Herbarium LE - Specimens

Selaginellaceae

Selaginella sp.

Binh Thuan Province, Tanh Linh District, Nui Ong Nature Reserve. Open grassy Dipterocarp woodland on thin degraded lateritic soil at elevation about 200 m a.s.l. Erect terrestrial herb to 20 cm tall in shady place. Leaves light green. Very common.

31 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3482.

Selaginellaceae

Selaginella sp.

Binh Thuan Province, Tanh Linh District, Nui Ong Nature Reserve. Open grassy Dipterocarp woodland on thin degraded lateritic soil at elevation about 200 m a.s.l. Erect terrestrial herb to 20 cm tall in shady place. Leaves light green. Very common.

31 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3482.

Burmanniaceae

Burmannia sp.

Binh Thuan Province, Tanh Linh District, Nui Ong Nature Reserve. Open grassy Dipterocarp woodland on thin degraded lateritic soil at elevation about 200 m a.s.l. Erect terrestrial achlorophyllous herb on open moist soil along trail. Flowers white. Locally common.

31 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3479.

Selaginellaceae

Selaginella sp.

Binh Thuan Province, Tanh Linh District, Nui Ong Nature Reserve. Open grassy Dipterocarp woodland on thin degraded lateritic soil at elevation about 200 m a.s.l. Erect terrestrial herb to 25 cm tall on moist soil along trail. Locally very common.

31 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3481.

Selaginellaceae

Selaginella sp.

Binh Thuan Province, Tanh Linh District, Nui Ong Nature Reserve. Open grassy Dipterocarp woodland on thin degraded lateritic soil at elevation about 200 m a.s.l. Prostrate terrestrial herb on open moist soil along trail. Locally common.

31 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3480.

Melastomataceae

Memecylon sp.

Lam Dong Province, Bao Lam District. Secondary grassy coniferous woodlands and open scrub on thin degraded volcanic lateritic rocky soil at elevation about 900 m a.s.l. Shrub or treelet to 3 m tall in open shrubs. Not common.

30 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3477.

Hamamelidaceae

Quangnamia syncarpa K.S.Nguyen, Aver., C.W.Lin, G.Q.Liu & H.H.Truong

Hamamelidaceae

Quangnamia syncarpa K.S.Nguyen, Aver., C.W.Lin, G.Q.Liu & H.H.Truong⁣

Quang Nam Province, Dai Loc District, Dai Hong Commune, Ngoc Thach Village, Huu Nien Mountain, evergreen broad-leaved forest on sandstone, around point 15.823°N, 107.966°E, at elevation of 400–600 m a. s. l., tree 7–15 m tall, growing on rocky places, not common. 13 VI 2024. Truong Hoang Hap, HH-QN 15”

ISOTYPE

Vietnam, Quang Nam Province: Dai Loc District, Dai Hong Commune, Ngoc Thach Village, Huu Nien Mountain, evergreen broad-leaved forest on sandstone, around point 15.823 °N, 107.966 °E, at elevation of 400–600 m a. s. l., tree 10–25 m tall, 10–20(25) cm in DBH, usually growing on rocky places nearby stream, not common. 9 VII 2024. Truong Hoang Hap, HH-QN 30

Plants of Dong Chau - Khe Nuoc Trong Nature Reserve (DCKNTNR)

Rubiaceae

Lasianthus sp.

Quang Binh Province, Le Thuy District, Kim Thuy Municipality. Secondary evergreen broad-leaved forest with bamboo on very steep river slopes composed by shaly limestone at elevation about 300-400 m a.s.l. around point 17º4’29’’N 106º43’5’’E. 22 November 2024, Nguyen Huu Cuong, Phan Thanh Quyet, Duong Van Nam, L. Averyanov, T. Maisak, DCKNTNR110.

Plants of Dong Chau - Khe Nuoc Trong Nature Reserve (DCKNTNR)

Verbenaceae

Lantana camara L.

