Herbarium LE - Specimens

Melastomataceae

Sonerila sp.

Gia Lai Province, Mang Yang District, Ayune Commune, Kon Ka Kinh National Park. Secondary and primary dry evergreen broad-leaved forest on shale and lava rocks at elevation of 900-1000 m a.s.l. around point 14º13N 108º20E. Erect herb to 10 cm tall on shady steep rocky stream slope. Leaves evenly green, nerves pale pink. Not common.

4 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4733.

Balsaminaceae

Impatiens sp.

Khanh Hoa Province, Khanh Vinh District, Khanh Le Pass. Primary evergreen broad-leaved forest on very steep rocky mountain slopes on granite and gneiss at elevation of 1000-1200 m a.s.l., around point 12º13’N 108º44’E. Erect terrestrial and lithophytic herb to 30 cm tall on wet mossy rocks and cliffs of small waterfall. Flowers bright purple. Not common.

19 April 2026, L. Averyanov, Nguyen Van Canh, T. Maisak, АL4973.

Balsaminaceae

Impatiens sp.

Khanh Hoa Province, Khanh Vinh District, Khanh Le Pass. Primary evergreen broad-leaved forest on very steep rocky mountain slopes on granite and gneiss at elevation of 1000-1200 m a.s.l., around point 12º13’N 108º44’E. Erect terrestrial and lithophytic herb to 30 cm tall on wet mossy rocks and cliffs of small waterfall. Flowers bright purple. Not common.

19 April 2026, L. Averyanov, Nguyen Van Canh, T. Maisak, АL4973.

Araceae

Typhonium sp.

Dac Lak Province, Lak District, Lieng Buk Sak Waterfall, evergreen broad-leaved forest on very steep slopes along stream. Terrestrial and lithophytic herb among open rocks. Not common.

June 2025, Nguyen Van Canh, AL4562.

Araceae

Arisaema sp.

Plant obtained from local collectors in Son La Province, Thuan Chau District. Cultivated and collected in Nguyen Van Canh Garden, July 2025, Nguyen Van Canh, AL4563.

Araceae

Typhonium sp.

Dac Lak Province, Lak District, Lieng Buk Sak Waterfall, evergreen broad-leaved forest on very steep slopes along stream. Terrestrial and lithophytic herb among open rocks. Not common.

June 2025, Nguyen Van Canh, AL4562.

Araceae

Thyphonium sp.

Dac Lak Province, Yok Don National Park, terrestrial herb in open dipterocarp forest at elevation of about 200 m a.s.l.

August 2025, Nguyen Van Canh, AL4564.

Araceae

Thyphonium sp.

Dac Lak Province, Yok Don National Park, terrestrial herb in open dipterocarp forest at elevation of about 200 m a.s.l.

August 2025, Nguyen Van Canh, AL4564.

Convallariaceae

Peliosanthes sp.

Lam Dong Province, Di Linh District, Brah Yang Mountain, primary and old secondary evergreen broad-leaved and mixed forest (with Pinus khasya) on granite around point 11º27.669ʹ N 108º03.543ʹ E at elevation of 1000-1100 m a.s.l. Terrestrial rhizomatous herb on shady rocky slope. Fruits ovoid to ellipsoid, glossy blue. Locally common.

14 November 2023, L. Averyanov, Nguyen Van Canh, T. Maisak, AL2685 (photo and seeds only).

Convallariaceae

Peliosanthes sp.

Ninh Thuan Province, Ninh Hai District, Vinh Hy Townlet. Very dry evergreen primary and secondary short-tall wind-formed coastal forest on very steep sea faced sandstone rocky slopes at elevation of 200–250 m a.s.l. Terrestrial herb in shady rocky places. Flowers dark violet. Not common.

16 May 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3235.

Convallariaceae

Ophiopogon muongnhensis K.S.Nguyen, N.Tanaka & Aver.

