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LE 01318680

LE 01318680

Adiantum caudatum L.⁣ det. Averyanov, Leonid Vladimirovich at 22.01.2026

Collectors: Tien Hiep Nguyen; Averyanov, Leonid Vladimirovich; et al. Field collecting number: LA-VN1312. Collection date: 11.04.2015.
Administrative regions: LA - Houaphan.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Laos
Original label text:
FLORA OF LAOS
Polypodiaceae s.l.
Adiantum sp.

Houphan Prov., Hem Distr., Na Puok village, Pu Tham Nhon mountain around point 20°10'00.0''N 103°24'37.0''E. Primary and secondary broad-leaved evergreen dry forest on very steep rocky mountain slopes composed by solid marble-like highly eroded limestone at elev. 1050-1150 m a.s.l. Terrestrial fern on steep shady slope. Common.
LA-VN 1312 11 April 2015
Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318680 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318680
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LE 01318679

LE 01318679

Adiantum philippense L.⁣ det. Averyanov, Leonid Vladimirovich at 22.01.2026

Collectors: Tien Hiep Nguyen; Averyanov, Leonid Vladimirovich; et al. Field collecting number: LA-VN1205. Collection date: 08.04.2015.
Administrative regions: LA - Houaphan.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Laos
Original label text:
FLORA OF LAOS
Polypodiaceae s.l.
Adiantum sp.

Houphan Prov., Viengxay Distr., Xieng Luong village, around point 20°22'06.1''N 104°17'18.6''E. Primary and secondary broad-leaved evergreen dry forest on very steep rocky slopes of highly eroded mountains composed by solid marble-like limestone at elev. 600-700 m a.s.l.
Terrestrial rosulate fern on shady rocky slope. Locally common.
LA-VN 1205 8 April 2015
Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Somneuk Bouthamounty.
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318679 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318679
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LE 01318677

LE 01318677

Davallia solida (G. Forst.) Sw.⁣ det. Averyanov, Leonid Vladimirovich at 22.01.2026

Collectors: Tien Hiep Nguyen; Averyanov, Leonid Vladimirovich; et al. Field collecting number: LA-VN1275. Collection date: 11.04.2015.
Administrative regions: LA - Houaphan.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Laos
Original label text:
FLORA OF LAOS
Polypodiaceae s.l.
Davallia sp.

Houphan Prov., Xon Distr., Nong Hua Pa Noi village, around point 20°13'02.1''N 103°24'23.3''E. Primary and secondary broad-leaved evergreen forest on very steep rocky mountain slopes composed by shaly limestone at elev. 1100-1200 m a.s.l. Creeping epiphyte on tall mossy tree along ridge edge. Locally very common.
LA-VN 1275 11 April 2015
Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318677 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318677
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LE 01318674

LE 01318674

Drynaria rigidula (Sw.) Bedd.⁣ ⟮prelim. ID⟯ det. Averyanov, Leonid Vladimirovich at 22.01.2026

Collectors: Tien Hiep Nguyen; Averyanov, Leonid Vladimirovich; et al. Field collecting number: LA-VN1228. Collection date: 11.04.2015.
Administrative regions: LA - Houaphan.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Laos
Original label text:
FLORA OF LAOS
Polypodiaceae s.l.
Pseudodrynaria sp.

Houphan Prov., Xon Distr., Nong Hua Pa Noi village, around point 20°13'02.1''N 103°24'23.3''E. Primary and secondary broad-leaved evergreen forest on very steep rocky mountain slopes composed by shaly limestone at elev. 1100-1200 m a.s.l. Short creeping epiphyte on tall trees. Very common.
LA-VN 1228 11 April 2015
Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318674 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318674
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LE 01318671

LE 01318671

Belvisia mucronata (Fée) Copel.⁣ ⟮prelim. ID⟯ det. Averyanov, Leonid Vladimirovich at 22.01.2026

Collectors: Tien Hiep Nguyen; Averyanov, Leonid Vladimirovich; et al. Field collecting number: LA-VN1339. Collection date: 11.04.2015.
Administrative regions: LA - Houaphan.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Laos
Original label text:
FLORA OF LAOS
Polypodiaceae s.l.
Belvisia sp.

Houphan Prov., Hem Distr., Na Puok village, Pu Tham Nhon mountain around point 20°10'00.0''N 103°24'37.0''E. Primary and secondary broad-leaved evergreen dry forest on very steep rocky mountain slopes composed by solid marble-like highly eroded limestone at elev. 1050-1150 m a.s.l. Epiphyte on old mossy trees among shady mossy rocks on mountain top. Common.
LA-VN 1339 11 April 2015
Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318671 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318671
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LE 01318670

LE 01318670

Asplenium rupestre Hope⁣ det. Tien Hiep Nguyen at 11.04.2015

Collectors: Tien Hiep Nguyen; Averyanov, Leonid Vladimirovich; et al. Field collecting number: LA-VN1338. Collection date: 11.04.2015.
Administrative regions: LA - Houaphan.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Laos
Original label text:
FLORA OF LAOS
Polypodiaceae s.l.
Asplenium rupestre Hope

