Herbarium LE - Specimens

Asclepiadaceae

Dischidia sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest along sea shore on limestone. Herbaceous vine to 1.5 m long on exposed rocks and vertical cliffs. Occasional.

17 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4372 (NHC172).

Asclepiadaceae

Hoya sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest along sea shore on limestone. Herbaceous vine to 3 m long on exposed rocks and vertical cliffs. Locally very common. Flowers white, corona red in center.

17 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4371 (NHC171).

Orchidaceae

Liparis distans C.B.Clarke

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone and limestone slope at elevation about 80-150 m a.s.l. along forest trail. Clustering herb on shady steep rocky slope. Not common.

17 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4370 (NHC170).

Gesneriaceae

Primulina drakei (B.L.Burtt) Mich.Möller & A.Weber

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone and limestone slope at elevation about 80-150 m a.s.l. along forest trail. Herbaceous plant with branching succulent stem to 0.5 m tall on open steep rocky slope. Not rare.

17 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4369 (NHC169).

Gesneriaceae

Primulina gemella (D.Wood) Yin Z.Wang

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone and limestone slope at elevation about 80-150 m a.s.l. along forest trail. Herbaceous rosulate herb on open steep rocky slope. Flowers white. Not common.

17 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4368 (NHC168).

Asclepiadaceae

Hoya parasitica (Wall. ex Hornem.) Wight?

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone and limestone slope at elevation about 80-150 m a.s.l. along forest trail. Herbaceous to semi-woody vine 3-4 m long on open rocky slope. Flowers white. Locally common.

17 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4367 (NHC167).

Rutaceae

Melicope sp.?

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone and limestone slope at elevation about 80-150 m a.s.l. along forest trail. Treelet 3-4 m tall on open rocky slope. Flowers white. Not common.

17 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4366 (NHC166).

Moraceae

Ficus sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone and limestone slope at elevation about 80-150 m a.s.l. along forest trail. Treelet to 5 m tall on open rocky slope. Syconia greenish to pink-purple. Not common.

17 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4365 (NHC165).

Annonaceae

Mitrephora maingayi Hook.f. & Thomson

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone and limestone slope at elevation about 80-150 m a.s.l. along forest trail. Treelet to 3 m tall on shady rocky slope. Flowers white. Not common.

17 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4364 (NHC164).

Orchidaceae

Vanilla aphylla Blume

Dak Lak Province, Lak District, Krong No Municipality, around point 12.1408 N 108.1325 E.

Dry evergreen broad-leaved secondary forest on hill slopes composed with granite at elevation 800–900 m a.s.l.

Epiphytic vine on small trees in dray secondary scrub on exposed granite cliffs along stream canyon. Locally common.

24 November 2022, L. Averyanov, Nguyen Van Canh, T. Maisak, Truong Ba Vuong, et al., AL 1754.

Fowered in cult. on 18 June 2025, L. Averyanov, T. Maisak, AL1754.

All photos © - L. Averyanov

Orchidaceae

Paphiopedilum emersonii Koop. & P.J.Cribb

Vietnam sine loc., sine date, sine coll.

Flowered in cultivation on 18 June 2025, L. Averyanov, AL4439.

Fam. Orchidaceae

Paphiopedilum concolor (Bateman) Pfitzer

N. Vietnam, prov. Son La, distr. Moc Chau, municipality Chieng Hac, vicinities of Co Liu village (N 20º51' E 104º38'). Remnants of primary evergreen broadleaved forests on vertical limestone bluffs along Xap river canyon at 550-600 m elev. Lithophyte forb on shady vertical limestone cliffs of N exposition. Flowers dull yellow; tepals and lip inside with few sparsely purple smallest dots. Leaves pale green with fine white tessellation above, dirty purple-violet spotted below. Not rare on inaccessible vertical bluffs.

No NTH 2922 28 May 1999.

Collectors: N.T.Hiep, P.H.Hoang, Averyanov L.

Flowered in cultivation on 18 June 2025, L. Averyanov, T. Maisak, NTH2922.

All photos © - L. Averyanov

Orchidaceae

Flickingeria parishii Seidenf.