Dien Bien Prov., Muong Nhe Distr., Muong Nhe Natural Reserve, Sin Thau Municipality, A Pa Chai Village around point about N22º22’51’’ E102º14’26”. Old humid secondary evergreen broad-leaved submontane forest of steep mountain slopes composed with sandstone at elevations 700–800 m a.s.l. along small stream. Terrestrial herb with stem to 0.8 m long on shady slope. Very common.

15 May 2022, L. Averyanov, Tran Huy Thai, Khang Sinh Nguyen, Cuong Huu Nguyen, T. Maisak, Bac Cam Khoa, VR1641a.

Flowered in culture on 12 Mat 2026, L. Averyanov, VR1641a, garden number GN 22174.

Convallariaceae

Ophiopogon muongnhensis K.S.Nguyen, N.Tanaka & Aver.

Dien Bien Prov., Muong Nhe Distr., Muong Nhe Natural Reserve, Sin Thau Municipality, A Pa Chai Village around point about N22º22’51’’ E102º14’26”. Old humid secondary evergreen broad-leaved submontane forest of steep mountain slopes composed with sandstone at elevations 700–800 m a.s.l. along small stream. Terrestrial herb with stem to 0.8 m long on shady slope. Very common.

15 May 2022, L. Averyanov, Tran Huy Thai, Khang Sinh Nguyen, Cuong Huu Nguyen, T. Maisak, Bac Cam Khoa, VR1641a.

Flowered in culture on 12 Mat 2026, L. Averyanov, VR1641a, garden number GN 22174.

Convallariaceae

Peliosanthes sp.

Gia Lai Province, G’Bang District, Kon Chu Rang Nature Reserve, around point 14º30.324´ N 108º29.762´ E. Primary evergreen broad-leaved forest with Dacrycarpus and Dacrydium at elevation about 1200 m a.s.l. Terrestrial clustering herb in shady place. Leaves grassy green. Not rare. 21 November 2023, L. Averyanov, Nguyen Van Canh, T. Maisak, AL2741B (living plant only).

Flowered in culture on 22 December 2025, L. Averyanov, AL2741B (garden number 2465).

Convallariaceae

Peliosanthes sp.

Gia Lai Province, G’Bang District, Kon Chu Rang Nature Reserve, around point 14º30.324´ N 108º29.762´ E. Primary evergreen broad-leaved forest with Dacrycarpus and Dacrydium at elevation about 1200 m a.s.l. Terrestrial clustering herb in shady place. Leaves grassy green. Not rare.

21 November 2023, L. Averyanov, Nguyen Van Canh, T. Maisak, AL2741A (living plant only).

Flowered in culture on 22 December 2025, L. Averyanov, AL2741A (garden number 2465).

Convallariaceae

Peliosanthes sp.

Lam Dong Province, Di Linh District, Brah Yang Mountain, primary and old secondary evergreen broad-leaved and mixed forest (with Pinus khasya) on granite around point 11º27.669ʹ N 108º03.543ʹ E at elevation of 1000-1100 m a.s.l. Terrestrial rhizomatous herb on shady rocky slope. Fruits ovoid to ellipsoid, glossy blue. Locally common.

14 November 2023, L. Averyanov, Nguyen Van Canh, T. Maisak, AL2685 (photo and seeds only).

Hypoxidaceae

Zeuxine nervosa (Lindl.) Trimen⁣

Hanoi Capital Region, Ba Vi National Park, old humid secondary and remnants of primary evergreen broad-leaved forest on steep mountain slopes composed by shale at elevation of about 1000 m a.s.l. Terrestrial rosulate herb on shady slope. Flowers yellow. Not rare.

10 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4263 (NHC066).

Orchidaceae

Bulbophyllum gibbosum

Lam Dong Province, Bao Lam District. Secondary grassy coniferous woodlands and open scrub on thin degraded volcanic lateritic rocky soil at elevation about 900 m a.s.l. Clustering epiphyte. Not rare.

30 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3489.

Fl. in culture 10 May 2026, L. Averyanov, AL3489.