Houphan Prov., Hem Distr., Na Puok village, Pu Tham Nhon mountain around point 20°10'00.0''N 103°24'37.0''E. Primary and secondary broad-leaved evergreen dry forest on very steep rocky mountain slopes composed by solid marble-like highly eroded limestone at elev. 1050-1150 m a.s.l. Terrestrial and epiphyte on old mossy trees among shady mossy rocks on mountain top. Common.
LA-VN 1338 11 April 2015
Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Hieu, T.Maisak, Pheng Phengsintham.
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318670 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318670
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LE 01318656

LE 01318656

Eria stricta Lindl.⁣ det. Averyanov, Leonid Vladimirovich at 12.01.2026

Collectors: Cuong, Huu Nguyen; et al. Field collecting number: AL4195. Collection date: 12.01.2026.
Administrative regions: VN - Thanh Pho Ha Noi.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Orchids of Vietnam, Laos and Cambodia
Original label text:
FLORA OF VIETNAM
Orchidaceae
Eria stricta Lindl.

Hanoi Capital Region, Ba Vi National Park, old humid secondary evergreen broad-leaved forest on very steep mountain slopes composed by granite and shale at elevation of 700-850 m a.s.l. Clustering epiphyte on tall mossy tree. Very common.
09 May 2025, Cuong Huu Nguyen, Son Van Le, Ha Van Dang, Ly Van Nguyen, L. Averyanov, T. Maisak, AL4195 (NHC006).

Flowered in culture on 12 January 2026, L. Averyanov, T. Maisak, AL 4195
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318656 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318656
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LE 01318655

LE 01318655

Aspidistra subrotata Y.Wan & C.C.Huang⁣ det. Averyanov, Leonid Vladimirovich at 05.01.2026

Collectors: Cuong, Huu Nguyen; et al. Field collecting number: AL3015. Collection date: 05.01.2026.
Administrative regions: VN - Tinh Thanh Hoa.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
FLORA OF VIETNAM
Convallariaceae
Aspidistra sp.

Thanh Hoa Province, Quan Hoa District, Nam Dong Community, Ban Lo Village, Ke Stream, the territory of Nam Dong National Reserve around point 22º25’46’’N 105º36’58’E. Old secondary broad-leaved evergreen humid forest on wet very steep alluvial slopes based on solid, marble-like highly eroded limestone at elevation of 600–800 m a.s.l. Terrestrial rhizomatous herb on steep shady rocky slope. Occasional.
04 May 2024, Nguyen Huu Cuong, Nguyen Van Ly, L. Averyanov, T. Maisak, AL3015 (living plants only).

Flowered in culture on 5 January 2026, L. Averyanov, AL 3015
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318655 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318655
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LE 01318654

LE 01318654

Nervilia muratana S.W.Gale & S.K.Wu⁣ det. Averyanov, Leonid Vladimirovich at 20.01.2026

Collectors: Averyanov, Leonid Vladimirovich; Nguyen Van Canh; Maisak, Tatiana Viktorovna Field collecting number: AL2704. Collection date: 20.01.2026.
Administrative regions: VN - Tinh Lam GJong.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Orchids of Vietnam, Laos and Cambodia
Original label text:
FLORA OF VIETNAM
Orchidaceae
Nervilia muratana S.W.Gale & S.K.Wu

Lam Dong Province, Di Linh District, Brah Yang Mountain, primary and old secondary evergreen broad-leaved and mixed forest (with Pinus khasya) on granite around point 11º27.669ʹ N 108º03.543ʹ E at elevation of 1000-1100 m a.s.l.Terrestrial her. Occasional.
14 November 2023, L. Averyanov, Nguyen Van Canh, T. Maisak, AL2704 (living plant only).

Flowered in culture 20 January 2026, L. Averyanov, AL 2704
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318654 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318654
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LE 01318653

LE 01318653

Campanulorchis longipes (Gagnep.) J.Ponert⁣ det. Averyanov, Leonid Vladimirovich at 12.01.2026

Collectors: Averyanov, Leonid Vladimirovich; et al. Field collecting number: AL1994. Collection date: 12.01.2026.
Administrative regions: VN - Tinh Lam GJong; VN - Tinh Khanh Hoa.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Orchids of Vietnam, Laos and Cambodia
Original label text:
FLORA OF VIETNAM
Orchidaceae
Eria longipes Gagnep.,

The border of Lam Dong and Khanh Hoa provinces, Khanh Le Pass, around point N12º10.7 E108º40.6. Remnants of primary evergreen broad-leaved forest on granite and gneiss at elevation about 600 m a.s.l. Creeping epiphyte on tall mossy trees. Very common.
4 December 2022, L. Averyanov, Nguyen Van Canh, T. Maisak, Nguyen Quoc Bao, AL 1994.

Flowered in culture on 12 January 2026, L. Averyanov, T. Maisak, AL 1994
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318653 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318653
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LE 01318652

LE 01318652

Tainia penangiana Hook.f.⁣ ⟮prelim. ID⟯ det. Averyanov, Leonid Vladimirovich at 12.02.2026

Collectors: Averyanov, Leonid Vladimirovich; Maisak, Tatiana Viktorovna Field collecting number: AL3144. Collection date: 12.01.2026.
Administrative regions: VN - Tinh GJak Lak.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Orchids of Vietnam, Laos and Cambodia
Original label text:
FLORA OF VIETNAM
Orchidaceae
Tainia penangiana Hook.f.

