Ha Giang Prov., Quan Ba Distr., Tung Vai Commune, Thang Village, around point 23.05349ºN 104.86446ºE, steep rocky slopes of river valley composed with eroded limestone mixed with sandstone at elevation 1000–1100 m a.s.l. Remnants of primary evergreen broad-leaved very humid forest along river. Prostrate creeping lithophytic herb on wet mossy stream rocks. Fruits pale dull blue to almost black. Locally very common. 7 October 2019, L. Averyanov, Nguyen Sinh Khang, T. Maisak, VR 1427 (HN, LE01061116, LE01067015).
Coordinates (geographic position):23° 3′ 12.56″ N, 104° 51′ 52.06″ E ±1000 mRecord creation: 2020-02-07, Leonid Averyanov, PhotoScan D1Citation: Specimen LE 01067015 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01067015
Ha Giang Prov., Quan Ba Distr., Can Ty Commune, Sin Suoi Ho Village, around point 23.09566ºN 105.02142ºE. Very steep mountain slopes composed of solid highly eroded karstic limestone at elevation 800–1000 m a.s.l. Remnants of primary evergreen broad-leaved and coniferous (with Calocedrus rupestris and Xanthocyparis vietnamensis) humid forest. Terrestrial herb or undershrub to 1 m tall in open rpcky place. Flowers purple. Occasional. 11 October 2019, L. Averyanov, Nguyen Sinh Khang, T. Maisak, VR 1558 (HN, LE01067021, LE01070220).
Coordinates (geographic position):23° 5′ 44.38″ N, 105° 1′ 17.11″ E ±1000 mRecord creation: 2020-02-07, Leonid Averyanov, PhotoScan D1Citation: Specimen LE 01067021 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01067021
Ha Giang prov., Quan Ba distr., Bat Dai Son Nature Reserve, Thanh Van commune, Mo Sai village, around point 23º07ʹ46.4ʺN 104º57ʹ56.2ʺE, at elevation 1000-1190 m a.s.l., karstic, highly eroded mountains composed of solid marble-like limestone, primary evergreen broad-leaved and mixed (with Pseudotsuga sinensis) humid forest on steep rocky slopes near mountain tops. Woody vine or ascendent shrub to 3 m tall on rocky mountain slope. Flowers white to yellowish. Occasional. 18 April 2018, L. Averyanov, Nguyen Sinh Khang, Chuong Quang Ngan, T. Maisak, VR 377 (HN, LE01061495 – photo).
Coordinates (geographic position):23° 7′ 46.4″ N, 104° 57′ 56.2″ E ±1000 mRecord creation: 2020-01-05, Leonid Averyanov, PhotoScan D1. Processing status: work_status_1Citation: Specimen LE 01067043 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01067043
Ha Giang prov., Quan Ba distr., Bat Dai Son Nature Reserve, Thanh Van commune, Mo Sai village, around point 23º07ʹ46.4ʺN 104º57ʹ56.2ʺE, at elevation 1000-1190 m a.s.l., karstic, highly eroded mountains composed of solid marble-like limestone, primary evergreen broad-leaved and mixed (with Pseudotsuga sinensis) humid forest on steep rocky slopes near mountain tops. Shrub 2-3 m tall on rocky mountain slope. Flowers white. Occasional. 18 April 2018, L. Averyanov, Nguyen Sinh Khang, Chuong Quang Ngan, T. Maisak, VR 378 (HN, LE01061496 – photo).
Coordinates (geographic position):23° 7′ 46.4″ N, 104° 57′ 56.2″ E ±1000 mRecord creation: 2020-01-05, Leonid Averyanov, PhotoScan D1. Processing status: work_status_1Citation: Specimen LE 01067048 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01067048
Ha Giang Prov., Quan Ba Distr., Can Ty Commune, Sin Suoi Ho Village, around point 23.09566ºN 105.02142ºE. Very steep mountain slopes composed of solid highly eroded karstic limestone at elevation 800–1000 m a.s.l. Remnants of primary evergreen broad-leaved and coniferous (with Calocedrus rupestris and Xanthocyparis vietnamensis) humid forest. Terrestrial herb to 1.5 m tall on shady very steep rocky slope. Flowers white with greenish apex. Very common. 11 October 2019, L. Averyanov, Nguyen Sinh Khang, T. Maisak, VR 1590 (HN, LE01067075, LE01070186).
