elevation 200–400 m a.s.l.03 March 2023, Nguyen Sinh Khang, AL 2205. Living plant.

Clusiaceae

Garcinia tinctoria W.Wight

Bac Kan Province, Ba Be District, Ba Be National Park. Dry evergreen broad-leaved secondary forest on steep rocky slope composed by white crystalline marble-like highly eroded limestone along lake shore on point 22º25’46’’N 105º36’58’E at elev. of about 200 m a.s.l. Tree to 20 m tall. Fruits light green. Occasional.

28 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2957.

Arecaceae

Arenga caudata (Lour.) H.E.Moore

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Dry secondary evergreen broad-leaved forest on steep rocky slope composed by white crystalline marble-like highly eroded limestone around point 22º27’42’’N 105º34’41’E at elevation of 700–800 m a.s.l. Miniature palm to 1.5 m tall in rocky open place between remnant hills. Occasional.

27 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2953.

Polypodiaceae

Tectaria sp.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Dry secondary evergreen broad-leaved forest on steep rocky slope composed by white crystalline marble-like highly eroded limestone around point 22º27’42’’N 105º34’41’E at elevation of 700–800 m a.s.l.Terrestrial fern in shady depression between limestone hills. Common.

27 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2951.

Ebenaceae

Diospyros sp.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Dry secondary evergreen broad-leaved forest on steep rocky slope composed by white crystalline marble-like highly eroded limestone around point 22º27’42’’N 105º34’41’E at elevation of 700–800 m a.s.l. Tree to 15 m tall in shady depression between limestone hills. Fruits and calyx green. Not common.

27 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2950.

Icacinaceae

Iodes vitiginea Hance

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Dry secondary evergreen broad-leaved forest on steep rocky slope composed by white crystalline marble-like highly eroded limestone around point 22º27’42’’N 105º34’41’E at elevation of 700–800 m a.s.l. Semi-woody vine 6-7 m long in open secondary forest along fiend border. Flowers yellowish. Common.

27 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2948.

Polypodiaceae (Apleniaceae)

Asplenium nidus L.,

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Dry secondary evergreen broad-leaved forest on steep rocky slope composed by white crystalline marble-like highly eroded limestone around point 22º27’42’’N 105º34’41’E at elevation of 700–800 m a.s.l. Lithophytic rosulate fern along rocky ridge edge. Not rare.

27 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2945.

Convallariaceae

Aspidistra babensis K.S.Nguyen, Aver. & Tillich

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Dry secondary evergreen broad-leaved forest on steep rocky slope composed by white crystalline marble-like highly eroded limestone around point 22º27’42’’N 105º34’41’E at elevation of 700–800 m a.s.l. Terrestrial herb with underground rhizome, leaves very rigid, almost black. Locally abundant.

27 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2944.

Cycadaceae

Cycas brachycantha K.D.Hill, T.H.Nguyen & L.K.Phan

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Dry secondary evergreen broad-leaved forest on steep rocky slope composed by white crystalline marble-like highly eroded limestone around point 22º27’42’’N 105º34’41’E at elevation of 700–800 m a.s.l. Lithophytic palm-like plant with erect stem to 0.5 m tall and leaves to 2.5 m long along rocky ridge edge. Locally very common. Male specimen.

27 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2942.

Rubiaceae

Psychotria sp.?

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Dry secondary evergreen broad-leaved forest on steep rocky slope composed by white crystalline marble-like highly eroded limestone around point 22º27’42’’N 105º34’41’E at elevation of 700–800 m a.s.l. Undershrub 1 m tall on steep shady rocky slope. Flowers white. Not common.

27 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2936.

Lauraceae

Neocinnamomum lecomtei H. Liu

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Dry secondary evergreen broad-leaved forest on steep rocky slope composed by white crystalline marble-like highly eroded limestone around point 22º27’42’’N 105º34’41’E at elevation of 700–800 m a.s.l. Treelet 1.5 2 m tall on steep rocky slope. Fruits bright red. Rare.

27 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2935.

Opiliaceae

Melientha suavis Pierre

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Dry secondary evergreen broad-leaved forest on steep rocky slope composed by white crystalline marble-like highly eroded limestone around point 22º27’42’’N 105º34’41’E at elevation of 700–800 m a.s.l. Treelet to 10 m in rather open place. Not rare.

27 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2934.

Vitaceae

Tetrastigma leucostaphylum (Dennst.) Alston?