Dak Lak Province, Lak District, around Lieng Bok Sak Waterfall at elevation about 400 m a.s.l. Dry evergreen broad-leaved forest along rocky stream on granite. Terrestrial herb among rocks along stream. Occasional.
11 May 2024, L. Averyanov, Nguyen Van Canh, T. Maisak, AL3144.

Flowered in culture on 12 January 2026, L. Averyanov, T. Maisak, AL3144.
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318652 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318652
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LE 01318651

LE 01318651

Hypericum uralum D.Don⁣ det. Averyanov, Leonid Vladimirovich at 05.01.2026

Collectors: Дубенская Г.И. Field collecting number: s.n.. Collection date: 28.04.2025.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of India
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318651 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318651
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LE 01318649

LE 01318649

Pararuellia sp.⁣ det. Averyanov, Leonid Vladimirovich at 05.01.2026

Collectors: Averyanov, Leonid Vladimirovich; Nguyen Van Canh; Maisak, Tatiana Viktorovna Field collecting number: AL3944. Collection date: 01.04.2026.
Administrative regions: VN - Tinh GJak Lak.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
FLORA OF VIETNAM
Acanthaceae
Gen. ?

Dak Lak Province, M’Drak District, W foothills of Chu Mu Mt., dry secondary and remnants of primary evergreen broad-leaved forest on granite on steep slopes along rocky river at elevation 600-650 m a.s.l. Terrestrial adpressed rosulate herb on soil cliff in shady place along stream. Flowers light blue violet. Not rare.
25 April 2025, L. Averyanov, N.V. Canh, T. Maisak, AL3944.
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318649 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318649
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LE 01318648

LE 01318648

Melia azedarach L.⁣ det. Averyanov, Leonid Vladimirovich at 05.01.2026

Collectors: Дубенская Г.И. Field collecting number: s.n.. Collection date: 04.05.2019.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of India
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318648 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318648
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LE 01318647

LE 01318647

Smilacina sp.⁣ det. Averyanov, Leonid Vladimirovich at 05.01.2026

Collectors: Дубенская Г.И. Field collecting number: s.n.. Collection date: 20.05.2023.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of India
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318647 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318647
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LE 01318646

LE 01318646

Bellevalia sp.⁣ ⟮prelim. ID⟯ det. Averyanov, Leonid Vladimirovich at 05.01.2026

Collectors: Дубенская Г.И. Field collecting number: s.n.. Collection date: 30.04.2019.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of India
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318646 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318646
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LE 01318645

LE 01318645

Bellevalia sp.⁣ ⟮prelim. ID⟯ det. Averyanov, Leonid Vladimirovich at 05.01.2026

Collectors: Дубенская Г.И. Field collecting number: s.n.. Collection date: 30.04.2019.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of India
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318645 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318645
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LE 01318470

LE 01318470

Peliosanthes crassicoronata K.S.Nguyen, Aver. & N.Tanaka⁣ det. Averyanov, Leonid Vladimirovich at 22.12.2025

Collectors: Averyanov, Leonid Vladimirovich; Nguyen Van Canh; Maisak, Tatiana Viktorovna Field collecting number: AL2741a. Collection date: 22.12.2025.
Administrative regions: VN - Gia Lai.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318470 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318470
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LE 01318469

LE 01318469

Peliosanthes argenteostriata Aver. & N.Tanaka⁣ ⟮prelim. ID⟯ det. Averyanov, Leonid Vladimirovich at 22.12.2025

Collectors: Cuong, Huu Nguyen; et al. Field collecting number: AL2385. Collection date: 22.12.2025.
Administrative regions: VN - Tinh Bac Kan.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
FLORA OF VIETNAM
Convallariaceae
Peliosanthes sp.

Bac Kan Province, Ba Be District, Ba Be National Park. Primary and old secondary evergreen broad-leaved forest on very steep rocky slopes composed by white crystalline marble-like highly eroded limestone around point 22º27’26’’N 105º36’12’E at elev. of 600-800 m a.s.l.
Terrestrial herb in shady rocky place. Locally common.
30 October 2023, Nguyen Huu Cuong, Chu Ngoc Quan, L. Averyanov, Nguyen Van Ly, T. Maisak, AL2385.
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318469 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318469
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LE 01318439

LE 01318439

Pyrrosia nummulariifolia (Sw.) Ching⁣ det. Averyanov, Leonid Vladimirovich at 26.12.2025

Collectors: D.K. Harder; et al. Field collecting number: DKH6652. Collection date: 22.02.2001.
Administrative regions: VN - Tinh Lao Cai.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
POLYPODIACEAE
Pyrrosia


Lao Cai
Van Ban District; Liem Phu Municipality;
Khuoi Ngoa Village. Collections on
ridges just west of village.
21°59'31"N 104°19'05"E 323 - 1050 m
Epiphytic, vine. Fronds light green;
sporangia orange-brown; abaxial leaf
surfaces very light brown.