Coordinates (geographic position):23° 5′ 44.38″ N, 105° 1′ 17.11″ E ±1000 mRecord creation: 2020-02-07, Leonid Averyanov, PhotoScan D1Citation: Specimen LE 01067075 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01067075
Ha Giang prov., Quan Ba distr., Bat Dai Son commune, Bat Dai Son Nature Reserve, San Chu village, around point 23º08ʹ55.1ʺN 104º59ʹ45.7ʺE, 1100-1290 m a.s.l., karstic, highly eroded mountains composed of solid marble-like limestone, primary evergreen coniferous humid forest with Pseudotsuga sinensis on rocky mountain tops and ridges. Terrestrial erect herb to 1 m tall in rather shady place. Inflorescence rachis and flower buds pink. Occasional. 13 April 2018, L. Averyanov, Nguyen Sinh Khang, Chuong Quang Ngan, T. Maisak, VR 017 (HN).
Record creation: 2020-01-05, Leonid Averyanov, PhotoScan D1. Processing status: work_status_1Citation: Specimen LE 01067084 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01067084
Ha Giang prov., Quan Ba distr., Bat Dai Son commune, Bat Dai Son Nature Reserve, San Chu village, around point 23º08ʹ55.1ʺN 104º59ʹ45.7ʺE, 1100-1200 m a.s.l., karstic, highly eroded mountains composed of solid marble-like limestone, primary evergreen broad-leaved humid forest on very steep rocky slopes. Terrestrial erect herb to 0.5 m tall in shady places. Occasional. 13 April 2018, L. Averyanov, Nguyen Sinh Khang, Chuong Quang Ngan, T. Maisak, VR 015 (HN, LE01067085).
Record creation: 2020-02-07, Leonid Averyanov, PhotoScan D1Citation: Specimen LE 01067085 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01067085
Ha Giang Prov., Quan Ba Distr., Thai An Commune, Seo Lung Two Village, around point 22.96988ºN 105.08646ºE. Very steep mountain slopes composed of stratified shale and clayey subsoil at elevation 800–1000 m a.s.l. Primary and secondary evergreen broad-leaved humid forest. Terrestrial herb to 1 m tall on steep shady rocky river slope. Flowers white. Common. 10 October 2019, L. Averyanov, Nguyen Sinh Khang, T. Maisak, VR 1512 (HN, LE01067105, LE01070179).
Coordinates (geographic position):22° 58′ 11.57″ N, 105° 5′ 11.26″ E ±1000 mRecord creation: 2020-02-07, Leonid Averyanov, PhotoScan D1Citation: Specimen LE 01067105 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01067105
Ha Giang Prov., Quan Ba Distr., Thai An Commune, Seo Lung Two Village, around point 22.96988ºN 105.08646ºE. Very steep mountain slopes composed of stratified shale and clayey subsoil at elevation 800–1000 m a.s.l. Primary and secondary evergreen broad-leaved humid forest. Treelet 6-8 m tall on open steep river slope. Inflorescence bract pure white. Common. 10 October 2019, L. Averyanov, Nguyen Sinh Khang, T. Maisak, VR 1540 (HN, LE01067132, LE01070157).
Coordinates (geographic position):22° 58′ 11.57″ N, 105° 5′ 11.26″ E ±1000 mRecord creation: 2020-02-07, Leonid Averyanov, PhotoScan D1Citation: Specimen LE 01067132 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01067132
Ha Giang prov., Quan Ba distr., Bat Dai Son Nature Reserve, Thanh Van commune, Mo Sai village, around point 23º07ʹ46.4ʺN 104º57ʹ56.2ʺE, at elevation 1000-1190 m a.s.l., karstic, highly eroded mountains composed of solid marble-like limestone, primary evergreen broad-leaved and mixed (with Pseudotsuga sinensis) humid forest on steep rocky slopes near mountain tops. Terrestrial herb to 0.5 m tall on shady rocky mountain slope. Flowers whitish. Common. 18 April 2018, L. Averyanov, Nguyen Sinh Khang, Chuong Quang Ngan, T. Maisak, VR 382 (HN, LE01061500 – photo).
Coordinates (geographic position):23° 7′ 46.4″ N, 104° 57′ 56.2″ E ±1000 mRecord creation: 2020-01-05, Leonid Averyanov, PhotoScan D1. Processing status: work_status_1Citation: Specimen LE 01067261 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01067261
Ha Giang prov., Quan Ba distr., Bat Dai Son commune, Bat Dai Son Nature Reserve, San Chu village, around point 23º08ʹ55.1ʺN 104º59ʹ45.7ʺE, 1000-1200 m a.s.l., karstic, highly eroded mountains composed of solid marble-like limestone, primary and partially degraded evergreen broad-leaved humid forest on rocky mountain slopes. Terrestrial and lithophytic herb in shady place. Common. 13 April 2018, L. Averyanov, Nguyen Sinh Khang, Chuong Quang Ngan, T. Maisak, VR 073 (HN, LE01058104, LE01067557).