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Dry secondary evergreen broad-leaved forest on steep rocky slope composed by white crystalline marble-like highly eroded limestone around point 22º27’42’’N 105º34’41’E at elevation of 700–800 m a.s.l. Herbaceous vine to 4 m long in rather open place. Fruits bright orange red. Not rare.

27 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2933.

Pandaceae

Microdesmis caseariifolia Planch. ex Hook.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Secondary evergreen broad-leaved forest on steep rocky slope composed by white crystalline marble-like highly eroded limestone around point 22º27’36’’N 105º39’18’E at elevation of about 250 m a.s.l. Shrub or small treelet in shady place. Flower buds yellow. Occasional. common.

26 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2927.

Annonaceae

Rauwenhoffia sp.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Secondary evergreen broad-leaved forest on steep rocky slope composed by white crystalline marble-like highly eroded limestone around point 22º27’36’’N 105º39’18’E at elevation of about 250 m a.s.l. Shrub or small woody vine. Not common.

26 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2926.

Solanaceae

Solanum sp.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Secondary evergreen broad-leaved forest on steep rocky slope composed by white crystalline marble-like highly eroded limestone around point 22º27’36’’N 105º39’18’E at elevation of about 250 m a.s.l. Treelet 4-5 m tall in open place, flowers white with violet tint, ripe fruits dull orange. Occasional.

26 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2925.

Rubiaceae

Ixora sp.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Secondary evergreen broad-leaved forest on steep rocky slope composed by white crystalline marble-like highly eroded limestone around point 22º27’36’’N 105º39’18’E at elevation of about 250 m a.s.l. Shrub 1 n tall in in open place, flowers white. Rare.

26 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2923.

Alangiaceae

Alangium sp.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Secondary evergreen broad-leaved forest on steep rocky slope composed by white crystalline marble-like highly eroded limestone around point 22º27’36’’N 105º39’18’E at elevation of about 250 m a.s.l. Tree 5-8 m tall in open place, flowers white. Occasional.

26 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2922.

Solanaceae

Solanum sp.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Secondary evergreen broad-leaved forest on steep rocky slope composed by white crystalline marble-like highly eroded limestone around point 22º27’36’’N 105º39’18’E at elevation of about 250 m a.s.l. Erect herb to 1 m tall in open place along road. Flowers white with violet tint. Occasional.

26 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2921.

Polypodiaceae (Pteridaceae)

Pteris cretica L.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Secondary evergreen broad-leaved forest on steep rocky slope composed by white crystalline marble-like highly eroded limestone around point 22º27’36’’N 105º39’18’E at elevation of about 250 m a.s.l. Terrestrial fern to 0.6 m tall on very steep rocky slope. Common.

26 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2920.

Euphorbiaceae

Antidesma sp.?

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Secondary evergreen broad-leaved forest on steep rocky slope composed by white crystalline marble-like highly eroded limestone around point 22º27’36’’N 105º39’18’E at elevation of about 250 m a.s.l. Tree 6-8 m tall in open place. Not rare.

26 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2919.

Arecaceae

Arenga caudata (Lour.) H.E.Moore

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Secondary evergreen broad-leaved forest on steep rocky slope composed by white crystalline marble-like highly eroded limestone around point 22º27’36’’N 105º39’18’E at elevation of about 250 m a.s.l. Palm to 1.5 tall on shady slope. Not common.

26 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2917.

Rutaceae

Evodia sp.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Secondary evergreen broad-leaved forest on steep rocky slope composed by white crystalline marble-like highly eroded limestone around point 22º27’36’’N 105º39’18’E at elevation of about 250 m a.s.l. Tree 10-15 m tall, to 0.3 m DBH on open place. Ripe fruits orange red. Common.

26 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2915.

Rutaceae

Evodia sp.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Secondary evergreen broad-leaved forest on steep rocky slope composed by white crystalline marble-like highly eroded limestone around point 22º27’36’’N 105º39’18’E at elevation of about 250 m a.s.l. Tree 10-15 m tall, to 0.3 m DBH on open place. Flowers white. Common.

26 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2914.

Fabaceae (Caesalpiniaceae)

Cassia sp.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Secondary evergreen broad-leaved forest on steep rocky slope composed by white crystalline marble-like highly eroded limestone around point 22º27’36’’N 105º39’18’E at elevation of about 250 m a.s.l. Tree to 15 m tall and 0.4 m DBH on steep slope along road. Common.

26 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2913.