22 February 2001
D.K. Harder, P.K. Loc, L.V. Averyanov,
G.E. Schatz & S. Bodine 6652
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318439 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318439
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LE 01318438

LE 01318438

Polypodium diversum Rosenst.⁣ det. Averyanov, Leonid Vladimirovich at 26.12.2025

Collectors: D.K. Harder; et al. Field collecting number: DKH6944. Collection date: 26.02.2001.
Administrative regions: VN - Tinh Lao Cai.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
POLYPODIACEAE
Lepisorus


Lao Cai
Van Ban District; Nam Xe Municipality;
Nam Xi Tan Village. Collection in forest
on steep W-facing slope along stream;
just above village; on N side of road
between Nam Xe and Khau Co Pass at near
40 km to Than Uyen mark.
22°02'31"N 103°58'32"E 700 - 1113 m
Epiphytic, on slope in shade,
occasional, creeper. Rhizome green with
26 February 2001
D.K. Harder, P.K. Loc, N.T. Hiep, G.E.
Schatz, S. Bodine & N.Q. Hieu 6944
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
----------------------------------------
Page 2
brown hairs; sporangia light
yellow-green.
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318438 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318438
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LE 01318437

LE 01318437

Polypodium diversum Rosenst.⁣ det. Averyanov, Leonid Vladimirovich at 26.12.2025

Collectors: D.K. Harder; et al. Field collecting number: DKH6650. Collection date: 22.02.2001.
Administrative regions: VN - Tinh Lao Cai.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
POLYPODIACEAE
Lepisorus subrostratus (C. Chr.) C. Chr.
& Tardieu
det. P.K. Loc, 2001

Lao Cai
Van Ban District; Liem Phu Municipality;
Khuoi Ngoa Village. Collections on
ridges just west of village.
21°59'31"N 104°19'05"E 323 - 1050 m
Occasional, epiphytic. Pinnae light
green; sporangia light brown.
22 February 2001
D.K. Harder, P.K. Loc, L.V. Averyanov,
G.E. Schatz & S. Bodine 6650
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318437 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318437
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LE 01318436

LE 01318436

Colysis digitata Ching⁣ det. Averyanov, Leonid Vladimirovich at 26.12.2025

Collectors: D.K. Harder; et al. Field collecting number: DKH7115. Collection date: 02.03.2001.
Administrative regions: VN - Tinh Son La.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
POLYPODIACEAE
Colysis


Son La
Yen Chau District; Muong Lum
Municipality; Lum Village. Collections
on ridge S of village. Collected with
P.K. Loc, N.T. Hiep, L.V. Averyanov,
G.E. Schatz, S. Bodine & N.Q. Hieu.
21°00'47"N 104°29'30"E 900 - 1134 m
Occasional, lithophytic. Rhizome green;
rachis green; sporangia dark brown.

2 March 2001
D.K. Harder & et al. 7115
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318436 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318436
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LE 01318435

LE 01318435

Colysis digitata Ching⁣ det. Averyanov, Leonid Vladimirovich at 26.12.2025

Collectors: D.K. Harder; et al. Field collecting number: DKH7215. Collection date: 03.03.2001.
Administrative regions: VN - Tinh Son La.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
POLYPODIACEAE
Colysis


Son La
Yen Chau District; Muong Lum
Municipality; Lum Village. Collection on
ridges S of village. Collected with P.K.
Loc, N.T. Hiep, B. Tan, G.E. Schatz,
L.V. Averyanov & S. Bodine.
21°00'17"N 104°29'08"E 1242 m
Occasional, lithophytic, on mossy rocks.
Rachis black; sporangia dark dull
black-brown; rhizome green with brown
hairs.
3 March 2001
D.K. Harder & et al. 7215
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318435 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318435
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LE 01318434

LE 01318434

Colysis digitata Ching⁣ det. Averyanov, Leonid Vladimirovich at 26.12.2025

Collectors: D.K. Harder; et al. Field collecting number: DKH6146. Collection date: 11.02.2001.
Administrative regions: VN - Tinh Ha Giang.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
POLYPODIACEAE
Colysis


Ha Giang
Quan Ba District; Bat Dai Son
Municipality; Bat Dai Son Provincial
Protected Area (2000); valley just W of
Bat Dai Son border station; valley of
slash and burn agriculture, then higher
up, degraded secondary vegetation in
shade.
23°09'04"N 104°59'53"E 1000 m

Occasional, among rocks. Hair on rhizome
11 February 2001
D.K. Harder, P.K. Loc, L.V. Averyanov,
G.E. Schatz & S. Bodine 6146
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)


----------------------------------------
Page 2
brown; sporangia dark brown.
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318434 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318434
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LE 01318433

LE 01318433

Colysis digitata Ching⁣ det. Averyanov, Leonid Vladimirovich at 26.12.2025

Collectors: D.K. Harder; et al. Field collecting number: DKH7412. Collection date: 07.03.2001.
Administrative regions: VN - Tinh Son La.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
POLYPODIACEAE
Colysis