Record creation: 2020-02-07, Leonid Averyanov, PhotoScan D1Citation: Specimen LE 01067557 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01067557
Ha Giang prov., Quan Ba distr., Bat Dai Son Nature Reserve, Can Ty commune, Lung Vay village, around point 23º05ʹ46.7ʺN 105º02ʹ53.1ʺE, 1100-1280 m a.s.l., karstic, highly eroded mountains composed of solid marble-like limestone, primary evergreen broad-leaved humid forest on steep rocky slopes near mountain tops. Erect terrestrial and lithophytic herb to 1.5 m tall among shady rocks. Young flower buds light green. Occasional. 15 April 2018, L. Averyanov, Nguyen Sinh Khang, Chuong Quang Ngan, T. Maisak, VR 168 (HN, LE01058128, LE01067558).
Record creation: 2020-02-07, Leonid Averyanov, PhotoScan D1Citation: Specimen LE 01067558 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01067558
Ha Giang Prov., Quan Ba Distr., Thai An Commune, Seo Lung Two Village, around point 22.96988ºN 105.08646ºE. Very steep mountain slopes composed of stratified shale and clayey subsoil at elevation 800–1000 m a.s.l. Primary and secondary evergreen broad-leaved humid forest. Treelet 6-8 m tall on open steep river slope. Inflorescence bract pure white. Common. 10 October 2019, L. Averyanov, Nguyen Sinh Khang, T. Maisak, VR 1540.
Coordinates (geographic position):22° 58′ 11.57″ N, 105° 5′ 11.26″ E ±1000 mComment:
Record creation: 2020-01-04, Аверьянов Леонид ВладимировичCitation: Specimen LE 01070157 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01070157
Ha Giang Prov., Quan Ba Distr., Thai An Commune, Seo Lung Two Village, around point 22.96988ºN 105.08646ºE. Very steep mountain slopes composed of stratified shale and clayey subsoil at elevation 800–1000 m a.s.l. Primary and secondary evergreen broad-leaved humid forest. Terrestrial herb to 1 m tall on steep shady rocky river slope. Flowers white. Common. 10 October 2019, L. Averyanov, Nguyen Sinh Khang, T. Maisak, VR 1512.
Coordinates (geographic position):22° 58′ 11.57″ N, 105° 5′ 11.26″ E ±1000 mComment:
Record creation: 2020-01-04, Аверьянов Леонид ВладимировичCitation: Specimen LE 01070179 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01070179
Ha Giang Prov., Quan Ba Distr., Can Ty Commune, Sin Suoi Ho Village, around point 23.09566ºN 105.02142ºE. Very steep mountain slopes composed of solid highly eroded karstic limestone at elevation 800–1000 m a.s.l. Remnants of primary evergreen broad-leaved and coniferous (with Calocedrus rupestris and Xanthocyparis vietnamensis) humid forest. Terrestrial herb to 1.5 m tall on shady very steep rocky slope. Flowers white with greenish apex. Very common. 11 October 2019, L. Averyanov, Nguyen Sinh Khang, T. Maisak, VR 1590.
Coordinates (geographic position):23° 5′ 44.38″ N, 105° 1′ 17.11″ E ±1000 mComment:
Record creation: 2020-01-04, Аверьянов Леонид ВладимировичCitation: Specimen LE 01070186 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01070186
Ha Giang Prov., Quan Ba Distr., Can Ty Commune, Sin Suoi Ho Village, around point 23.09566ºN 105.02142ºE. Very steep mountain slopes composed of solid highly eroded karstic limestone at elevation 800–1000 m a.s.l. Remnants of primary evergreen broad-leaved and coniferous (with Calocedrus rupestris and Xanthocyparis vietnamensis) humid forest. Terrestrial herb or undershrub to 1 m tall in open rpcky place. Flowers purple. Occasional. 11 October 2019, L. Averyanov, Nguyen Sinh Khang, T. Maisak, VR 1558.