FLORA OF VIETNAM

Orchidaceae

Phalaenopsis chibae T.Yukawa⁣

Lam Dong Province, Don Duong District, Thanh My Town area, secondary evergreen forest on granite along rocky stream at elevation about 1150 m a.s.l. Lithophyte. Not common.

28 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3418.

Orchidaceae

Cymbidium nanulum Y.S.Wu & S.C.Chen

Living plant obtained at 25 Nov. 2022, Nguyen Van Canh, L. Averyanov, T. Maisak, AL 1696. Plant originated from Dak Lak Province, Ea Hleo District, Yok Don NP, Dipterocarp forest at elevation 200–300 m a.s.l., along stream, locally common, 2020, Nguyen Van Canh.

(also common in Gia Lai and Dak Nong provinces).

Fl. in culture 12 May 2026, L. Averyanov, AL1696.

Orchidaceae

Tainia penangiana Hook.f.

Gia Lai Province, K’bang District, dry primary and old secondary evergreen broad-leaved forest on lava rocks along rocky river at elevation 600-650 m a.s.l. Canopy epiphyte in open places along stream, flowers light green, very small, about 0.5 mm across. Not common.

27 April 2025, L. Averyanov, N.V. Canh, T. Maisak, AL4006.

Orchidaceae

Acanthophippium sylhetense Lindl.

Dien Bien Prov., Muong Nhe Distr., Muong Nhe Natural Reserve, Leng Su Sin Municipality, A Suoi Voi Village around point about N22º20’16’’ E102º21’35”. Secondary evergreen broad-leaved lowland and submontane forest on mountain stream slopes composed with sandstone at elevations 500–600 m a.s.l. Terrestrial herb in shady place along stream valley. Flowers yellowish mottled with red. Not common.

18 May 2022, L. Averyanov, Tran Huy Thai, Khang Sinh Nguyen, T. Maisak, VR 1746.

Orchidaceae

Lam Dong Province, Bidoup Nui Ba National Park, Bidoup Mountains, primary evergreen broad-leaved forest at elev. 1700-1800 m a.s.l. Terrestrial herb in shady place. Locally very common. May 2019, Nguyen Van Canh, AL 1114.

Herbarium and photos prepared in 26 May 20121, L. Averyanov, T. Maisak, AL 1114.

Orchidaceae

Eria sp.

Thanh Hoa Province, Quan Hoa District, Nam Dong Community, Ban Lo Village, Ke Stream, the territory of Nam Dong National Reserve around point 22º25’46’’N 105º36’58’E. Primary and old secondary broad-leaved evergreen humid forest on very steep rocky slopes on solid, marble-like highly eroded limestone at elevation 900–1200 m a.s.l. on steep rocky slope. Clustering epiphyte and lithophyte on mossy trees on rocky mossy mountain top. Not rare.

04 May 2024, Nguyen Huu Cuong, Nguyen Van Ly, L. Averyanov, T. Maisak, AL3066.

Orchidaceae

Mycaranthes oblitterata Blume⁣.

Lam Dong Province, Bidoup Nui National Park, humid primary evergreen broad- leaved and secondary mixed ( with Pinus kesiya) forest at elevation of 1200- 2100 m.a.s.l.

14-26 january 2026, A. Petrov, ( living plant only)

Flowered in culture on 26 May 2026, L.Averyanov, AL 4524

Orchidaceae

Taeniophyllum sp.

Binh Dinh Province, Van Canh District. Secondary evergreen broad-leaved forest with bamboo on granite at elevation 250-350 m a.s.l., around point 13º38’N 108º57’E. Leafless epiphyte on small tree along rocky river in shady place. Rare.

12 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4892.

Flowered in culture on 26 May 2026, L. Averyanov, AL4892.

Orchidaceae

Erythrodes hirsuta (Griff.) Ormerod

Lam Dong Province, Dam Rong District, Phi Lieng Village, Phu Son Pass. Shady secondary forest with bamboo on very steep slope at elevation 1000-1400 m a.s.l. Terrestrial herb on shady steep road-cut. Rare.