Son La
Moc Chau District; Van Ho Municipality,
Hua Tat Village. Collections on
limestone ridge from degraded forests to
cultivation on lower slopes to primary
broadleaved, closed wet forest along
limestone ridge and slopes near tops.
20°46'13"N 104°46'12"E 1100 - 1300 m

Lithophyte herb in shady wet places.
Common.
7 March 2001
D.K. Harder, P.K. Loc, L.V. Averyanov &
N.X. Tam 7412
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318433 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318433
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LE 01318432

LE 01318432

Colysis digitata Ching⁣ det. Averyanov, Leonid Vladimirovich at 26.12.2025

Collectors: Averyanov, Leonid Vladimirovich; et al. Field collecting number: VH6040. Collection date: 11.05.2000.
Administrative regions: VN - Tinh GJak Lak.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
FLORA OF VIETNAM
Polypodiaceae s.l.
Asplenium sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of
Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary wet closed mountain broad-leaved and mixed forest on
granites along very steep shady slopes of river canyon at elev. 1500-1700 m. Creeping epiphyte at the base of mossy trees. Very common.
No VH 6040 11.05.2000
International Botanical Expedition of the U.S.A. National Geographic
Society (grant No 6383-98)
___________________________________________________________
Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318432 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318432
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LE 01318431

LE 01318431

Colysis digitata Ching⁣ det. Averyanov, Leonid Vladimirovich at 26.12.2025

Collectors: Averyanov, Leonid Vladimirovich; et al. Field collecting number: VH6240. Collection date: 07.05.2000.
Administrative regions: VN - Tinh GJak Lak.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
FLORA OF VIETNAM
Polypodiaceae s.l.
Asplenium sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of
Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary closed wet broad-leaved and mixed forest (with Pinus,
Fokienia, Dacrydium and Dacrycarpus) on steep granite slopes at elev. 1200-1300 m. Creeping epiphyte at base of large trees. Very common.
No VH 6240 08.05.2000
International Botanical Expedition of the U.S.A. National Geographic
Society (grant No 6383-98)
___________________________________________________________
Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318431 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318431
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LE 01318430

LE 01318430

Plagiogyria adnata (Blume) Bedd.⁣ det. Averyanov, Leonid Vladimirovich at 26.12.2025

Collectors: Averyanov, Leonid Vladimirovich; et al. Field collecting number: VH6368. Collection date: 11.05.2000.
Administrative regions: VN - Tinh GJak Lak.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
FLORA OF VIETNAM
Polypodiaceae s.l.
Plagyogeria sp. ?

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of
Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary wet closed mountain broad-leaved and mixed forest on
granites along very steep shady slopes of river canyon at elev. 1700-1750 m. Terrestrial fern on very steep shady slopes of river
canyon. Not rare.
No VH 6368 11.05.2000
International Botanical Expedition of the U.S.A. National Geographic
Society (grant No 6383-98)
___________________________________________________________
Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318430 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318430
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LE 01318429

LE 01318429

Plagiogyria adnata (Blume) Bedd.⁣ det. Averyanov, Leonid Vladimirovich at 26.12.2025

Collectors: Averyanov, Leonid Vladimirovich; et al. Field collecting number: VH6248. Collection date: 07.05.2000.
Administrative regions: VN - Tinh GJak Lak.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
FLORA OF VIETNAM
Polypodiaceae s.l.
Plagyogeria sp. ?

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of
Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary closed wet broad-leaved and mixed forest (with Pinus,
Fokienia, Dacrydium and Dacrycarpus) on steep granite slopes at elev. 1200-1550 m. Terrestrial fern on shady very steep slope of river
canyon. Not rare.
No VH 6248 07.05.2000
International Botanical Expedition of the U.S.A. National Geographic
Society (grant No 6383-98)
___________________________________________________________
Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318429 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318429
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LE 01318428

LE 01318428

Asplenium nidus L.⁣ det. Averyanov, Leonid Vladimirovich at 26.12.2025

Collectors: D.K. Harder; et al. Field collecting number: DKH6464. Collection date: 17.02.2001.
Administrative regions: VN - Tinh Ha Giang.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
ASPLENIACEAE
Asplenium

Ha Giang
Vi Xuyen District; Minh Tan Municipality
Ban Ngay Village. Collection on E
facing ridge above river; on exposed
limestone rocks; collections in
disturbed agricultural land leading into
primary forest with some deforestation
of Burretiodendron (Tiliaceae) trees.
23°00'09"N 104°55'21"E 225 - 621 m
23°00'37"N 104°54'46"E
Frequent epiphyte and occasional
17 February 2001
D.K. Harder, P.K. Loc, N.T. Hiep, G.E.
Schatz & S. Bodine 6464
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
----------------------------------------
Page 2
lithophyte. Fronds light green;
sporangia dull, light brown; midrib of
frond purple-brown, producing "nests"
when large.
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318428 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318428
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LE 01318427

LE 01318427

Asplenium antrophyoides Christ⁣ det. Averyanov, Leonid Vladimirovich at 26.12.2025

Collectors: D.K. Harder; et al. Field collecting number: DKH7392. Collection date: 07.03.2001.
Administrative regions: VN - Tinh Son La.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
ASPLENIACEAE
Asplenium