Coordinates (geographic position):23° 5′ 44.38″ N, 105° 1′ 17.11″ E ±1000 mComment:
Record creation: 2020-01-04, Аверьянов Леонид ВладимировичCitation: Specimen LE 01070220 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01070220
FLORA OF VIETNAM Rubiaceae Leptomischus hiepii L. Wu, K.S. Nguyen & Aver. sp nov. Son La province, Muong La district, Muong La N.R., Ngoc Chien commune. Primary evergreen broad-leaved forest along streams at the base and slope of sandstone mountain, around point 21.61032° N 104.10576° E, at elevation of 1350 m a.s.l. Lithophytic perennial herbs around 30-45 cm tall, stem sub-erect, flower pure white, calyx and corolla glabrous adaxially, white pubescence abaxially, very common at waterfalls and humid places in a restricted area (around 2500 m2). March 2, 2019, Nguyen Sinh Khang, Nguyen Duc Thang et al., NSK 1153 (holotype CSFI; isotype CSFI, HN, LE 01058686, LE 01058688, LE 01073034 - photo), including living plants and DNA samples (HN)
Coordinates (geographic position):21° 36′ 37.15″ N, 104° 6′ 20.74″ E ±1000 mComment:
Record creation: 2020-03-18, Аверьянов Леонид ВладимировичCitation: Specimen LE 01073034 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01073034
FLORA OF VIETNAM Rubiaceae Gaertnera vaginans (DC.) Merr.
Quang Binh Prov., Minh Hoa Distr., Thuong Hoa municipality, around point 17°41′28”N 105°53′42.7”E.
Primary closed evergreen broad-leaved forest on middle part of slope of remnant mountain composed with solid crystalline limestone at elevation about 650 m a.s.l.
Shrub about 4 m tall on mountain top. Fruits dark green. Common on mountain top.
FLORA OF VIETNAM Rubiaceae Lasianthus biflorus (Blume) M.Gangop. et Chakrab.
Quang Binh Prov., Minh Hoa Distr., Thuong Hoa municipality, environs of Mo O O O village, around point 17°39′31.7”N 105°54′48.2”E.
Closed broad-leaved primary forest on very steep rocky slope of stream valley composed with highly eroded solid crystalline limestone at elevation about 422 m a.s.l.
Shrub 1-2 m tall. Flowers white. Common.
CPC 3888 28 July 2011 Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Vinh.
Coordinates (geographic position):17° 39′ 31.7″ N, 105° 54′ 48.2″ E ±1000 mRecord creation: 2021-11-24, Tatiana Maisak, PhotoScan D2. Processing status: work_status_1Citation: Specimen LE 01077481 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01077481
FLORA OF VIETNAM Rubiaceae Hedyotis acutangula Champ. ex Benth.
Quang Binh Prov., Minh Hoa Distr., Thuong Hoa municipality, environs of Mo O O O village, around point 17°39′21.1”N 105°54′41.7”E.
Closed broad-leaved evergreen primary forest on steep rocky slope of middle part of remnant mountain composed with highly eroded solid crystalline limestone at elevation 400-450 m a.s.l. (mt summit at about 570 m).
Terrestrial and lithophytic herb to 0.5 m tall on rocky slope. Flowers white. Very common.
CPC 3879 28 July 2011 Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Vinh.
Coordinates (geographic position):17° 39′ 21.1″ N, 105° 54′ 41.7″ E ±1000 mRecord creation: 2021-11-24, Tatiana Maisak, PhotoScan D2. Processing status: work_status_1Citation: Specimen LE 01077482 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01077482
Quang Binh Prov., Minh Hoa Distr., Thuong Hoa municipality, around point 17°41′28”N 105°53′42.7”E.
Wet primary evergreen broad-leaved forest on shady humid rocky slope of narrow stream canyon composed with solid crystalline limestone at elevation about 600 m a.s.l.
Woody vine about 4 m long in open place. Fruits green. Occasional.
Quang Binh Prov., Minh Hoa Distr., Thuong Hoa municipality, environs of Mo O O O village, around point 17°39′11.6”N 105°54′53”E.
Primary broad-leaved closed forest on very steep rocky slope of remnant mountain composed with highly eroded solid crystalline limestone at elevation 400-500 m a.s.l.
Shrub 2-2.5 m tall. Fruits green. Very common.
CPC 3675 24 July 2011 Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Vinh.
Coordinates (geographic position):17° 39′ 11.6″ N, 105° 54′ 53″ E ±1000 mRecord creation: 2021-11-24, Tatiana Maisak, PhotoScan D2. Processing status: work_status_1Citation: Specimen LE 01077485 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01077485
FLORA OF VIETNAM Rubiaceae Hedyotis acutangula Champ. ex Benth.
Quang Binh Prov., Minh Hoa Distr., Thuong Hoa municipality, environs of Mo O O O village, around point 17°39′36.7”N 105°54′55.7”E.