Quang Binh Province, Le Thuy District, Kim Thuy Municipality. Secondary evergreen broad-leaved forest with bamboo on very steep river slopes composed by shaly limestone at elevation about 160 m a.s.l. around point 17º4’29’’N 106º43’5’’E. Climbing shrub to 1.5 m tall on open road cut among secondary shrubs. Flowers orange. Very common aggressive weed.

24 November 2024, Nguyen Huu Cuong, Phan Thanh Quyet, Duong Van Nam, L. Averyanov, T. Maisak, DCKNTNR149 (AL3719).

Gesneriaceae

Chirita sp.?

Plant obtained from local dealers from Ninh Binh Province, Cuc Phuong National Park and cultivated by Loc Bao Sang in Lam Dong, Di Linh Town. Photo and herbarium prepared in 28 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3430.

FLORA OF VIETNAM

Convallariaceae

Aspidistra marasmioides Tillich

(A. erecta Yan Liu & C.-I Peng)

Ha Giang prov., Quan Ba distr., Bat Dai Son Nature Reserve, Can Ty commune, Lung Vay village, around point 23º06ʹ02.6ʺN 105º03ʹ04.5ʺE, 1200-1305 m a.s.l., karstic, highly eroded mountains composed of solid marble-like limestone, primary evergreen broad-leaved humid forest on steep rocky slopes near mountain tops. Erect terrestrial herb with semiwoody stem to 0.40 cm tall among shady rocks. Leaves uniform green. Fruits brown, globular prickly-papillose, 5-7 mm in diameter. Not common. 15 April 2018, L. Averyanov, Nguyen Sinh Khang, Chuong Quang Ngan, T. Maisak, VR 201 (HN, LE01054358).

Gesneriaceae

Chirita sp.?

Plant obtained from local dealers from Lai Chau Province, Bat Xay District and cultivated by Loc Bao Sang in Lam Dong, Di Linh Town. Photo and herbarium prepared in 28 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3423.

Plants of Dong Chau - Khe Nuoc Trong Nature Reserve (DCKNTNR)

Polypodiaceae s.l.

Leptochilus chilangensis (V.N.Tu) Liang Zhang & Li Bing Zhang?

Quang Binh Province, Le Thuy District, Kim Thuy Municipality. Secondary evergreen broad-leaved forest with bamboo on very steep river slopes composed by shaly limestone at elevation about 160 m a.s.l. around point 17º4’29’’N 106º43’5’’E. Creeping lithophytic fern on shady wet stream rocks. Locally common.

24 November 2024, Nguyen Huu Cuong, Phan Thanh Quyet, Duong Van Nam, L. Averyanov, T. Maisak, DCKNTNR155 (AL3728).

Plants of Dong Chau - Khe Nuoc Trong Nature Reserve (DCKNTNR)

Casuarinacea

Casuarina equisetifolia L.

Cultivated tree about 18 m tall in Hanoi, on the territory of National Forestry University. Many clustered fascicles in the tree canopy. Observed in alone tree only.

27 November 2024, Nguyen Huu Cuong, L. Averyanov, T. Maisak, AL3735.

Asclepiadaceae

Dischidia sp.

Dak Lak Province, Chu Mu Mountain, evergreen broad-leaved submontane forest at elevation about 1000 m a.s.l. Epiphytic vine to 2 m long, flowers white. Ly Ngoc Quan sine n. Herbarium specimen collected from cultivated plant. 26 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3392.

Balsaminaceae

Impatiens coerulea sp. nov.?

Dak Lak Province, Lac District, southern part of Nam Kar Nature Reserve around point 12º16’10’’N 108º05’24’’E at elevations of about 600 m a.s.l. Open steep rocky slope in secondary evergreen broad-leaved forest with bamboo along road. Lithophytic herb with fleshy stem to 0.8 m tall among large rocks. Flowers bluish with yellow. Locally very common.

26 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3388.

Balsaminaceae

Impatiens coerulea sp. nov.?

Dak Lak Province, Lac District, southern part of Nam Kar Nature Reserve around point 12º16’10’’N 108º05’24’’E at elevations of about 600 m a.s.l. Open steep rocky slope in secondary evergreen broad-leaved forest with bamboo along road. Lithophytic herb with fleshy stem to 0.8 m tall among large rocks. Flowers bluish with yellow. Locally very common.