18 April 2025, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3802a. Living plant only.

Flowered in culture on April 2026, L. Averyanov, AL4532a.

Orchidaceae

Panisea sp.

Lam Dong Province, Bidoup Nui Ba National Park, humid primary evergreen broad-leaved and secondary mixed (with Pinus kesiya) forest at elevation of 1200-2100 m a.s.l.

14-26 January 2026, A. Petrov, AL4530 (living plant only).

Flowered in culture on April 2026, L. Averyanov, AL4530.

Orchidaceae

Eria acervata Lindl.

Lam Dong Province, Bidoup Nui Ba National Park, humid primary evergreen broad-leaved and secondary mixed (with Pinus kesiya) forest at elevation of 1200-2100 m a.s.l.

14-26 January 2026, A. Petrov, AL4532 (living plant only).

Flowered in culture on April 2026, L. Averyanov, AL4532.

Orchidaceae

Eria sp.

Lam Dong Province, Bao Lam District. Secondary grassy coniferous woodlands and open scrub on thin degraded volcanic lateritic rocky soil at elevation about 900 m a.s.l. Clustering epiphyte. Not rare.

30 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3491a.

Fl. in culture April 2026, L. Averyanov, AL3491a.

Orchidaceae

Dendrobium faulhaberianum Schltr.

Kon Tum Province, Dak Gley District, W slopes of Ngoc Linh Mountain system. Remnants of primary and secondary evergreen broad-leaved forest on granite and shale at elevation of about 1100 m a.s.l. Epiphyte. Occasional.

02 May 2025, L. Averyanov, N.V. Canh, T. Maisak, AL4124.

Flowered in culture in April 2026, L. Averyanov, AL4124.

Orchidaceae

Corybas annamensis Aver.

Lam Dong Province, Bi Doup - Nui Ba National Park, evergreen forest at elevation of about 1600 m a.s.l.

July 2025, Nguyen Van Canh, AL4585.

Orchidaceae

Eria pannea

Lam Dong Province, Bidoup Nui Ba National Park, humid primary evergreen broad-leaved and secondary mixed (with Pinus kesiya) forest at elevation of 1200-2100 m a.s.l.

14-26 January 2026, A. Petrov, AL4520 (living plant only).

Flowered in culture on April 2026, L. Averyanov, AL4522.

Orchidaceae

Diploprora carinata Aver. et V.C.Nguyen, sp. nov.

Living plant obtained at 25 Nov. 2022, Nguyen Van Canh, L. Averyanov, T. Maisak, AL 1692.

Plant originated from Quang Nam Province, Nong Son District, evergreen broad-leaved forest at elevation 100–150 m a.s.l. 2022, Nguyen Van Canh.

Plant flowered in culture on 23 March 2026, L. Averyanov, AL1692.

Orchidaceae

Thrixspermum flammeum

Lam Dong Province, Bao Lam District. Secondary grassy coniferous woodlands and open scrub on thin degraded volcanic lateritic rocky soil at elevation about 900 m a.s.l. Epiphyte.

30 October 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3503.

Flowered in cultivation 12 May 2026, L. Averyanov, AL3503.

Orchidaceae

Octarrhena sp.

Lam Dong Province, Bidoup Nui Ba National Park, humid primary evergreen broad-leaved and secondary mixed (with Pinus kesiya) forest at elevation of 1200-2100 m a.s.l. Epiphyte on tall tree. Not common.

14-26 January 2026, A. Petrov, AL4539 (living plant only).

Flowered in culture on 6 May 2026, L. Averyanov, AL4539.

Orchidaceae

Trachoma rhopalorrhachis Garay?

Living plant obtained at 25 Nov. 2022, Nguyen Van Canh, L. Averyanov, T. Maisak, AL 1690.

Plant originated from Dak Lak Province, Lak District, Krong No Municipality, around point 12.2520 N 108.3449 E. Old closed evergreen broad-leaved secondary forest on hill slopes composed with granite at elevation about 650 m a.s.l. 2022, Nguyen Van Canh.

Flowered in 06 May 2025, L. Averyanov, 1690.

