Herbarium LE - Specimens

Euphorbiaceae

Breynia phuongiana N.K.T.Tram, Vuong, Aver. & V.C.Nguyen

Originally collected in Dak Lak Province, Ea H'leo District, evergreen forest at elevation of about 700 m a.s.l., 15 Feb. 2023, Nguyen Van Canh, AL 2456

Clonotype. Collected in Nguyen Van Canh Garden on 26 March 2026, Nguyen Van Canh, L. Averyanov, Truong Ba Vuong, T. Maisak, AL4552.

Convallariaceae

Aspidistra opaca Tillich

Dac Lak Province, Lac District, Evergreen broad-leaved forest on very steep slopes along stream. Locally very common. Flowers dark violet, stigma white.

Collected in Nguyen Van Canh Garden on 26 March 2026, Nguyen Van Canh, L. Averyanov, Truong Ba Vuong, T. Maisak, AL4550.

Convallariaceae

Peliosanthes “subsessiliflora” sp. new.? (P. micrantha aff.)

Cultivated plant originated from Binh Dinh Province, Van Canh District, lowland forest along stream, at elevation of about 300 m a.s.l., locally common.

Collected in Nguyen Van Canh Garden on 26 March 2026, Nguyen Van Canh, L. Averyanov, Truong Ba Vuong, T. Maisak, AL4548. Flowers dark violet, fruits glossy blue.

Selaginellaceae

Selaginella sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900-1000 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial erect to sub-prostrate herb to 0.5 m tall in open place along trail. Common.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4621.

Rubiaceae

Gaertnera sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900-1000 m a.s.l. around the point 12º10’N 107º46’E. Treelet or shrub to 2 m tall in open place along trail. Flowers white. Not common.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4620.

Rubiaceae

Brachytome walichii Hook.f

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900-1000 m a.s.l. around the point 12º10’N 107º46’E. Treelet or shrub to 2.5 m tall in open place along trail. Flowers pale pink to pink orange. Not rare.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4619.

Apocynaceae

Wrightia dubia (Sims) Spreng.?

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900-1000 m a.s.l. around the point 12º10’N 107º46’E. Treelet or shrub to 3 m tall in open place along trail. Flowers red to red orange. Not rare.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4618.

Pandanaceae

Pandanus sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Wetland thicket in marshland depression with domination of Pandanus and Livistona along stream. Palm-like tree to 4 m tall and leaves to 6 m long. Locally common, co-dominant of plant community.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4610.

Convallariaceae

Tupistra thangii N.Tanaka & D.D.Nguyen?

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial herb with erect stem bearing many stout stilt roots on shady stream slope. Locally common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4609.

Oleaceae

Jasminum sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Scandent shrub of semi-woody vine to 3 m long in open place along road. Flowers white. Common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4606.

Potamogetonaceae

Aponogeton robinsonii A.Camus?

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Aquatic herb with immersed linear leaves in shallow slow water of rocky forest stream. Flowers white. Locally common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4605.

Potamogetonaceae

Aponogeton robinsonii A.Camus?

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Aquatic herb with immersed linear leaves in shallow slow water of rocky forest stream. Flowers white. Locally common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4605.

Acanthaceae

Acanthus sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Creeping ascending herb on shady wet stream slope. Flowers white. Locally very common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4604.

Rubiaceae

Psychotria langbianensis Wernham

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial herb to 1 m tall in rather open place along trail. Flowers white. Not rare.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4603.

Selaginellaceae

Selaginella sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Wetland thicket in marshland depression with domination of Pandanus and Livistona along stream. Terrestrial scandent herb to 1.5 m long in wet open place. Common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4602.

Rubiaceae

Brachytome walichii Hook.f?

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Treelet or shrub to 2 m tall in rather open place along trail. Flowers yellow to yellow orange. Occasional.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4601.

Araceae

Typhonium sp. “simplicifolium”

Dac Lak Province without location. Plant cultivated in Nguyen Van Canh Garden. Spathe reddish brown about 3 cm long, inflorescence very short.

July 2025, Nguyen Van Canh, AL4561.

Orchidaceae

Dendrobium sp.

Gia Lai Province, K'Bang District. Highly degraded secondary evergreen broad-leaved dry forest at elevation 600-800 m a.s.l., around point 14º26N 108º22E. Clustering epiphyte on tall tree. Not rare.

7 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4810.

Passifloraceae

Adina sp.?

Gia Lai Province, K'Bang District. Secondary and remnants of primary evergreen broad-leaved dry forest with Dacrydium on lava rocks (red soils) at elevation about 1000 m a.s.l., around point 14º15N 108º35E. Herbaceous vine 3-4 m long in open place. Fruits bright red. Not common.

6 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4785.

Solanaceae

Solanum sp.

Gia Lai Province, K'Bang District. Highly degraded secondary evergreen broad-leaved dry forest at elevation 600-800 m a.s.l., around point 14º26N 108º22E. Shrub to 2 m tall in open place along road. Flowers white, anthers yellow. Common weed in the area

7 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4764.