Son La
Moc Chau District; Van Ho Municipality,
Hua Tat Village. Collections on
limestone ridge from degraded forests to
cultivation on lower slopes to primary
broadleaved, closed wet forest along
limestone ridge and slopes near tops.
20°46'13"N 104°46'12"E 1100 - 1300 m
Lithophytic fern. Common.
7 March 2001
D.K. Harder, P.K. Loc, L.V. Averyanov &
N.X. Tam 7392
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318427 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318427
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LE 01318426

LE 01318426

Asplenium antrophyoides Christ⁣ det. Averyanov, Leonid Vladimirovich at 26.12.2025

Collectors: Cuong, Huu Nguyen Field collecting number: CLL592. Collection date: 2019.
Administrative regions: VN - Tinh Thanh Hoa.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
FLORA OF VIETNAM
Asplenium antrophyoides Christ

Northern Vietnam, Thanh Hoa, Pu Luong Nature Reserve.
2019
Coll. : Cuong Huu Nguyen CLL 529
Det. Averyanov L. 26.12.2025
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318426 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318426
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LE 01318425

LE 01318425

Prosaptia barathrophylla (Baker) M.G.Price⁣ det. Averyanov, Leonid Vladimirovich at 26.12.2025

Collectors: D.K. Harder; et al. Field collecting number: DKH7411. Collection date: 07.03.2001.
Administrative regions: VN - Tinh Son La.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
GRAMMITIDACEAE
Grammitis


Son La

Moc Chau District; Van Ho Municipality,
Hua Tat Village. Collections on
limestone ridge from degraded forests to
cultivation on lower slopes to primary
broadleaved, closed wet forest along
limestone ridge and slopes near tops.
20°46'13"N 104°46'12"E 1100 - 1300 m

Epiphyte in shady places. Common.

7 March 2001
D.K. Harder, P.K. Loc, L.V. Averyanov &
N.X. Tam 7411
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318425 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318425
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LE 01318424

LE 01318424

Prosaptia barathrophylla (Baker) M.G.Price⁣ det. Averyanov, Leonid Vladimirovich at 26.12.2025

Collectors: D.K. Harder; et al. Field collecting number: DKH7335. Collection date: 06.03.2001.
Administrative regions: VN - Tinh Son La.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
GRAMMITIDACEAE
Grammitis

Son La
Moc Chau District; Van Ho Municipality,
Hua Tat Village. Collections on
limestone ridge from degraded forests to
cultivation on lower slopes to primary
evergreen, closed forest on limestone
ridge and cliffs, some coniferous forest
with Pinus kwangtungensis and Calocedrus
macrolepis.
20°46'35"N 104°46'31"E 1400 - 1500 m
6 March 2001
D.K. Harder, P.K. Loc, L.V. Averyanov &
N.X. Tam 7335
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
---------------------------------------
Page 2
Epiphyte in shady place. Not rare.
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318424 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318424
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LE 01318423

LE 01318423

Prosaptia barathrophylla (Baker) M.G.Price⁣ det. Averyanov, Leonid Vladimirovich at 26.12.2025

Collectors: D.K. Harder; et al. Field collecting number: DKH6092. Collection date: 04.05.2000.
Administrative regions: VN - Tinh GJak Lak.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
POLYPODIACEAE
Pyrrosia

Ha Giang
Quan Ba District; Bat Dai Son
Municipality; Bat Dai Son Provincial
Protected Area (2000). On limestone
ridge of degraded forest just northwest
of Bai Dai Son border station;
collections in forest above cultivations
(Forest possibly burned ca. 25 years ago
or so, only small trees and few fire
tolerant herbs).
13°09'13"N 104°59'36"E 1064 m
10 February 2001
D.K. Harder, Hiep, P.K. Loc, L.V.
Averyanov, G.E. Schatz & S.A. Bodin 6092
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
----------------------------------------
Page 2
Epiphytic. Fronds light green; sporangia
rusty-brown.
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318423 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318423
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LE 01318422

LE 01318422

Prosaptia barathrophylla (Baker) M.G.Price⁣ det. Averyanov, Leonid Vladimirovich at 26.12.2025

Collectors: D.K. Harder; et al. Field collecting number: DKH6359. Collection date: 15.02.2001.
Administrative regions: VN - Tinh Ha Giang.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
GRAMMITIDACEAE
Grammitis

Ha Giang
Vi Xuyen District; Phong Quang
Municipality; Man Village. Limestone
peaks and ridges NE of Village.
22°53'21"N 104°56'45"E 595 - 787 m
Rare, lithophytic, in shade. Sporangia
brown.