26 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3388.

Malvaceae

Hibiscus sp.

Dak Lak Province, Lac District, southern part of Nam Kar Nature Reserve around point 12º16’10’’N 108º05’24’’E at elevations of about 600 m a.s.l. Secondary open shrubs along road. Shrub or small tree to 4 m tall in open place, Flowers yellow, later to orange red. Occasional.

26 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3387.

Malvaceae

Hibiscus sp.

Dak Lak Province, Lac District, southern part of Nam Kar Nature Reserve around point 12º16’10’’N 108º05’24’’E at elevations of about 600 m a.s.l. Secondary open shrubs along road. Shrub or small tree to 4 m tall in open place, Flowers yellow, later to orange red. Occasional.

26 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3387.

Solanaceae

Solanum sp.

Dak Lak Province, Lac District, southern part of Nam Kar Nature Reserve around point 12º16’10’’N 108º05’24’’E at elevations of about 600 m a.s.l. Secondary open shrubs along road. Shrub to 2 m tall in open place, Flowers white. Common.

26 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3386.

Solanaceae

Solanum sp.

Dak Lak Province, Lac District, southern part of Nam Kar Nature Reserve around point 12º16’10’’N 108º05’24’’E at elevations of about 600 m a.s.l. Secondary open shrubs along road. Shrub to 2 m tall in open place, Flowers white. Common.

26 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3386.

Convallariaceae

Aspidistra sp.

Dak Lak Province, Lac District, southern part of Nam Kar Nature Reserve around point 12º16’10’’N 108º05’24’’E at elevations of about 600 m a.s.l. Steep slope in secondary evergreen broad-leaved forest with bamboo along road near small stream. Terrestrial herb on shady slope with thick stout rhizome to 30 cm long. Not common.

26 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3385.

Gesneriaceae

Billilovia sp.

Dak Lak Province, Lac District, southern part of Nam Kar Nature Reserve around point 12º16’10’’N 108º05’24’’E at elevations of about 600 m a.s.l. Steep slope in secondary evergreen broad-leaved forest with bamboo along road near small stream. Terrestrial herb on shady slope. Locally common.

26 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3384.

Begoniaceae

Begonia sp.

Dak Lak Province, Lac District, southern part of Nam Kar Nature Reserve around point 12º16’10’’N 108º05’24’’E at elevations of about 600 m a.s.l. Steep slope in secondary evergreen broad-leaved forest with bamboo along road near small stream. Terrestrial herb on shady slope, leaves to 40 cm long and wide. Locally common.

26 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3383.

Begoniaceae

Begonia sp.

Dak Lak Province, Lac District, southern part of Nam Kar Nature Reserve around point 12º16’10’’N 108º05’24’’E at elevations of about 600 m a.s.l. Steep slope in secondary evergreen broad-leaved forest with bamboo along road near small stream. Terrestrial herb on shady slope, leaves to 40 cm long and wide. Locally common.

26 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3383.

Arecaceae

Arenga caudata (Lour.) H.E.Moore

Phu Yen Province, Dong Ha District, Deo Ca Pass. Dense sclerophyllous short wind formed forest and scrub on steep rocky sea-faced slopes among big granite boulders at elevation 100-150 m a.s.l. Terrestrial palm to 1.5 tall on shady rocky steep slope. Not rare.

04 November 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3550.

Rubiaceae

Ophiorrhiza sp.

Binh Dinh Province, Van Canh District, Canh Lien Municipality. Fractionally logged dense dry primary evergreen broad-leaved forest on steep mountain slopes composed by granite with quartzite at elevation 600-700 m a.s.l. Prostrate rosulate terrestrial herb on shady rocky stream slope. Locally common.

05 November 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3583.

Plants of Dong Chau - Khe Nuoc Trong Nature Reserve (DCKNTNR)

Urticaceae

Debregeasia squamata Hook.

