Herbarium LE - Specimens

FLORA OF LAOS Nakai-Nam Theun Protected Area Polypodiaceae s.l. (Pteridaceae) Pteris venusta Kunze Laos, Khammoune prov., Boualapa distr., Langkhang village, steep southern slopes of sandstone ridge allied to the road from the north between points 17°34'02.2"N 105°42'42.6"E, 200 m and 17°34'51.1"N 105°43'01.2"E, 535 m a.s.l. Primary lowland semideciduous and evergreen dry forest on very steep rocky slopes. Terrestrial rhizomatous fern on steep rocky slope. Occasional. 15 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, AL 1077.

FLORA OF LAOS Hin Nam No Nature Protected Area Polypodiaceae s.l. (Dryopteridaceae) Bolbitis virens (Hook. & Grev.) Schott Laos, Khammouane prov., Boualapha distr., Nong Seng village, southern sandstone slopes of Phou Chuang (Poujeuang) Mt. around point 17°33'14.4"N, 105°47'50.4"E. Primary semideciduous and evergreen broad-leaved dry forest at elev. 300-400 m a.s.l. Lithophytic creeping fern on shady mossy rocks. Occasional. 6 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 739.

FLORA OF LAOS Nakai-Nam Theun Protected Area Polypodiaceae s.l. (Pteridaceae) Calciphilopteris ludens (Hook.) Yesilyurt & H. Schneid. ( - Doryopteris ludens (Hook.) J. Sm.) Laos, Khammoune prov., Boualapa distr.. Langkhang village, steep southern slopes of sandstone ridge allied to the road from the north betw een points 17°34'02.2"N 105°42'42.6"E, 200 m and 17°34'5L1"N 105°43’01.2"E, 535 m a.s.l. Primary lowland semideciduous and evergreen dry forest on very steep rocky slopes. Terrestrial rhizomatous fern on very steep rocky slopes and cliffs. Locally very common. 15 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak. AL 1090.

FLORA OF LAOS Hin Nam No Nature Protected Area Polypodiaceae s.l. (Dryopteridaceae) Polystichum hookerianum (C.Presl) C.Chr. Laos, Khammoune prov., Boualapha distr., Vang Maner village, Palai mountain area, around point 17°30'13.6"N, 105°49'23.9"E, 520 m a.s.l. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 300-500 m a.s.l. Terrestrial fern in shady place. Occasional. 8 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 756.

FLORA OF LAOS Hin Nam No Nature Protected Area Polypodiaceae s.l. (Dryopteridaceae) Bolbitis appendiculata (Willd.) K.Iwats. Laos, Khammouane prov., Boualapha distr., Nong Seng village, southern sandstone slopes of Phou Chuang (Poujeuang) Mt. around point 17°33'14.4"N, 105°47'50.4"E. Primary semideciduous and evergreen broad-leaved dry forest at elev. 400-600 m a.s.l. Creeping lithophytic fern on shady mossy rocks. Very common. 6 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 735.

FLORA OF LAOS Hin Nam No Nature Protected Area Polypodiaceae s.l. (Pteridaceae) Pteris venusta Kunze Laos, Khammouane prov., Boualapha distr., Nong Seng village, southern sandstone slopes of Phou Chuang (Poujeuang) Mt. around point 17°33'14.4"N, 105°47'50.4"E. Primary semideciduous and evergreen broad-leaved dry forest at elev. 350-450 m a.s.l. Terrestrial fern on steep shady rocky slope. Very common. 6 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 724.

FLORA OF LAOS Hin Nam No Nature Protected Area Polypodiaceae s.l. (Pteridaceae) Antrophyum coriaceum (D. Don) T. Moore Laos, Khammoune prov., Boualapha distr., Vang Maner village, Palai mountain area, around point 17°30'13.6"N, 105°49'23.9"E, 520 m a.s.l. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 300-500 m a.s.l. Epiphyte on shady place. Occasional. 8 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 744.