Orchidaceae

Bulbophyllum semiteretifolium Gagnep.

Lam Dong Province, Bidoup Nui Ba National Park, humid primary evergreen broad-leaved and secondary mixed (with Pinus kesiya) forest at elevation of 1200-2100 m a.s.l.

14-26 January 2026, A. Petrov, AL4520 (living plant only).

Flowered in culture on 11 March 2026, L. Averyanov, AL4520.

FAGACEAE

Castanopsis fissa (Champ. ex Benth.)

Rehder & E.H. Wilson

det. P. Manos, 2000

Vinh Phu

Tam Pao district, Tam Dao townlet.

Collections in degraded and highly

impacted vegetation (logged, trails,

over-use), along upper road to east of

town, approx. 100 m below ridge,

lateritic substrate.

21°27'07"N 105°38'31"E 991 m

Tree, occasional, to 6 m x 60 cm dbh;

bark fissured, leaves tan abaxially;

18 October 2000

D.K. Harder, P.K. Loc, D. Stone, P.

Manos, N.Q. Hieu & A.L. Thomas 6043

Supported by National Science Foundation

and National Geographic Society

MISSOURI BOTANICAL GARDEN HERBARIUM (MO)

Asteraceae

Aster trinervis Desf.⁣?

N. Vietnam, Hamrong.

19.01.1975, A. Takhtajan, 8450

Convallariaceae

Peliosanthes teta Andrews

Living plant obtained and herbarium prepared at 25 Nov. 2022, Nguyen Van Canh, L. Averyanov, T. Maisak, AL 1688. Plant originated from Dak Lak Province, Chu Yang Sinh Mountains, broad-leaved evergreen forest at elevations about 700 m a.s.l., 2018, Nguyen Van Can.

Flowered in culture in 16 February 2026, L. Averyanov , AL1688.

Ochidaceae

Oberonia falcata King & Pantl.

Lam Dong Province, Lac Duong District, Bidoup Nui Ba National Par, southern slopes of Bidoup Mountain about 5 km to the south from Bidoup peak around point 12º04.076’N 108º38.927E. Primary broad-leaved evergreen forest and Pinus woodlands on steep granite slopes at elevation 1900-2000 m a.s.l. Epiphyte on mossy trees. 08 November 2023, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, T. Maisak. AL2611.

Flowered in culture on 13 February 2026, L. Averyanov, AL2611.

Fam. Asteraceae

Aensleya sp.

N. Vietnam, prov. Cao Bang, distr. Bao Lac, municipality Dinh Phung, about 5 km to NW from Yen Lac village (N 22ø44' E 105ø50'), about 50

km to NWW from Cao Bang town. Old secondary closed broadleaved wet cloud forest on steep slopes and cliffs along top of Nam Linh

limestone ridge at 1500-1580 m alt. Epilite and terrestrial herb on steep slopes. Not common.

No CBL 469 19 Nov. 1998

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 6300-98)

Collectors: Averyanov L., P.K.Loc, N.X.Tam.

Fam. Asteraceae

Aensleya sp.

N. Vietnam, prov. Cao Bang, distr. Bao Lac, municipality Dinh Phung, about 5 km to NW from Yen Lac village (N 22ø44' E 105ø50'), about 50

km to NWW from Cao Bang town. Old secondary closed broadleaved wet cloud forest on steep slopes and cliffs along top of Nam Linh

limestone ridge at 1500-1580 m alt. Terrestrial and epilite herb. Leaves light green, violet veined. Rare.

No CBL 492 19 Nov. 1998

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 6300-98)

Collectors: Averyanov L., P.K.Loc, N.X.Tam.

Fam. Asteraceae

Aensleya sp.

N. Vietnam, prov. Cao Bang, distr. Nguyen Binh, municipality Nguyen Binh (N 22ø37' E 105ø52'), about 41 km to W from Cao Bang town.

Elevation from Le A pass to Na Ni mt. peak at 1400-1900 m alt. Old secondary closed evegreen wet mountain forest. Terrestrial herb on

open granite steep outcrops. Flowers white. Common.