Quang Binh Province, Le Thuy District, Kim Thuy Municipality. Secondary and primary fractionally logged evergreen broad-leaved forest on very steep hill slopes composed by gray and yellow solid marble-like limestone at elevation about 200–250 m a.s.l. around point 16º59.5 N 106º38.2 E. Shrub to 2 m tall in open place along trail. Common.

09 November 2024, Nguyen Huu Cuong, Nguyen Van Ly, Phan Thanh Quyet, Duong Van Nam, Le Thi Phuong Thao, L. Averyanov, T. Maisak, AL3615.

Plants of Dong Chau - Khe Nuoc Trong Nature Reserve (DCKNTNR)

Convallariaceae

Aspidistra phongdiensis D.Dien, T.A.Le & Vislobokov

Quang Binh Province, Le Thuy District, Kim Thuy Municipality. Secondary and primary fractionally logged evergreen broad-leaved forest on very steep hill slopes composed by gray and yellow solid marble-like limestone at elevation about 200–250 m a.s.l. around point 16º59.5 N 106º38.2 E. Terrestrial creeping herb with thick plagiotropic or ascending rhizome with

many short stilt roots on steep shady stream slope. Leaves with long petiole and elliptic blade. Peduncle erect, bent down at apex. Flowers nutant, obovoid, container shaped, with 6 short, triangular, recurved lobes and small opening, dull pale greenish, yellowish or pinkish outside, pink purple inside, stamens yellow. Locally common.

09 November 2024, Nguyen Huu Cuong, Nguyen Van Ly, Phan Thanh Quyet, Duong Van Nam, Le Thi Phuong Thao, L. Averyanov, T. Maisak, AL3632.

Plants of Dong Chau - Khe Nuoc Trong Nature Reserve (DCKNTNR)

Convallariaceae

Aspidistra phongdiensis D.Dien, T.A.Le & Vislobokov

Quang Binh Province, Le Thuy District, Kim Thuy Municipality. Secondary and primary fractionally logged evergreen broad-leaved forest on very steep hill slopes composed by gray and yellow solid marble-like limestone at elevation about 200–250 m a.s.l. around point 16º59.5 N 106º38.2 E. Terrestrial creeping herb with thick plagiotropic or ascending rhizome with

many short stilt roots on steep shady stream slope. Leaves with long petiole and elliptic blade. Peduncle erect, bent down at apex. Flowers nutant, obovoid, container shaped, with 6 short, triangular, recurved lobes and small opening, dull pale greenish, yellowish or pinkish outside, pink purple inside, stamens yellow. Locally common.

09 November 2024, Nguyen Huu Cuong, Nguyen Van Ly, Phan Thanh Quyet, Duong Van Nam, Le Thi Phuong Thao, L. Averyanov, T. Maisak, AL3632.

Plants of Dong Chau - Khe Nuoc Trong Nature Reserve (DCKNTNR)

Convallariaceae

Aspidistra phongdiensis D.Dien, T.A.Le & Vislobokov

Quang Binh Province, Le Thuy District, Kim Thuy Municipality. Secondary and primary fractionally logged evergreen broad-leaved forest on very steep hill slopes composed by gray and yellow solid marble-like limestone at elevation about 200–250 m a.s.l. around point 16º59.5 N 106º38.2 E. Terrestrial creeping herb with thick plagiotropic or ascending rhizome with

many short stilt roots on steep shady stream slope. Leaves with long petiole and elliptic blade. Peduncle erect, bent down at apex. Flowers nutant, obovoid, container shaped, with 6 short, triangular, recurved lobes and small opening, dull pale greenish, yellowish or pinkish outside, pink purple inside, stamens yellow. Locally common.

09 November 2024, Nguyen Huu Cuong, Nguyen Van Ly, Phan Thanh Quyet, Duong Van Nam, Le Thi Phuong Thao, L. Averyanov, T. Maisak, AL3632.