15 February 2001
D.K. Harder, P.K. Loc, L.V. Averyanov,
G.E. Schatz & S. Bodine 6359
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318422 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318422
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LE 01318421

LE 01318421

Prosaptia barathrophylla (Baker) M.G.Price⁣ det. Averyanov, Leonid Vladimirovich at 26.12.2025

Collectors: D.K. Harder; et al. Field collecting number: DKH7488. Collection date: 08.03.2001.
Administrative regions: VN - Tinh Son La.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
GRAMMITIDACEAE
Grammitis

Son La
Moc Chau District; Long Luong
Municipality; Sang Cai Village.
Collections in primary, wet,
broad-leaved, evergreen and
semi-deciduous, mossy forest along
limestone ridge situated between
1400-1500 m elevation.
20°45'51"N 104°51'26"E 1300 - 1500 m
Epiphyte in shady places. Common.
8 March 2001
D.K. Harder, P.K. Loc, L.V. Averyanov &
N.X. Tam 7488
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318421 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318421
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LE 01318420

LE 01318420

Prosaptia barathrophylla (Baker) M.G.Price⁣ det. Averyanov, Leonid Vladimirovich at 26.12.2025

Collectors: D.K. Harder; et al. Field collecting number: DKH7261. Collection date: 04.03.2001.
Administrative regions: VN - Tinh Son La.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
Grammitideae C. Presl

Son La
Yen Chau District; Muong Lum
Municipality; On Oc Village; collection
along N-S ridge along western side.
Collection along trail from agricultural
field to secondary vegetation to primary
forest vegetation with some trees
removed along ridge top. Limestone
substrate (crystalline). GPS coordinates
at top of ridge.
20°59'06"N 104°28'46"E 1468 m
4 March 2001
D.K. Harder, P.K. Loc & N.T. Huong 7261
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
----------------------------------------
Page 2
Epiphytic. Root brown; raised sporangiabrown.
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318420 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318420
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LE 01318419

LE 01318419

Prosaptia barathrophylla (Baker) M.G.Price⁣ det. Averyanov, Leonid Vladimirovich at 25.12.2025

Collectors: D.K. Harder; et al. Field collecting number: DKH6379. Collection date: 15.02.2001.
Administrative regions: VN - Tinh Ha Giang.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM

Pteridophyta J.Y. Bergen & B.M. Davis

Ha Giang
Vi Xuyen District; Phong Quang
Municipality; Man Village. Limestone
peaks and ridges NE of Village.
22°53'21"N 104°56'45"E 595 - 787 m

Epiphytic, occasional, near top of small
hill. Rhizome and roots dark, dull
brown; sporangia light brown in small
light green cups.
15 February 2001
D.K. Harder, P.K. Loc, L.V. Averyanov,
G.E. Schatz & S. Bodine 6379
Supported by National Science Foundation
and National Geographic Society
MISSOURI BOTANICAL GARDEN HERBARIUM (MO)
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318419 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318419
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LE 01318418

LE 01318418

Prosaptia barathrophylla (Baker) M.G.Price⁣ det. Averyanov, Leonid Vladimirovich at 25.12.2025

Collectors: Averyanov, Leonid Vladimirovich; et al. Field collecting number: VH6021. Collection date: 05.05.2000.
Administrative regions: VN - Tinh GJak Lak.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
FLORA OF VIETNAM
Polypodiaceae
Asplenium sp. ?

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) along Dak Tour
river on N macroslope of Chu Yang Sinh mt system to main peak (12ш24'N, 108ш26'E). Primary broad-leaved mountain evergreen closed
forest on granites along river slopes at elev. 1000-1200 m. Epiphyte fern with pendent fronds up to 30 cm lg on mossy trees along shady
waterfall sides. Very common.
No VH 6021 05-06.05.2000
International Botanical Expedition of the U.S.A. National Geographic
Society (grant No 6383-98)
___________________________________________________________
Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318418 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318418
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LE 01318417

LE 01318417

Prosaptia barathrophylla (Baker) M.G.Price⁣ det. Averyanov, Leonid Vladimirovich at 25.12.2025

Collectors: Averyanov, Leonid Vladimirovich; et al. Field collecting number: VH6215. Collection date: 09.05.2000.
Administrative regions: VN - Tinh GJak Lak.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
FLORA OF VIETNAM
Polypodiaceae s.l.
Asplenium sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of
Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary mossy cloud mixed forest (with Pinus, Fokienia,
Dacrydium and Dacrycarpus) on very steep granite slopes at elev. 2100-2300 m. Epiphyte on mossy trees in shady place. Very common.
VH 6215 09.05.2000
International Botanical Expedition of the U.S.A. National Geographic
Society (grant No 6383-98)
___________________________________________________________
Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318417 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318417
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LE 01318416

LE 01318416

Pyrrosia adnascens (Sw.) Ching⁣ det. Averyanov, Leonid Vladimirovich at 25.12.2025

Collectors: Kudryavtzeva, Ekaterina Petrovna; Ogureeva G. Field collecting number: Kudryavtzeva728. Collection date: 12.10.1990.
Administrative regions: VN - Tinh Quang Ninh.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
POLYPODIACEAE
Pyrrosia adnascens (Sw.) Ching

Quang Ninh Prov.
Kudryavtzeva E., Ogureeva G.
Kudryavtzeva 728, 12.10.1990
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318416 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318416
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LE 01318415

LE 01318415

Pyrrosia adnascens (Sw.) Ching⁣ det. Averyanov, Leonid Vladimirovich at 25.12.2025

Collectors: Kudryavtzeva, Ekaterina Petrovna; Ogureeva G. Field collecting number: Kudryavtzeva728. Collection date: 12.10.1990.
Administrative regions: VN - Tinh Quang Ninh.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
POLYPODIACEAE
Taenitis blechnoides (Willd.) Sw.⁣

