Herbarium LE - Specimens

Solanaceae

Solanum camranhense Dy Phon et Hul

Ninh Thuan Province, Ninh Hai District, Vinh Hy Townlet. Very dry evergreen secondary scrub and short-tall wind-formed coastal forest on very steep sea faced sandstone rocky slopes and cliffs at elevation 100–200 m a.s.l. Erect terrestrial herb or undershrub to 1.5 m tall in open rocky place along road. Flowers white with light violet tint. Occasional.

15 May 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3233.

Typhaceae

Typha domingensis Pers.

(= T. angustata Bory & Chaub.)

Ninh Thuan Province, Ninh Hai District, Vinh Hy Townlet. Very dry evergreen secondary scrub and short-tall wind-formed coastal forest on very steep sea faced sandstone rocky slopes and cliffs at elevation 100–200 m a.s.l. Terrestrial and lithophytic herb to 1.8 m tall in wet open places along dry rocky stream. Rare.

15 May 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3232.

Cycadaceae

Cycas sp.

Ninh Thuan Province, Ninh Hai District, Vinh Hy Townlet. Very dry evergreen secondary scrub and short-tall wind-formed coastal forest on very steep sea faced sandstone rocky slopes and cliffs at elevation 150–200 m a.s.l. Palm like treelet to 4 m tall on steep rocky slopes. Not rare.

15 May 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3234.

Orchidaceae

Micropera pallida Lindl.

Khanh Hoa Province, Khanh Son District, Ta Gu Waterfall. Secondary evergreen, broad-leaved dry forest on granite along rocky stream at elevation 420–450 m a.s.l. Epiphyte. Occasional. 23 February 2023, L. Averyanov, Nguyen Van Canh, AL 2130. Living plants only.

Fl. in cult in 13 June 2024, L. Averyanov, AL2130.

Litus orientale maris Albi: Puschlachta

Kyrgyzstan, Jalal-Abad region, Toktogul

district, near the confluence of the Kara-

Luldzha (Alatay) and Uzunakhmat rivers,

rocky habitats with a predominace of

wormwood, altitude 1090 m, 41.930145° N,

72.572899° E, 12 V 2016, G. A. Lazkov

Кыргызстан, Джалал-Абадская область,

Токтогульский район, близ слияния рек

Кара-Кульджа (Алатай) и Узунахмат,

каменистые местообитания с

преобладанием полыни, высота 1090 м.,

41.930145° с.ш., 72.572899° в.д.,

12 V 2016, Г. А. Лазьков

Orchidaceae

Eulophia zollingeri J.J.Sm.

Gia Lai Province, near the Lam Truong 2 K'Bang, close to the Kon Chu Rang nature reserve. (Lâm trường, I am not sure about translation: like the place they grow tree to collect wood). Grow on decay wood, leafless orchid, under forest canopy. Lip with 2 short small lamellae in middle of lip.

11 June 2024, Truong Ba Vuong, Van Doan, BV 1897.

Arecaceae

Korthalsia laciniosa (Griff.) Mart.?

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Woody vine to 5 m long in shady humid place on rocky slope. Not rare.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2877 (photo only).

Convallariaceae

Tupistra wattii Hook.f.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Terrestrial creeping herb in shady place. Fruits red. Not rare.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2876e.

Asclepiadaceae

Dischidia acutifolia Maingay ex Hook.f.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Herbaceous vine 1-2 m long on mossy tree along ridge edge. Flowers dull light yellowish. Not rare.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2876.

Aceraceae

Acer tonkinense Lecomte

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l.

Tree to 20 m tall and 0.4 m in diameter along ridge edge. Young leaves dull reddish purple. Not rare.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2875.

Anacardiaceae

Pistacia weinmanniifolia J.Poiss. ex Franch.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l.

Tree to 25 m tall and 0.5 m in diameter along ridge edge. Not rare.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2874.

Celastraceae

Euonymus sp.?

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Treelet 2–2.5 m tall on shady rocky slope. Leaves uniform green. Occasional.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2873.

Celastraceae

Euonymus sp.?

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Treelet 2–2.5 m tall on shady rocky slope. Leaves above uniform green, below bright shiny purple. Very rare.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2872.

Sterculiaceae

Sterculia lanceolata Blume

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Tree to 10 m tall in rather open dry rocky slope. Flowers dull reddish. Common.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2871.

Stapgylleaceae?

Staphylea cochinchinensis?

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Erect terrestrial shrub or small treelet in rather open dry rocky slope. Occasiomnal.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2870.

Fabaceae (Mimosaceae)

Archidendron lucidum?

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l.

Tree about 10 m tall in shady place along rocky ridge edge. Flowers white. Common? Particularly in secondary forest.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2869.

Fabaceae

Ormosia sp.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Tree about 10 m tall in shady place along rocky ridge edge. Common.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2868.

Lauraceae

Cinnamomum sp.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Tree about 10 m tall in shady place along rocky ridge edge. Flowers light yellow. Occasional.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2867.