Plants of Dong Chau - Khe Nuoc Trong Nature Reserve (DCKNTNR)

Convallariaceae

Aspidistra phongdiensis D.Dien, T.A.Le & Vislobokov

Quang Binh Province, Le Thuy District, Kim Thuy Municipality. Secondary and primary fractionally logged evergreen broad-leaved forest on very steep hill slopes composed by gray and yellow solid marble-like limestone at elevation about 200–250 m a.s.l. around point 16º59.5 N 106º38.2 E. Terrestrial creeping herb with thick plagiotropic or ascending rhizome with many short stilt roots on steep shady stream slope. Leaves with long petiole and elliptic blade. Peduncle erect, bent down at apex. Flowers nutant, obovoid, container shaped, with 6 short, triangular, recurved lobes and small opening, dull pale greenish, yellowish or pinkish outside, pink purple inside, stamens yellow. Locally common.

09 November 2024, Nguyen Huu Cuong, Nguyen Van Ly, Phan Thanh Quyet, Duong Van Nam, Le Thi Phuong Thao, L. Averyanov, T. Maisak, AL3632.

Plants of Dong Chau - Khe Nuoc Trong Nature Reserve (DCKNTNR)

Convallariaceae

Aspidistra phongdiensis D.Dien, T.A.Le & Vislobokov

Quang Binh Province, Le Thuy District, Kim Thuy Municipality. Secondary and primary fractionally logged evergreen broad-leaved forest on very steep hill slopes composed by gray and yellow solid marble-like limestone at elevation about 200–250 m a.s.l. around point 16º59.5 N 106º38.2 E.

Terrestrial creeping herb with thick plagiotropic or ascending rhizome with many short stilt roots on steep shady stream slope. Leaves with long petiole and elliptic blade. Peduncle erect, bent down at apex. Flowers nutant, obovoid, container shaped, with 6 short, triangular, recurved lobes and small opening, dull pale greenish, yellowish or pinkish outside, pink purple inside, stamens yellow. Locally common.

09 November 2024, Nguyen Huu Cuong, Nguyen Van Ly, Phan Thanh Quyet, Duong Van Nam, Le Thi Phuong Thao, L. Averyanov, T. Maisak, AL3632.

Plants of Dong Chau - Khe Nuoc Trong Nature Reserve (DCKNTNR)

Polypodiaceae s.l.

Vittaria sp. (Haplopteris sp.)

Quang Binh Province, Le Thuy District, Kim Thuy Municipality. Secondary evergreen broad-leaved forest with bamboo on very steep river slopes composed by shaly limestone at elevation about 250 m a.s.l. around point 17º4’29’’N 106º43’5’’E.

Clustering epiphyte almost exclusively on Lewistona chinensis. Not common.

22 November 2024, Nguyen Huu Cuong, Phan Thanh Quyet, Duong Van Nam, L. Averyanov, T. Maisak, AL3694.

Plants of Dong Chau - Khe Nuoc Trong Nature Reserve (DCKNTNR)

Convallariaceae

Aspidistra letreae Aver., Tillich & Le Tuan Anh

Quang Binh Province, Le Thuy District, Kim Thuy Municipality. Secondary evergreen broad-leaved forest with bamboo on very steep river slopes composed by shaly limestone at elevation about 100 m a.s.l. around point 17º4’29’’N 106º43’5’’E.

Terrestrial herb with suberect stem with many stilt roots on shady rocky stream slope and on hill tops. Locally common.

22 November 2024, Nguyen Huu Cuong, Phan Thanh Quyet, Duong Van Nam, L. Averyanov, T. Maisak, AL3692.

Plants of Dong Chau - Khe Nuoc Trong Nature Reserve (DCKNTNR)

Convallariaceae

Aspidistra letreae Aver., Tillich & Le Tuan Anh

Quang Binh Province, Le Thuy District, Kim Thuy Municipality. Secondary evergreen broad-leaved forest with bamboo on very steep river slopes composed by shaly limestone at elevation about 100 m a.s.l. around point 17º4’29’’N 106º43’5’’E.

Terrestrial herb with suberect stem with many stilt roots on shady rocky stream slope and on hill tops. Locally common.