Laos, s.loc.,

04 April 2015, T.M. Maisak, TM1252.

Flowered in cultivation on 18 June 2025, L. Averyanov, T. Maisak, TM1252.

Orchidaceae

Flickingeria sp.

Dak Nong Province, Dak Song District, Nam Nung Nature Reserve around point 12.1200N 107.4233E.

Fractionally logged primary evergreen broad-leaved and mixed forest with Dacrydium, Dacrycarpus and Pinus latteri on granite at elevation about 900 m a.s.l. Epiphyte on tall tree. Not rare.

27 November 2022, L. Averyanov, Nguyen Van Canh, T. Maisak, Truong Ba Vuong, Nguyen Quoc Bao, AL 1833.

Flowered in cultivation on 17 June 2025, L. Averyanov, AL1833.

Б.Н. Городков. Экспедиция в Тобольскую губернию. 1912 г. B. Gorodkov. Provintia Tobolsk, distr, Jschim. 1912.

Тобольская губ., Ишимский уезд, между с. Пигановским и д. Крашеневой. Солончак.

Orchidaceae

Vanilla baosangii Vuong, Aver. & V.C.Nguyen⁣

Di Linh District (or Duc Trong District).

17 June 2025, Nguyen Van Canh, AL4437.

Petrosaviaceae

Petrosavia stellaris Becc.

The border of Lam Dong and Khanh Hoa provinces, Khanh Le Pass, around point N12º10.7 E108º40.6. Remnants of primary evergreen broad-leaved forest on granite and gneiss at elevation about 1600 m a.s.l. Terrestrial achlorophyllous herb to 15 cm tall in shady place on mountain summit. Flowers white to white with yellowish tint. Not common.

17 Jube 2025, Nguyen Van Canh, AL4438.

Convallariaceae

Aspidistra sp.

Khanh Hoa Provinces, road to Khanh Le Pass, east slope of Hon Giao Ridge, dry evergreen broad-leaved forest on granite at elevation 600-700 m a.s.l. Lithophytic herb on steep rocky slope of small stream. Locally common. Flower buds dark violet. 21 April 2025, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3902.

Flowered in cultivation on 17 June 2025, Nguyen Van Canh, AL3902

Euphorbiaceae

Phyllanthus reticulatus Poir.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone and limestone slope at elevation about 80-150 m a.s.l. along forest trail. Shrub or treelet to 5 m tall on open rocky slope. Fruits reddish-purple. Not common.

17 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4363 (NHC163).

Ebenaceae

Diospyros sp. (Maba sp.)

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone and limestone slope at elevation about 80-150 m a.s.l. along forest trail. Treelet to 2 m tall in shady rocky slope. Flowers white. Not common.

17 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4362 (NHC162).

Convallariaceae

Polygonatum punctatum Royle ex Kunth?

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone and limestone slope at elevation about 80-150 m a.s.l. along forest trail. Terrestrial and lithophytic rhizomatous herb to 1m talon rocky slope. Flowers white. Not common.

17 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4361 (NHC161).

Euphorbiaceae

Suregada multiflora (A.Juss.) Baill.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone and limestone slope at elevation about 80-150 m a.s.l. along forest trail. Tree 6-7 m tall on limestone slope. Flower buds greenish Not common.

17 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4360 (NHC160).

Cyperaceae

Gahnia sp.?

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone and limestone slope at elevation about 80-150 m a.s.l. along forest trail. Terrestrial sedge in wet open place. Not common.

17 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4359 (NHC159).

Moraceae

Ficus sp.

Quang Ninh Province, Van Don District, Van Yen Commune, Bai Tu Long National Park, Cai Lim protection sector|, around point 21º06’30’’N 107º34’42’’E. Young secondary evergreen broad-leaved forest with bamboo on steep sandstone and limestone slope at elevation about 80-150 m a.s.l. along forest trail. Tree 6-7 m tall on limestone slope. Syconia green to yellowish-green. Common.

17 May 2025, Cuong Huu Nguyen, Ha Van Dang, Ly Van Nguyen, Nam Van Duong, Tuan Thanh Tran, Hai Xuan Cao, L. Averyanov, T. Maisak, AL4358 (NHC158).