Magnoliaceae

Magnolia coco DC.?

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l.

Tree about 10 m tall. Flower buds green. Occasional.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2866.

Clusiaceae

Garcinia fragrans Elmer ex Merr.,

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l.

Tree to 25 m tall and to 0.6 m in diameter, with very hard timber. Young leaves dull purple brown, fruits green. Not rare.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2865.

Begoniaceae

Begonia sp.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Lithophytic herb on shady mossy vertical cliff. Flowers white to light pink, anthers and stigmas yellow. Leaves in young plants pale pink brown with white spots, leaves on flowering plants pure light green. Not rare.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2864.

Gesneriaceae

Paraboea nutans D.Fang & D.H.Qin

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Lithophytic herb on shady vertical cliff. Not common.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2863.

Pandanaceae

Pandanus sp.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Erect palm like plant with distinct internodes to 20 cm long ant stem to 2.5 m tall, and leaves to 3 m long in shady wet place between limestone hills. Not rare/

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2862.

Asteraceae

Vernonia sp.?

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Terrestrial herb to 1 m tall on old rice field. Flowers light pink purple. Common.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2861.

Podocarpaceae

Nageia wallichiana (C.Presl) Kuntze

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Treelet about 3 m tall in shady, humid rocky place. Rare.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2860.

Araceae

Aglaonema ovatum Engl.?

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Erect terrestrial and lithophytic herb to 0.5 m tall in shady, humid rocky place. Spathe and inflorescence white. Not rare.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2859.

Araceae

Amorphophallus sp.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Terrestrial and lithophytic herb with erect inflorescence to 30 cm tall in shady, humid rocky place. Peduncle pure white. Spathe pure light greenish. Occasional.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2858.

Araceae

Amorphophallus tonkinensis Engl. & Gehrm.?

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Terrestrial and lithophytic herb with erect inflorescence to 1 m tall in shady, humid rocky place. Peduncle dull olive green white spotted. Spathe pure light green, inflorescence axis pure white, male flowers white, female flowers light green. Occasional.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2857.

Orchidaceae

Corymborkis veratrifolia (Reinw.) Blume

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Terrestrial herb to 1.5 m tall in shady, humid rocky place. Not common.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2856.

Orchidaceae

Vanda sp.?

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Epiphyte on tall trees along ridge edge above tall cliff. Occasional.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2855.

Cactaceae

Opuntia sp.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Lithophytic and occasionally terrestrial herb to 1 m tall on and among open rosks. Locally abundant. Flowers fright yellow. Aggressive weed.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2854.

Zingiberaceae

Hedychium speciosum Wall.?

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Terrestrial and lithophytic herb to 1.2 m tall in shady place on very steep rocky slope. Fruita bright yellow orange, seeds bright red. Common.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2853.

Orchidaceae

Malaxis acuminata D.Don

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l.

Terrestrial and lithophytic herb in shady place on very steep rocky slope. Leaves dull purple. Flowers light green. Not common.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2852.

Piperaceae

Piper sp.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Erect terrestrial and lithophytic herb or undershrub to 1.5 tall in shady place on very steep rocky slope. Inflorescences white. Common.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2851.

Vitaceae

Tetrasigma sp.?

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Epiphytic vine about 1.5 m long in shady place on very steep rocky slope. Flower buds white. Not common.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2850.

Primulaceae

Lysimachia insignis Hemsl.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Terrestrial and occasionally lithophytic herb to 1 m tall on shady rocks on very steep rocky slope. Flowers yellow, fruits white, translucent. Not rare.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2849.

Convallariaceae

Disporum sp.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Terrestrial and occasionally lithophytic herb to 1 m tall on shady rocks along ridge edge above tall cliff. Flowers white. Not common.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2848.

Orchidaceae

Calanthe argenteo-striata C.Z.Tang & S.J.Cheng

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Lithophytic and occasionally terrestrial herb on shady rocks along ridge edge above tall cliff. Flowers white, later pale yellow. Leaves with contrast silvery stripes. Locally very common.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2847.

Lamiaceae

Leonurus sibiricus L.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Terrestrial herb to 1.2 m tall on old rice field. Flowers purple-violet. Occasional.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2846.

Selaginellaceae

Selaginella sp.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Terrestrial herb in shady wet place among big rocks Locally very common.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2845.

Commelinaceae

Pollia sp.

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Terrestrial and lithophytic herb in shady wet place among big rocks Flowers white. Locally common.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2844.

Cycadaceae

Cycas brachycantha K.D.Hill, T.H.Nguyen & L.K.Phan

Bac Kan Province, Ba Be District, Cao Thuong Community, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’20’’N 105º36’06’E at elevation of about 750 m a.s.l. Terrestrial palm-like plant with underground tuber-like stem and 1-5 leaves to 2 m long on very steep rocky slope. Young plants locally common. Mature samples not seen.

23 April 2024, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, T. Maisak, AL2843.