22 November 2024, Nguyen Huu Cuong, Phan Thanh Quyet, Duong Van Nam, L. Averyanov, T. Maisak, AL3692.

Plants of Dong Chau - Khe Nuoc Trong Nature Reserve (DCKNTNR)

Polypodiaceae s.l.

Pteris sp. s.l.

Quang Binh Province, Le Thuy District, Kim Thuy Municipality. Secondary evergreen broad-leaved forest with bamboo on very steep river slopes composed by shaly limestone at elevation about 100 m a.s.l. around point 17º4’29’’N 106º43’5’’E.

Terrestrial creeping fern with fronds to 1 m tall on shady rocky stream slope. Locally common.

22 November 2024, Nguyen Huu Cuong, Phan Thanh Quyet, Duong Van Nam, L. Averyanov, T. Maisak, AL3691.

Plants of Dong Chau - Khe Nuoc Trong Nature Reserve (DCKNTNR)

Polypodiaceae s.l.

Colysis sp.

Quang Binh Province, Le Thuy District, Kim Thuy Municipality. Secondary evergreen broad-leaved forest with bamboo on very steep river slopes composed by shaly limestone at elevation about 100 m a.s.l. around point 17º4’29’’N 106º43’5’’E.

Lithophytic creeping fern on shady wet stream rocks along water. Locally very common.

22 November 2024, Nguyen Huu Cuong, Phan Thanh Quyet, Duong Van Nam, L. Averyanov, T. Maisak, AL3690.

Plants of Dong Chau - Khe Nuoc Trong Nature Reserve (DCKNTNR)

Polypodiaceae s.l.

Pteris sp. s.l.

Quang Binh Province, Le Thuy District, Kim Thuy Municipality. Secondary evergreen broad-leaved forest with bamboo on very steep river slopes composed by shaly limestone at elevation about 100 m a.s.l. around point 17º4’29’’N 106º43’5’’E.

Terrestrial creeping fern with fronds to 1 m tall on shady rocky stream slope. Locally common.

22 November 2024, Nguyen Huu Cuong, Phan Thanh Quyet, Duong Van Nam, L. Averyanov, T. Maisak, AL3691.

Plants of Dong Chau - Khe Nuoc Trong Nature Reserve (DCKNTNR)

Convallariaceae

Aspidistra coccigera Aver. & Tillich ?

Quang Binh Province, Le Thuy District, Kim Thuy Municipality. Secondary evergreen broad-leaved forest with bamboo on very steep river slopes composed by shaly limestone at elevation about 100 m a.s.l. around point 17º4’29’’N 106º43’5’’E.

Terrestrial rhizomatous herb on shady rocky stream slope. Occasional.

22 November 2024, Nguyen Huu Cuong, Phan Thanh Quyet, Duong Van Nam, L. Averyanov, T. Maisak, AL3688.

Plants of Dong Chau - Khe Nuoc Trong Nature Reserve (DCKNTNR)

Lycopodiaceae

Lycopodium cernuum L.

Quang Binh Province, Le Thuy District, Kim Thuy Municipality. Secondary evergreen broad-leaved forest with bamboo on very steep river slopes composed by shaly limestone at elevation about 100 m a.s.l. around point 17º4’29’’N 106º43’5’’E.

Terrestrial herb to 0.6 m tall in open wet road cut. Very common.

22 November 2024, Nguyen Huu Cuong, Phan Thanh Quyet, Duong Van Nam, L. Averyanov, T. Maisak, AL3687.

Plants of Dong Chau - Khe Nuoc Trong Nature Reserve (DCKNTNR)

Polypodiaceae s.l.

Blechnum orientale L.

Quang Binh Province, Le Thuy District, Kim Thuy Municipality. Secondary evergreen broad-leaved forest with bamboo on very steep river slopes composed by shaly limestone at elevation about 100 m a.s.l. around point 17º4’29’’N 106º43’5’’E.

Terrestrial and lithophytic fern with fronds to 1.5 long on open rocky stream cliffs. Very common.