Quang Ninh Prov.
Kudryavtzeva E., Ogureeva G.
Kudryavtzeva 728, 12.10.1990
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318415 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318415
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LE 01318414

LE 01318414

Pyrrosia adnascens (Sw.) Ching⁣ ⟮prelim. ID⟯ det. Averyanov, Leonid Vladimirovich at 25.12.2025

Collectors: Kudryavtzeva, Ekaterina Petrovna; Ogureeva G. Field collecting number: Kudryavtzeva987. Collection date: 19.10.1990.
Administrative regions: VN - Tinh Quang Ninh.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
POLYPODIACEAE
Pyrrosia adnascens (Sw.) Ching

Quang Ninh Prov.
Kudryavtzeva E., Ogureeva G.
Kudryavtzeva 987, 19.10.1990
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318414 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318414
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LE 01318413

LE 01318413

Pyrrosia adnascens (Sw.) Ching⁣ det. Averyanov, Leonid Vladimirovich at 25.12.2025

Collectors: Kudryavtzeva, Ekaterina Petrovna; Ogureeva G. Field collecting number: Kudryavtzeva1062. Collection date: 20.10.1990.
Administrative regions: VN - Tinh Quang Ninh.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
POLYPODIACEAE
Pyrrosia adnascens (Sw.) Ching

Quang Ninh Prov.
Kudryavtzeva E., Ogureeva G.
Kudryavtzeva 1062, 20.10.1990
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318413 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318413
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LE 01318412

LE 01318412

Taenitis blechnoides (Willd.) Sw.⁣ det. Averyanov, Leonid Vladimirovich at 25.12.2025

Collectors: Kudryavtzeva, Ekaterina Petrovna; Ogureeva G. Field collecting number: Kudryavtzeva679. Collection date: 11.10.1990.
Administrative regions: VN - Tinh Quang Ninh.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
VIETNAM
POLYPODIACEAE
Taenitis blechnoides (Willd.) Sw.⁣

Quang Ninh Prov.
Kudryavtzeva E., Ogureeva G.
Kudryavtzeva 679, 11.10.1990
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318412 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318412
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LE 01318411

LE 01318411

Taenitis blechnoides (Willd.) Sw.⁣ det. Averyanov, Leonid Vladimirovich at 25.12.2025

Collectors: Averyanov, Leonid Vladimirovich; et al. Field collecting number: VH6291. Collection date: 06.05.2000.
Administrative regions: VN - Tinh GJak Lak.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
FLORA OF VIETNAM
Polypodiaceae s.l.
Pteris sp.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of
Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary closed wet broad-leaved forest on steep granite slopes
at elev. 1300 m. Terrestrial fern on steep shady slope. Not common.
No VH 6291 07.05.2000
International Botanical Expedition of the U.S.A. National Geographic
Society (grant No 6383-98)
___________________________________________________________
Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318411 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318411
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LE 01318410

LE 01318410

Woodwardia harlandii Hook⁣ det. Averyanov, Leonid Vladimirovich at 25.12.2025

Collectors: Averyanov, Leonid Vladimirovich; et al. Field collecting number: VH6290. Collection date: 06.05.2000.
Administrative regions: VN - Tinh GJak Lak.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
FLORA OF VIETNAM
Polypodiaceae s.l.
Woodwardtia sp. ?

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of
Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary closed wet broad-leaved forest on steep granite slopes
at elev. 1300 m. Terrestrial fern on steep shady slope. Not common.
No VH 6290 07.05.2000
International Botanical Expedition of the U.S.A. National Geographic
Society (grant No 6383-98)
___________________________________________________________
Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318410 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318410
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LE 01318409

LE 01318409

Leptochilus chilangensis (V.N.Tu) Liang Zhang & Li Bing Zhang⁣ det. Averyanov, Leonid Vladimirovich at 25.12.2025

Collectors: Averyanov, Leonid Vladimirovich; et al. Field collecting number: VH6070. Collection date: 04.05.2000.
Administrative regions: VN - Tinh GJak Lak.
Groups of specimens: L. V. Averyanov specimens; L. V. Averyanov herbarium; Flora of Vietnam
Original label text:
FLORA OF VIETNAM
Polypodiaceae s.l.

S. Vietnam, Dak Lac Prov., Krong Bong Distr., Cu Pui Municipality, elevation from Dak Tour village (12ш30'N, 108ш30'E) to main peak of
Chu Yang Sinh mt system (12ш24'N, 108ш26'E) along Dak Tour river (N slope). Primary wet closed mountain broad-leaved forest on granites
along river slopes at elev. 500-900 m. Lithophytic fern on shady mossy river rocks. Common.
No VH 6070 04.05.2000
International Botanical Expedition of the U.S.A. National Geographic
Society (grant No 6383-98)
___________________________________________________________
Collectors: L.Averyanov, N.T.Hiep, N.Q.Hieu, P.H.Hoang, D.X.Du, N.T.Vinh
Record creation: 2026-01-26 Tatiana Maisak, PhotoScan D2
Citation: Specimen LE 01318409 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01318409
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