No CBL 208 11-12 Nov. 1998

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 6300-98)

Collectors: Averyanov L., N.Q.Binh, N.T.Hiep, P.K.Loc, N.X.Tam.

Fam. Asteraceae

Aensleya sp.

N. Vietnam, prov. Cao Bang, distr. Nguyen Binh, municipality Nguyen Binh (N 22ø37' E 105ø52'), about 41 km to W from Cao Bang town.

Elevation from Le A pass to Na Ni mt. peak at 1400-1900 m alt. Old secondary closed evegreen wet mountain forest. Terrestrial herb up to

50 cm hg on open granite eutcrops. Common at 1700-1900 m alt.

No CBL 201 11-12 Nov. 1998

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 6300-98)

Collectors: Averyanov L., N.Q.Binh, N.T.Hiep, P.K.Loc, N.X.Tam.

Fam. Asteraceae

Aensleya sp.

N. Vietnam, prov. Ha Giang, distr. Dong Van, municipality Ho Quang Phin, vicinities of Ta Xa village, 14-16 km to SSW of Dong Van town (N

23ø16' E 105ø22'). Primary very wet mossy cloud mixed forest (with domination of Tsuga chinensis and Pinus kwangtungensis) along tops of

karst remnant limestone ridge at 1550-1600 m elev. Lithophyte herb on mossy shady vertical cliffs. Common.

No CBL 1788 28 April 1999

International Botanical Expedition of the U.S.A. National Geographic Society (grant No 6300-98)

Collectors: P.K.Loc, P.H.Hoang, Averyanov L.

Fam. Asteraceae

Ainsliaea sp.

Northern Vietnam, Lao Cai Prov., Van Ban Distr., Khanh Yen Ha municipality, to the SW of Na Nheo village (in the vicinity of point

21ø57'41''N, 104ø13'29''E). Primary evergreen broad-leaved and mixed montane forests (with Fokienia hodginsii) along granite and quartzite

ridge at elev. 2000-2100 m a.s.l. Terrestrial herb on steep open mossy slope. Flowers white, fragrant.

No HAL 2387 (Photo) 16 March 2002

Collectors: L.Averyanov, P.K.Loc, D.T.Doan (FFI Project).

Fam. Asteraceae

Ainsliaea sp.

Northern Vietnam, Lao Cai Prov., Van Ban Distr., Khanh Yen Ha municipality, to the SW of Na Nheo village (in the vicinity of point

21ø57'41''N, 104ø13'29''E). Primary evergreen broad-leaved and mixed montane forests (with Fokienia hodginsii) along granite and quartzite

ridge at elev. 2000-2100 m a.s.l. Terrestrial herb on steep open mossy slope. Flowers white, fragrant.

No HAL 2386 16 March 2002

Collectors: L.Averyanov, P.K.Loc, D.T.Doan (FFI Project).

Fam. Asteraceae

Ainsliaea sp.

Northern Vietnam, Lao Cai Prov., Van Ban Distr., Khanh Yen Ha municipality, to the SW of Na Nheo village (in the vicinity of point

21ø57'26''N, 104ø13'00''E). Primary evergreen broad-leaved montane forest with Fokienia hodginsii along granite/quartzite ridge at elev.

2200 m a.s.l. Terrestrial herb on very steep rocky slopes. Flowers white, fragrant. Common.

No HAL 2631 22 March 2002

Collectors: L.Averyanov, P.K.Loc, D.T.Doan (FFI Project).

Fam. Asteraceae

Ainsliaea sp.

Northern Vietnam, Lao Cai Prov., Van Ban Distr., to the west of Nam Xay village (21ø58'18''N, 104ø00'24''E). Primary coniferous (with

Fokienia hodginsii) evergreen montane forest at elev. 1700 m a.s.l. Terrestrial herb. Rare.

No HAL 2029 05 March 2002

Collectors: L.Averyanov, P.K.Loc, D.T.Doan (FFI Project).