FLORA OF LAOS Hin Nam No Nature Protected Area Polypodiaceae s.l. (Pteridaceae) Cheilosoria belangeri (Bory) Ching & Shing ( = Cheilanthes belangeri (Bory) C. Chr.) Laos, Khammoune prov., Boualapha distr., Nong Seng village, Chanh Daeng mountain, around point 17°31'11.1"N, 105°46'41.4"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 400-600 m a.s.l. Lithophytic fern on exposed rocks near mountain top. Not common. 11 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 1000.

FLORA OF LAOS Nakai-Nam Theun Protected Area Polypodiaceae s.l. (Pteridaceae) Pteris venusta Kunze Laos, Khammoune prov., Boualapa distr., Langkhang village, steep southern slopes of sandstone ridge allied to the road from the north between points 17°34'02.2"N 105°42'42.6"E, 200 m and 17°34'51.1"N 105°43'01.2"E, 535 m a.s.l. Primary lowland semideciduous and evergreen dry forest on very steep rocky slopes. Terrestrial rhizomatous fern on steep rocky slope. Occasional. 15 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, AL 1077.

FLORA OF LAOS Hin Nam No Nature Protected Area Polypodiaceae s.l. (Pteridaceae) Adiantum flabellulatum L. Laos, Khammoune prov., Boualapha distr., Nong Seng village, Vang Phae Lay mountain, around point 17°31'02.7"N, 105°47'17.2"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 300-530 m a.s.l. Terrestrial fern on shady rocky slope. Occasional. 11 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 931.

FLORA OF LAOS Hin Nam No Nature Protected Area Polypodiaceae s.l. (Pteridaceae) Calciphilopteris ludens (Hook.) Yesilyurt & H. Schneid. ( = Doryopteris ludens (Hook.) J. Sm.) Laos, Khammoune prov., Boualapha distr., Nong Seng village, Vang Phae Lay mountain, around point 17°31'02.7"N, 105°47'17.2"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 300-530 m a.s.l. Terrestrial and lithophytic fern on very steep rocky slope near mountain top. Not rare. 11 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 917.

FLORA OF LAOS Hin Nam No Nature Protected Area Polypodiaceae s.l. (Pteridaceae) Antrophyum coriaceum (D. Don) T. Moore Laos, Khammoune prov., Boualapha distr., Nong Seng village, Vang Phae Lay mountain, around point 17°31'02.7"N, 105°47'17.2"E. Primary dry evergreen and semideciduous broad-leaved forest on very steep rocky slope near karstic hill top composed with solid, marble-like highly eroded limestone at 300-530 m a.s.l. Epiphyte on shady rocky slope. Not common. 11 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 913.

FLORA OF LAOS Hin Nam No Nature Protected Area Polypodiaceae s.l. (Thelypteridaceae) Cyclosorus parasiticus (L.) Farw. Laos, Khammouane prov., Boualapha distr., Nong Seng village, southern sandstone slopes of Phou Chuang (Poujeuang) Mt. around point 17°34'13.3"N 105°47'57.5"E. Primary evergreen broad-leaved dry forest at elev. 700-1000 m a.s.l. Terrestrial and lithophytic fern among shady stream rocks. Very common. 3 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 589.

FLORA OF LAOS Hin Nam No Nature Protected Area Polypodiaceae s.l. (Dryopteridaceae) Bolbitis sinensis (Baker) K.Iwats. Laos, Khammouane prov., Boualapha distr., Nong Seng village, southern sandstone slopes at summit of Phou Chuang (Poujeuang) Mt. around point 17°35'18.1"N, 105°47'47.5"E. Primary evergreen mixed and coniferous dense wind-formed forest at elev. 1000-1400 m a.s.l. Lithophytic fern on shady rocks. Common. 5 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 651a.

FLORA OF LAOS Hin Nam No Nature Protected Area Polypodiaceae s.l. (Pteridaceae) Haplopteris flexuosa (Fee) E.H. Crane ( = Vittaria flexuosa Fee) Laos, Khammouane prov., Boualapha distr., Nong Seng village, southern sandstone slopes at summit of Phou Chuang (Poujeuang) Mt. around point 17°35'18.1"N, 105°47'47.5"E. Primary evergreen mixed and coniferous dense wind-formed forest at elev. 1000-1400 m a.s.l. Epiphytic and occasionally lithophytic fern in shady places on mountain top. Locally common. 5 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 659a.