Zingiberaceae

Alpinia sp.

Gia Lai Province, K'Bang District. Highly degraded secondary evergreen broad-leaved dry forest at elevation 600-800 m a.s.l., around point 14º26N 108º22E. Terrestrial herb to 3 m tall in open place along road. Flowers white, median tepal red. Common.

7 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4767.

Selaginellaceae

Selaginella sp.

Gia Lai Province, K'Bang District. Secondary and remnants of primary evergreen broad-leaved dry forest with Dacrydium on lava rocks (red soils) at elevation about 1000 m a.s.l., around point 14º15N 108º35E. Prostrate herb on shady slope along trail. Common.

6 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4781.

Podocarpaceae

Dacrydium elatum (Roxb.) Wall.

Gia Lai Province, K'Bang District. Highly degraded secondary evergreen broad-leaved dry forest at elevation 600-800 m a.s.l., around point 14º26N 108º22E. Sapling about 2 m tall of tree to 35 m tall. Former co-dominant of the first forest stratum.

7 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4778.

Selaginellaceae

Selaginella sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Wetland thicket in marshland depression with domination of Pandanus and Livistona along stream. Terrestrial scandent herb to 2 m long in wet open place. Common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4600.

Selaginellaceae

Selaginella sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 900 m a.s.l. around the point 12º10’N 107º46’E. Wetland thicket in marshland depression with domination of Pandanus and Livistona along stream. Terrestrial scandent herb to 2 m long in wet open place. Common.

28 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4600.

Convallariaceae

Aspidistra sp. (“pumila”)

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 1000-1200 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial rhizomatous herb in shady place. Not rare.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4639.

Convallariaceae

Aspidistra paucitepala Vislobokov, Nuraliev & D.D.Sokoloff

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 1000-1200 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial herb with erect stem bearing numerous stilt roots. Fruits dirty olive. Locally abundant.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4638.

Convallariaceae

Aspidistra paucitepala Vislobokov, Nuraliev & D.D.Sokoloff

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 1000-1200 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial herb with erect stem bearing numerous stilt roots. Fruits dirty olive. Locally abundant.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4638.

Zingiberaceae

Etlingera sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 1000-1200 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial herb to 2 m tall in shady place. Flowers bright red. Occasional.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4627.

Zingiberaceae

Etlingera sp.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 1000-1200 m a.s.l. around the point 12º10’N 107º46’E. Terrestrial herb to 2 m tall in shady place. Flowers bright red. Occasional.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4627.

Podocarpaceae

Podocarpus annamiensis N.E.Gray

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 1000-1200 m a.s.l. around the point 12º10’N 107º46’E. Treelet about 2 m tall on steep shady slope. Tree to 20 m tall. Occasional.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4626.

Pinaceae

Keteleeria evelyniana Mast.

Dak Nong Province, Dak Song District, Quang Son Commune, Nam Nung Nature Reserve, primary dry evergreen broad-leaved forest on basalt and lava rocks at elevation of about 1000-1200 m a.s.l. around the point 12º10’N 107º46’E. Treelet about 3 m tall on steep shady slope. Tree to 40 m tall and to 3 m DBH. Occasional emergent at elevation of about 1200 m a.s.l.

29 March 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Tran Thi Thuy Nhan, Nguyen Quoc Bao, T. Maisak, АL4624.

Convallariaceae

Peliosanthes macrostegia Hance⁣ var. subcapitata Aver.

Gia Lai Province, K'Bang District. Highly degraded secondary evergreen broad-leaved dry forest at elevation 600-800 m a.s.l., around point 14º26N 108º22E. Terrestrial herb on shady slope. Flowers dirty violet, fruits glossy blue. Locally common.

7 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4768.

Convallariaceae

Peliosanthes macrostegia Hance⁣ var. subcapitata Aver.

Gia Lai Province, K'Bang District. Highly degraded secondary evergreen broad-leaved dry forest at elevation 600-800 m a.s.l., around point 14º26N 108º22E. Terrestrial herb on shady slope. Flowers dirty violet, fruits glossy blue. Locally common.

7 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4768.

Convallariaceae

Peliosanthes macrostegia Hance⁣ var. subcapitata Aver.

Gia Lai Province, K'Bang District. Highly degraded secondary evergreen broad-leaved dry forest at elevation 600-800 m a.s.l., around point 14º26N 108º22E. Terrestrial herb on shady slope. Flowers dirty violet, fruits glossy blue. Locally common.

7 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4768.

Convallariaceae

Tupistra fungilliformis F.T.Wang & S.Yun Liang

Lao Cai Province, Bat Xat district, Y Ty commune

28 November 2025. Truong Ba Vuong, BV 1955.

Epiphytic plant, inflorescence oblique to pendulous, rachis square (4 angles can be see at number IMG_1110), flower sessile, odorless.