Remnants of short tall broad-leaved primary forest on rocky top of remnant mountain composed with highly eroded solid crystalline limestone at elevation about 612 m a.s.l.
Lithophytic herb on shady very steep rocky slope near to mountain top. Flowers white. Common.
CPC 3769 26 July 2011 Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Vinh.
Coordinates (geographic position):17° 39′ 36.7″ N, 105° 54′ 55.7″ E ±1000 mRecord creation: 2021-11-24, Tatiana Maisak, PhotoScan D2. Processing status: work_status_1Citation: Specimen LE 01077486 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01077486
Quang Binh Prov., Minh Hoa Distr., Thuong Hoa municipality, environs of Mo O O O village, around point 17°39′36.7”N 105°54′55.7”E.
Remnants of short tall broad-leaved primary forest on rocky top of remnant mountain composed with highly eroded solid crystalline limestone at elevation about 612 m a.s.l.
Woody vine to 5 m long on rocky mountain top. Not rare.
CPC 3781 26 July 2011 Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Vinh.
Coordinates (geographic position):17° 39′ 36.7″ N, 105° 54′ 55.7″ E ±1000 mRecord creation: 2021-11-24, Tatiana Maisak, PhotoScan D2. Processing status: work_status_1Citation: Specimen LE 01077487 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01077487
FLORA OF VIETNAM Rubiaceae Brachytome wallichii Hook.f.?
Quang Binh Prov., Minh Hoa Distr., Thuong Hoa municipality, environs of Mo O O O village, around point 17°39′36.7”N 105°54′55.7”E.
Remnants of short tall broad-leaved primary forest on rocky top of remnant mountain composed with highly eroded solid crystalline limestone at elevation about 612 m a.s.l.
Shrub about 2 m tall on rocky mountain top. Fruits dark green. Very common. Co-dominant of short mountain top forest.
CPC 3784 26 July 2011 Coll.: N.T.Hiep, L.Averyanov, N.S.Khang, N.Q.Vinh.
Coordinates (geographic position):17° 39′ 36.7″ N, 105° 54′ 55.7″ E ±1000 mRecord creation: 2021-11-24, Tatiana Maisak, PhotoScan D2. Processing status: work_status_1Citation: Specimen LE 01077488 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01077488
Da Nang City, Hoa Vang Distr., Hoa Ninh municipality, Ba Na – Nui Chua protected area, around point 15°59′52”N 107°59′04”E.
Highly destroyed primary and secondary broadleaved evergreen forest on very steep mountain slopes with granite mother rocks at elev. 1200-1500 m a.s.l.
Creeping semi-woody vine to 5 m long in open slope along road cut. Flowers white. Common.
FLORA OF VIETNAM NEW Rubiaceae Spiradiclis loana R. J. Wang
Thanh Hoa Prov., Thuong Xuan Distr., Van Xuan Municipality, Hang Cao village, Xuan Lien Natural Reserve.
Remnants of primary and secondary broad-leaved evergreen forest on highly eroded rocky limestone hills at elevation 100-200 m a.s.l. around point 19°50′47.2″N 105°14′42.7″E.
Lithophytic herb on vertical shady cliff. Locally common.
FLORA OF VIETNAM Rubiaceae Uncaria macrophylla Wall. Verde.Ha Giang Prov., Vi Xuyen Distr., Tung Ba municipality, Khau Ca NR. Primary broad-leaved humid evergreen forest on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation 700-1150 m a.s.l. around point 22°50′43.9″N 105°07′18.5″E. Shrub climbering 4-5 m high. Common. 09 May 2015 CPC 7849 Coll.: N.Q. Hieu, N.T.Hiep, T.B. Ngan, C.Q. Ngan, D.K. Harder, C. P. Tuy, G.C.Pao
Coordinates (geographic position):22° 50′ 43.9″ N, 105° 7′ 18.5″ E ±1000 mRecord creation: 2021-11-24, Tatiana Maisak, PhotoScan D2. Processing status: work_status_1Citation: Specimen LE 01077503 // Virtual herbarium of Komarov Botanical Institute RAS — http://re.herbariumle.ru/01077503
Border area between Phieng Luong municipality (Ha Giang Prov., Bac Me Distr.) and Sinh Long municipalitiy (Tuen Quang Prov., Na Hang Distr.).
Primary broad-leaved humid evergreen and mixed forest on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation 1100-1170 m a.s.l. around point 22°38′24.3″N 105°20′21.3″E.
Scandent shrub about 2 m tall on slopes and on base of rocky limestone mountain. Fruits dark green. Not common.