FLORA OF LAOS Hin Nam No Nature Protected Area Polypodiaceae s.l. (Dryopteridaceae) Polystichum biaristatum (Blume) T. Moore Laos, Khammouane prov., Boualapha distr., Nong Seng village, southern sandstone slopes of Phou Chuang (Poujeuang) Mt. around point 17°34'13.3"N 105°47'57.5"E. Primary evergreen broad-leaved dry forest at elev. 700-1000 m a.s.l. Terrestrial and lithophytic fern in shady places. Very common. 3 May 2018, L. Averyanov, Khang Sinh Nguyen, T. Maisak, Littideth Xaiyavongsa, Siphanom Keovankham, AL 556.

FLORA OF VIETNAM Polypodiaceae s.l. Dryopteridaceae Bolbitis sp. Ha Giang prov., Quan Ba distr., Tung Vai commune, Thang village, around point 23º03ʹ41.5ʺN 104º50ʹ41.6ʺE, steep rocky slopes near mountain top composed with eroded stratified highly eroded limestone at elevation 1200-1400 m a.s.l. Primary evergreen broad-leaved very humid forest. Creeping lithophyte on mossy rocks. Rare. 22 April 2018, L. Averyanov, Nguyen Sinh Khang, Nguyen Tien Hiep, Nguyen Quang Hieu, Chuong Quang Ngan, T. Maisak, VR 706.

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM Polypodiaceae s.l. Dryopteridaceae Cyrtomium fortunei J. Sm. Ha Giang prov., Quan Ba distr., Bat Dai Son Nature Reserve, Can Ty commune, Dau Cau 1 village, around point 23º05ʹ26.3ʺN 105º01ʹ03.0ʺE, 1100-1150 m a.s.l., karstic, highly eroded mountains composed of solid marble-like limestone, primary evergreen broad-leaved humid forest on steep rocky slopes near mountain tops. Terrestrial and lithophytic rosulate fern on very steep slope. Not rare. 14 April 2018, L. Averyanov, Nguyen Sinh Khang, Chuong Quang Ngan, T. Maisak, VR 128.

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM Polypodiaceae s.l. Dryopteridaceae Cyrtomium fortunei J. Sm. Ha Giang prov., Quan Ba distr., Bat Dai Son Nature Reserve, Can Ty commune, Dau Cau 1 village, around point 23º05ʹ26.3ʺN 105º01ʹ03.0ʺE, 1100-1150 m a.s.l., karstic, highly eroded mountains composed of solid marble-like limestone, primary evergreen broad-leaved humid forest on steep rocky slopes near mountain tops. Terrestrial and lithophytic rosulate fern on very steep slope. Not rare. 14 April 2018, L. Averyanov, Nguyen Sinh Khang, Chuong Quang Ngan, T. Maisak, VR 128.

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM Polypodiaceae s.l. Dryopteridaceae Polystichum hookerianum (C.Presl) C.Chr. Ha Giang prov., Quan Ba distr., Cao Ma Po commune, Chin Chu Lin village, around point 23º04ʹ47.2ʺN 104º48ʹ55.6ʺE, steep slopes and flattened top of mountain composed with sandstone at elevation 1800–1950 m a.s.l. Primary evergreen broad-leaved very humid forest. Terrestrial fern on very steep mossy slope. Common. 21 October 2018, L. Averyanov, Nguyen Sinh Khang, T. Maisak, Truong Duc Thieu, VR 1139.

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM.

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM.

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM Dryopteridaceae Cyrtomium hemionitis Christ Ha Giang province, Quan Ba district, Cao Ma Po commune, Vang Cha Phin village, around point point 23°05?01??N, 104°48?38??E, at elevation of 1770 m a. s. l. Secondary evergreen broad-leaved forest on silicate mountain mixed limestone rock. Terrestrial herb around 18-35 cm, leaf scare brown. Not common. NSK 1101, July 4, 2018. Nguyen Sinh Khang & Pin.