Euphorbiaceae

Mallotus sp.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Old secondary evergreen broad-leaved forest and scrub on very steep rocky slope composed by white crystalline marble-like highly eroded limestone above tall cliff around point 22º27’36’’N 105º39’18’E at elevation of about 300 m a.s.l. Tree to 10 m tall on rather open mossy rocky steep slope. Common.

25 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2910.

Gesneriaceae

Paraboea sp.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Old secondary evergreen broad-leaved forest and scrub on very steep rocky slope composed by white crystalline marble-like highly eroded limestone above tall cliff around point 22º27’36’’N 105º39’18’E at elevation of about 300 m a.s.l. Terrestrial and lithophytic herb to 0.5 m tall on rather open mossy rocky steep slope. Locally common.

25 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2907.

Urticaceae

Procris rhizantha Gagnep.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Old secondary evergreen broad-leaved forest and scrub on very steep rocky slope composed by white crystalline marble-like highly eroded limestone above tall cliff around point 22º27’36’’N 105º39’18’E at elevation of about 300 m a.s.l. Erect herb to 0.5 m tall on rocky, mossy, steep rather open slope. Not rare.

25 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2904.

Euphporbiaceae

Strophioblachia fimbricalyx Boerl.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Old secondary evergreen broad-leaved forest and scrub on very steep rocky slope composed by white crystalline marble-like highly eroded limestone above tall cliff around point 22º27’36’’N 105º39’18’E at elevation of about 300 m a.s.l. Treelet 4–5 m tall on rather open place in valley between limestone hills. Fruits and calyx green. Not rare.

25 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2903.

Menispermaceae

Stephania sp.?

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Old secondary evergreen broad-leaved forest and scrub on very steep rocky slope composed by white crystalline marble-like highly eroded limestone above tall cliff around point 22º27’36’’N 105º39’18’E at elevation of about 300 m a.s.l. Herbaceous vine to 5 m long with globular tuber 20-25 cm in diam., among scrub on rocky, mossy, steep rather open slope. Fruits light green. Not rare.

25 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2902.

Asclepiadaceae

Dischidia acutifolia Maingay ex Hook.f.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Old secondary evergreen broad-leaved forest and scrub on very steep rocky slope composed by white crystalline marble-like highly eroded limestone above tall cliff around point 22º27’36’’N 105º39’18’E at elevation of about 300 m a.s.l. Small herbaceous vine to 2 m long among scrub on mossy steep, rather open slope. Flowers pale yellowish. Common.

25 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2901.

Polypodiaceae

Pyrrosia adnascens (Sw.) Ching

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Old secondary evergreen broad-leaved forest and scrub on very steep rocky slope composed by white crystalline marble-like highly eroded limestone above tall cliff around point 22º27’36’’N 105º39’18’E at elevation of about 300 m a.s.l. Creeping epiphytic and lithophytic fern on mossy steep, rather open slope. Abundant.

25 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2900.

Asclepiadaceae

Hoya diversifolia Blume

Dak Lak Province, Chu Yang Sing Mountain, Chu Yang Sinh National Park, evergreen broad-leaved forest at elevation of 400-500 m a.s.l., lithophytic vine, obtained from local collectors in 2014.

10 May 2024, Nguyen Van Canh garden, AL3138.

Asclepiadaceae

Hoya thuhae Aver., Vuong & V.C.Nguyen

HOLOTYPE

Dak Lak Province, Yok Don National Park, Dipterocarp forest at elevation of about 200 m a.s.l., living plant obtained from local collectors in 2018 by Nguyen Van Canh. Herbarium specimen and photos were made from plants cultivated in Buon Ma Thuot on 10 May 2024, Nguyen Van Canh, L. Averyanov, T. Maisak, AL3137.

Zingiberaceae

Alpinia vietnamica

Da Nang, Son Tra, evergreen broad-leaved forest at elevation of about 400 m a.s.l., collected in 2021.

10 May 2024, Nguyen Van Canh garden, AL3133.

Lowiaceae

Orchidantha sp.

Dong Nai Province, Nam Cat Thien National Park, evergreen broad-leaved forest at elevation of about 200 m a.s.l., collected in 2019.

10 May 2024, Nguyen Van Canh garden, AL3132.

Zingiberaceae

Curcuma sp.

Dak Lak Province, Dipterocarp forest at elevation of about 200 m a.s.l., obtained from local collectors, collected in 2019.

10 May 2024, Nguyen Van Canh, AL3130.

Asclepiadaceae

Hoya burmanica Rolfe

Son La Province, obtained from local collectors,

10 May 2024, Nguyen Van Canh, AL3126.