Convallariaceae

Tupistra fungilliformis F.T.Wang & S.Yun Liang

Lao Cai Province, Bat Xat district, Y Ty commune

28 November 2025. Truong Ba Vuong, BV 1955.

Epiphytic plant, inflorescence oblique to pendulous, rachis square (4 angles can be see at number IMG_1110), flower sessile, odorless.

Convallariaceae

Aspidistra bicolor Tillich

Cao Bang Province, Phuc Hoa District (Phục Hòa), Hanh Phuc Commune (Hạnh Phúc),

Lung Luong Village (Lũng Luông). Plants in the population with 2 types of inflorescences (?), one is shorter quite close to the stem, the other form with much longer, erect inflorescence. Flower odorless. Leaves sometimes with sparsely yellowish dots.

17 December 2025, Truong Ba Vuong, BV 1956.

Gesneriaceae

Didymocarpus hanhnguyenae V.C.Nguyen, Vuong, Aver. & C.V.Lin⁣.

Kon Tum Province, Tu MA Rong District.

20 September 2025, Nguyen Van Canh, Nguyen Van Khuong, Truong Ba Vuong BV 1990

Convallariaceae

Ophiopogon tonkinensis L.Rodr.?

Cao Bang Province, Trung Khanh District.

16 November 2025, Truong Ba Vuong, BV 1994

Asclepiadaceae

Hoya globulosa Dean (= H. villosa Costantin).

Gia Lai Province, K'Bang District. Secondary and remnants of primary evergreen broad-leaved dry forest with Dacrydium on lava rocks (red soils) at elevation about 1000 m a.s.l., around point 14º15N 108º35E. Herbaceous to semi-woody vine to 4 m long on tall trees. Occasional.

6 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4791.

Cycadaceae

Cycas micholitzii Dyer

Gia Lai Province, K'Bang District. Secondary and remnants of primary evergreen broad-leaved dry forest with Dacrydium on lava rocks (red soils) at elevation about 1000 m a.s.l., around point 14º15N 108º35E. Terrestrial caudex palm-like plant with leaves to 1.5 m long on steep river slope. Occasional.

6 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4790.

Podocarpaceae

Dacrycarpus imbricatus (Blume) de Laub.

Gia Lai Province, K'Bang District. Secondary and remnants of primary evergreen broad-leaved dry forest with Dacrydium on lava rocks (red soils) at elevation about 1000 m a.s.l., around point 14º15N 108º35E. Sapling of tree to 35 m tall. Occasional co-dominant of the first forest stratum.

6 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4788.

Podocarpaceae

Dacrydium elatum (Roxb.) Hook.

Gia Lai Province, K'Bang District. Secondary and remnants of primary evergreen broad-leaved dry forest with Dacrydium on lava rocks (red soils) at elevation about 1000 m a.s.l., around point 14º15N 108º35E. Sapling of three to 35 m tall. Occasional co-dominant of the first forest stratum.

6 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, Nguyen Quoc Bao, T. Maisak, АL4783.

Rubiaceae

Psychotria sp.

Lam Dong Province, Lac Duong District. Humid evergreen broad-leaved forest with Ducampopinus on granite along tops of ridge at elevation of about 1750 m a.s.l. around point 12º06’N 108º22’E. Terrestrial undershrub about 1.5 m tall along forest trail. Flowers white. Common.

18 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, T. Maisak, АL4950.

Rubiaceae

Lasianthus sp.

Lam Dong Province, Lac Duong District. Humid evergreen broad-leaved forest with Ducampopinus on granite along tops of ridge at elevation of about 1750 m a.s.l. around point 12º06’N 108º22’E. Shrub 1.5-2 m tall in open place along forest trail. Flowers white with pink-violet tint. Common.

18 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, T. Maisak, АL4948.

Lycopodiaceae

Lycopodium cernuum?

Lam Dong Province, Lac Duong District. Humid evergreen broad-leaved forest with Ducampopinus on granite along tops of ridge at elevation of about 1750 m a.s.l. around point 12º06’N 108º22’E. Scandent suberect vine-like herb to 2.5 m long in open shrubs along roadcut. Common.

18 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, T. Maisak, АL4946.

Gnetaceae

Gnetum sp.

Lam Dong Province, Lac Duong District. Humid evergreen broad-leaved forest with Ducampopinus on granite along tops of ridge at elevation of about 1750 m a.s.l. around point 12º06’N 108º22’E. Woody vine to 30 m long on tall trees along forest trail. Occasional.

18 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, T. Maisak, АL4945.

Rubiaceae

Ophiorrhiza sp.

Lam Dong Province, Lac Duong District. Humid evergreen broad-leaved forest with Ducampopinus on granite along tops of ridge at elevation of about 1750 m a.s.l. around point 12º06’N 108º22’E. Erect herb to 0.5 m tall along wet rocky slope of small stream. Flowers white. Common.

18 April 2026, L. Averyanov, Nguyen Van Canh, Truong Ba Vuong, T. Maisak, АL4944.