FLORA OF VIETNAM

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM Polypodiaceae s.l. (Thelypteridaceae) Cyclosorus ciliatus (Wall. ex Benth.) Panigrahi Ha Giang Prov., Quan Ba Distr., Tung Vai Commune, Thang Village, around point 23.05349ºN 104.86446ºE, steep rocky slopes of river valley composed with eroded limestone mixed with sandstone at elevation 1000–1100 m a.s.l. Remnants of primary evergreen broad-leaved very humid forest along river. Lithophytic fern on wet mossy stream rocks. Very common. 7 October 2019, L. Averyanov, Nguyen Sinh Khang, T. Maisak, VR 1411.

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM Dipteridaceae Dipteris chinensis Christ Lam Dong Province, Bao Loc City Area, old secondary dry evergreen broad-leaved and mixed (with Pinus kesiya) submontane forest on granite at elevation about 1000 m a.s.l. on top of hill. Terrestrial fern with fronds to 1.5 m tall on steep wet shady road cut. Not common. 27 October 2019, L. Averyanov, Nguyen Van Canh, T. Maisak, AL 1204.4 (photo only).

All photos: © L. Averyanov

FLORA OF VIETNAM Polypodiaceae s.l. (Pteridaceae) Coniogramme intermedia Hieron. Ha Giang Prov., Quan Ba Distr., Cao Ma Po Commune, Then Van 1 Village, Binh Mountain, around point 23.09452ºN 104.84831ºE, steep rocky slopes composed with highly eroded limestone at elevation 1400–1600 m a.s.l. Remnants of primary evergreen broad-leaved very humid forest with plantation of “wild-grown” Amomum. Terrestrial fern on shady rocky slope. Common. 4 October 2019, L. Averyanov, Nguyen Sinh Khang, T. Maisak, VR 1307.

FLORA OF VIETNAM Polypodiaceae s.l. (Thelypteridaceae) Cyclosorus erubescens (Wall. ex Hook.) C.M. Kuo Ha Giang Prov., Quan Ba Distr., Tung Vai Commune, Thang Village, around point 23.05349ºN 104.86446ºE, steep rocky slopes of river valley composed with eroded limestone mixed with sandstone at elevation 1000–1100 m a.s.l. Remnants of primary evergreen broad-leaved very humid forest along river. Terrestrial fern on very steep river slope. Fronds to 1.6 m tall. Common. 7 October 2019, L. Averyanov, Nguyen Sinh Khang, T. Maisak, VR 1443.

FLORA OF VIETNAM Polypodiaceae s.l. (Thelipteridaceae) Cyclosorus subpubescens (Blume) Ching Ha Giang Prov., Quan Ba Distr., Thai An Commune, Seo Lung Two Village, around point 22.96988ºN 105.08646ºE. Very steep mountain slopes composed of stratified shale and clayey subsoil at elevation 800–1000 m a.s.l. Primary and secondary evergreen broad-leaved humid forest. Terrestrial fern on very steep wet shady slope. Fronds to 2 m long. Common. 10 October 2019, L. Averyanov, Nguyen Sinh Khang, T. Maisak, VR 1501.

FLORA OF VIETNAM Polypodiaceae s.l. (Dryopteridaceae) Peranema aspidioides (Blume) Mett. Ha Giang Prov., Quan Ba Distr., Thai An Commune, Seo Lung Two Village, around point 22.96988ºN 105.08646ºE. Very steep mountain slopes composed of stratified shale and clayey subsoil at elevation 800–1000 m a.s.l. Primary and secondary evergreen broad-leaved humid forest. Terrestrial ferb on open shrubby place on steep stream slope. Fronds to 1.2 m long. Common. 10 October 2019, L. Averyanov, Nguyen Sinh Khang, T. Maisak, VR 1505.

FLORA OF VIETNAM Polypodiaceae s.l. (Dryopteridaceae) Bolbitis heteroclita (Pr.) Ching Ha Giang Prov., Quan Ba Distr., Tung Vai Commune, Thang Village, around point 23.05349ºN 104.86446ºE, steep rocky slopes of river valley composed with eroded limestone mixed with sandstone at elevation 1000–1100 m a.s.l. Remnants of primary evergreen broad-leaved very humid forest along river. Terrestrial and lithophytic fern on steep shady river slope near to water line. Very common. 7 October 2019, L. Averyanov, Nguyen Sinh Khang, T. Maisak, VR 1422.