22 November 2024, Nguyen Huu Cuong, Phan Thanh Quyet, Duong Van Nam, L. Averyanov, T. Maisak, AL3686.

Plants of Dong Chau - Khe Nuoc Trong Nature Reserve (DCKNTNR)

Polypodiaceae s.l.

Colysis digitata Ching

Quang Binh Province, Le Thuy District, Kim Thuy Municipality. Secondary evergreen broad-leaved forest with bamboo on very steep river slopes composed by shaly limestone at elevation about 100 m a.s.l. around point 17º4’29’’N 106º43’5’’E.

Lithophytic creeping fern on wet shady stream rocks. Very common.

22 November 2024, Nguyen Huu Cuong, Phan Thanh Quyet, Duong Van Nam, L. Averyanov, T. Maisak, AL3685.

Plants of Dong Chau - Khe Nuoc Trong Nature Reserve (DCKNTNR)

Euphorbiaceae

Mallotus sp.

Quang Binh Province, Le Thuy District, Kim Thuy Municipality. Secondary evergreen broad-leaved forest with bamboo on very steep river slopes composed by shaly limestone at elevation about 100 m a.s.l. around point 17º4’29’’N 106º43’5’’E.

Tree to 15 m tall in open place of stream valley. Not rare.

22 November 2024, Nguyen Huu Cuong, Phan Thanh Quyet, Duong Van Nam, L. Averyanov, T. Maisak, AL3684.

Plants of Dong Chau - Khe Nuoc Trong Nature Reserve (DCKNTNR)

Myrsinaceae

Ardisia sp.

Quang Binh Province, Le Thuy District, Kim Thuy Municipality. Secondary evergreen broad-leaved forest with bamboo on very steep river slopes composed by shaly limestone at elevation about 100 m a.s.l. around point 17º4’29’’N 106º43’5’’E.

Small erect shrub to 1.5 m tall on open wet stream rocks. Fruits bright green. Locally very common.

22 November 2024, Nguyen Huu Cuong, Phan Thanh Quyet, Duong Van Nam, L. Averyanov, T. Maisak, AL3683.

Plants of Dong Chau - Khe Nuoc Trong Nature Reserve (DCKNTNR)

Euphorbiaceae

Phyllanthus sp.

Quang Binh Province, Le Thuy District, Kim Thuy Municipality. Secondary evergreen broad-leaved forest with bamboo on very steep river slopes composed by shaly limestone at elevation about 100 m a.s.l. around point 17º4’29’’N 106º43’5’’E.

Small erect undershrub to 1 m tall on open wet stream rocks. Flowers red-purple. Locally common.

22 November 2024, Nguyen Huu Cuong, Phan Thanh Quyet, Duong Van Nam, L. Averyanov, T. Maisak, AL3682.

Plants of Dong Chau - Khe Nuoc Trong Nature Reserve (DCKNTNR)

Polypodiaceae s.l.

Cibotium barometz (L.) J.Sm.

Quang Binh Province, Le Thuy District, Kim Thuy Municipality. Secondary evergreen broad-leaved forest with bamboo on very steep river slopes composed by shaly limestone at elevation about 100 m a.s.l. around point 17º4’29’’N 106º43’5’’E.

Woody fern with erect stem to 0.5 m tall and fronds to 2 m long on rocky stream slope. Locally common.

22 November 2024, Nguyen Huu Cuong, Phan Thanh Quyet, Duong Van Nam, L. Averyanov, T. Maisak, AL3680.

Plants of Dong Chau - Khe Nuoc Trong Nature Reserve (DCKNTNR)

Polypodiaceae s.l.

Cibotium barometz (L.) J.Sm.

Quang Binh Province, Le Thuy District, Kim Thuy Municipality. Secondary evergreen broad-leaved forest with bamboo on very steep river slopes composed by shaly limestone at elevation about 100 m a.s.l. around point 17º4’29’’N 106º43’5’’E.

Woody fern with erect stem to 0.5 m tall and fronds to 2 m long on rocky stream slope. Locally common.

22 November 2024, Nguyen Huu Cuong, Phan Thanh Quyet, Duong Van Nam, L. Averyanov, T. Maisak, AL3680.