FLORA OF VIETNAM Polypodiaceae s.l. Dryopteridaceae Cyrtogonellum fraxinellum (Christ) Ching Ha Giang prov., Quan Ba distr., Bat Dai Son Nature Reserve, Thanh Van commune, Mo Sai village, around point 23º07ʹ46.4ʺN 104º57ʹ56.2ʺE, at elevation 1000-1190 m a.s.l., karstic, highly eroded mountains composed of solid marble-like limestone, primary evergreen broad-leaved and mixed (with Pseudotsuga sinensis) humid forest on steep rocky slopes near mountain tops. Terrestrial and lithophytic fern on mossy rocky mountain top. Common. 18 April 2018, L. Averyanov, Nguyen Sinh Khang, Chuong Quang Ngan, T. Maisak, VR 343 (HN, LE01058181 – photo).

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM Polypodiaceae s.l. Dryopteridaceae Cyrtomium hemionitis Christ Ha Giang prov., Quan Ba distr., Bat Dai Son Nature Reserve, Thanh Van commune, Mo Sai village, around point 23º07ʹ46.4ʺN 104º57ʹ56.2ʺE, at elevation 1000-1190 m a.s.l., karstic, highly eroded mountains composed of solid marble-like limestone, primary evergreen broad-leaved and mixed (with Pseudotsuga sinensis) humid forest on steep rocky slopes near mountain tops. Terrestrial and lithophytic fern on mossy rocky mountain top. Common. 18 April 2018, L. Averyanov, Nguyen Sinh Khang, Chuong Quang Ngan, T. Maisak, VR 349 (HN, LE01058187, LE01058188 – photos).

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM Polypodiaceae s.l. Dryopteridaceae Polystichum dielsii Christ Ha Giang prov., Quan Ba distr., Bat Dai Son Nature Reserve, Can Ty commune, Lung Vay village, around point 23º05ʹ46.7ʺN 105º02ʹ53.1ʺE, 1100-1280 m a.s.l., karstic, highly eroded mountains composed of solid marble-like limestone, primary evergreen broad-leaved humid forest on steep rocky slopes near mountain tops. Creeping lithophytic or epiphytic fern in shady place. Occasional. 15 April 2018, L. Averyanov, Nguyen Sinh Khang, Chuong Quang Ngan, T. Maisak, VR 170 (HN, LE01058130 – photo).

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM Polypodiaceae s.l. Dryopteridaceae Polystichum dielsii Christ Ha Giang prov., Quan Ba distr., Bat Dai Son Nature Reserve, Thanh Van commune, Tan village, Thong (Conifer) Mountain, around point 23º07ʹ40.3ʺN 104º57ʹ11.2ʺE, at elevation 1000-1195 m a.s.l., karstic, highly eroded mountains composed of solid marble-like limestone, primary evergreen broad-leaved and coniferous (with Pseudotsuga sinensis) humid forest on steep rocky slopes near mountain tops. Terrestrial and lithophytic fern among mossy rocks. Not rare. 19 April 2018, L. Averyanov, Nguyen Sinh Khang, Chuong Quang Ngan, T. Maisak, VR 484 (HN, LE01061407 – photo).

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM

All photos © L. Averyanov & K.S. Nguyen

FLORA OF VIETNAM
Dryopteridaceae
Diacalpe aspidioides Blume

Northern Vietnam, Lao Cai Prov., Van Ban Distr., Khanh Yen Ha
municipality, to the SW of Na Nheo village (in the vicinity of point
21°57′34′'N, 104°13′49′'E). Primary evergreen broad-leaved wet montane
forest with Fokienia hodginsii on steep granite and quartzite slopes
of SW exposition at elev. 1800-1900 m a.s.l. Terrestrial fern in shady
place. Not rare. HAL 2507 18 March 2002

Collectors: L.Averyanov, P.K.Loc, D.T.Doan (FFI Project).

FLORA OF VIETNAM

All photos © Nguyen Sinh Khang